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<document id="BA1D3C06661A8D4CEADB5C717E994AA0" ID-DOI="10.5281/zenodo.203960" ID-GBIF-Dataset="24889de9-856a-41ba-a31b-3faf1828f314" ID-ISSN="1175-5326" ID-Zenodo-Dep="203960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460322587115" checkinUser="plazi" docAuthor="Kaydan, Bora" docDate="2011" docId="2B77423D341EFFBBD99AFE636730016D" docLanguage="en" docName="zt02970p062.pdf" docOrigin="Zootaxa 2970" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterobrevennia Kaydan, gen. nov." docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="D74E3A45341AFFBED90DFFF76674037A" masterDocTitle="Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey" masterLastPageNumber="62" masterPageNumber="49" pageNumber="53" updateTime="1698256943753" updateUser="plazi">
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<mods:title id="17DBA1665B8F76223C63610899AB8604">Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey</mods:title>
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<mods:name id="4C95649514ADCA3C4580F552FABAA167" type="personal">
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<mods:namePart id="F063012D6D66DBA42BBF9DE7187AAEA0">Kaydan, Bora</mods:namePart>
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<mods:title id="DB6C108EEFA23D92891496EE29E66D07">Zootaxa</mods:title>
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<mods:date id="9534AE9D40C1CEA5FC9C9AAB55EBF302">2011</mods:date>
<mods:detail id="59C91998AFDB55E3980988D0E151F1B5" type="volume">
<mods:number id="1CF1EAE70C2C317877FE3F527C12FD3E">2970</mods:number>
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<mods:classification id="270D3795894019D0F5116A93BF68F601">journal article</mods:classification>
<mods:identifier id="CA4AFEE15EA7D3F6AB9F334E82F6D5B5" type="DOI">10.5281/zenodo.203960</mods:identifier>
<mods:identifier id="A2C06FA330A700D413C326105013C9D0" type="GBIF-Dataset">24889de9-856a-41ba-a31b-3faf1828f314</mods:identifier>
<mods:identifier id="0DE83E21AEF4E73D6622642D8C47E5E1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="10D1A0C10CD91627A04B9E6D76A8D901" type="Zenodo-Dep">203960</mods:identifier>
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<treatment id="2B77423D341EFFBBD99AFE636730016D" ID-DOI="http://doi.org/10.5281/zenodo.6187396" ID-GBIF-Taxon="119402715" ID-Zenodo-Dep="6187396" LSID="urn:lsid:plazi:treatment:2B77423D341EFFBBD99AFE636730016D" httpUri="http://treatment.plazi.org/id/2B77423D341EFFBBD99AFE636730016D" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
<subSubSection id="EBC4A0A0341EFFBAD99AFE63643D02D7" box="[151,585,403,430]" pageId="4" pageNumber="53" type="nomenclature">
<paragraph id="A361F32B341EFFBAD99AFE63643D02D7" blockId="4.[151,585,403,430]" box="[151,585,403,430]" pageId="4" pageNumber="53">
<heading id="F8294447341EFFBAD99AFE63643D02D7" bold="true" box="[151,585,403,430]" fontSize="11" level="1" pageId="4" pageNumber="53" reason="1">
<emphasis id="91AA2F39341EFFBAD99AFE63643D02D7" bold="true" box="[151,585,403,430]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD99AFE6367BB02D7" authority="Kaydan" authorityName="Kaydan" box="[151,463,403,430]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBAD99AFE63671702D4" bold="true" box="[151,355,404,430]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
Kaydan
</taxonomicName>
,
<taxonomicNameLabel id="8A999242341EFFBAD8D1FE64643D02D7" box="[476,585,403,429]" pageId="4" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EBC4A0A0341EFFBAD99AFE2C643907E6" pageId="4" pageNumber="53" type="materials_examined">
<paragraph id="A361F32B341EFFBAD99AFE2C6576028B" blockId="4.[151,770,474,497]" box="[151,770,474,497]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD99AFE2C6752028B" bold="true" box="[151,294,474,497]" pageId="4" pageNumber="53">
<typeStatus id="7C654D89341EFFBAD99AFE2C66B8028A" box="[151,204,475,496]" pageId="4" pageNumber="53">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="64DE88A8341EFFBAD826FE2D648F028B" authority="Borchsenius, 1949" authorityName="Borchsenius" authorityYear="1949" box="[299,763,474,497]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="opertus">
<emphasis id="91AA2F39341EFFBAD826FE2D6449028B" bold="true" box="[299,573,474,497]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD826FE2D645B028A" bold="true" box="[299,559,474,496]" italics="true" pageId="4" pageNumber="53">Heterobrevennia opertus</emphasis>
(
</emphasis>
Borchsenius, 1949
</taxonomicName>
)
</paragraph>
<paragraph id="A361F32B341EFFBAD99AFDEA6446004E" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD99AFDEA6469014C" bold="true" box="[151,541,541,566]" pageId="4" pageNumber="53">Generic diagnosis. Adult female:</emphasis>
Body elongate,
<quantity id="64265ECE341EFFBADBD8FDEA650C014F" box="[725,888,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="3.01" metricValueMax="3.2" metricValueMin="2.82" pageId="4" pageNumber="53" unit="mm" value="3.01" valueMax="3.2" valueMin="2.82">2.82–3.20 mm</quantity>
long,
<quantity id="64265ECE341EFFBADAB3FDEA6215014F" box="[958,1121,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" metricValueMax="2.2" metricValueMin="1.56" pageId="4" pageNumber="53" unit="mm" value="1.88" valueMax="2.2" valueMin="1.56">1.56–2.20 mm</quantity>
wide. Antennae generally 8 or 9 segmented (but occasionally 7 when 2nd and 3rd segments fused). Eyes present on margin. Labium 3 segmented. Posterior spiracles slightly larger than anterior spiracles. Legs well developed, with translucent pores on hind femur and tibia; claw with a denticle; tarsal digitules setose, claw digitules capitate, broader than tarsal digitules. Both anterior and posterior ostioles well developed. Multilocular disc pores limited to margins of both body surfaces and to posterior abdominal segments. Quinquelocular pores smaller than multilocular pores, scattered over both body surfaces. Trilocular pores present on dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae.
</paragraph>
<paragraph id="A361F32B341EFFBAD9CAFCB766B60092" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD9CAFCB7675B0022" bold="true" box="[199,303,832,856]" pageId="4" pageNumber="53">Dorsum.</emphasis>
Cerarii numbering always 5 or more pairs, situated on posterior abdominal segments and on head. Dorsal body setae spine-like. Multilocular pores situated on body margin. Quinquelocular pores scattered throughout. Trilocular pores scattered throughout, sometimes few, but always present at least on cerarii, ostioles and near spiracle opening. Minute discoidal pores scattered throughout. Oral-collar tubular ducts elongate scattered throughout.
</paragraph>
<paragraph id="A361F32B341EFFBAD9CAFC03643907E6" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD9CAFC03676D0776" bold="true" box="[199,281,1012,1036]" pageId="4" pageNumber="53">Venter.</emphasis>
Body setae slender, longest setae present medially on head; apical setae long; spine-like setae present submarginally on body. Multilocular disc pores present on posterior abdominal segments and on margin of thorax and head. Quinquelocular pores scattered over body. Trilocular pores present around atrium of spiracle, absent elsewhere. Minute discoidal pores scattered over body. Oral-collar tubular ducts in rows on posterior abdominal segment and scattered on thorax and head.
</paragraph>
</subSubSection>
<subSubSection id="EBC4A0A0341EFFBAD9CAFB5067780536" pageId="4" pageNumber="53" type="discussion">
<paragraph id="A361F32B341EFFBAD9CAFB506231062A" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<emphasis id="91AA2F39341EFFBAD9CAFB50673007BA" bold="true" box="[199,324,1191,1216]" pageId="4" pageNumber="53">Comment.</emphasis>
<taxonomicName id="64DE88A8341EFFBAD846FB5F647307BA" box="[331,519,1192,1216]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBAD846FB5F647307BA" box="[331,519,1192,1216]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8A999242341EFFBADB00FB5F640107BA" box="[525,629,1192,1216]" pageId="4" pageNumber="53" rank="genus">
<emphasis id="91AA2F39341EFFBADB00FB5F641A07BA" bold="true" box="[525,622,1192,1216]" pageId="4" pageNumber="53">gen. nov</emphasis>
.
</taxonomicNameLabel>
is most similar to
<taxonomicName id="64DE88A8341EFFBADA4EFB5E658907BA" box="[835,1021,1193,1216]" class="Insecta" family="Pseudococcidae" genus="Heterococcopsis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBADA4EFB5E658907BA" box="[835,1021,1193,1216]" italics="true" pageId="4" pageNumber="53">Heterococcopsis</emphasis>
</taxonomicName>
in having trilocular pores on the dorsum and 8- or 9-segmented antennae. The two genera differ as follows (features of
<taxonomicName id="64DE88A8341EFFBADD5FFB3A6378079E" box="[1106,1292,1229,1252]" class="Insecta" family="Pseudococcidae" genus="Heterococcopsis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBADD5FFB3A6378079E" box="[1106,1292,1229,1252]" italics="true" pageId="4" pageNumber="53">Heterococcopsis</emphasis>
</taxonomicName>
in parentheses): i) no circuli (with 3 circuli), ii) trilocular pores scattered on dorsum (trilocular pores on dorsum restricted to cerarii, ostioles and posterior abdominal segments), iii) at least five cerarii, of which 3 are present on head (only 2 cerarii, restricted to posterior abdominal segments), and iv) legs slender (legs stout).
</paragraph>
<paragraph id="A361F32B341EFFBAD9C8FAAB656506E2" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD9C8FAAB67F5060E" box="[197,385,1372,1396]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBAD9C8FAAB67F5060E" box="[197,385,1372,1396]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
is also similar to all the other genera in the
<taxonomicName id="64DE88A8341EFFBADA80FAAA6252060E" box="[909,1062,1373,1396]" class="Insecta" family="Pseudococcidae" genus="Heterococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBADA80FAAA6252060E" box="[909,1062,1373,1396]" italics="true" pageId="4" pageNumber="53">Heterococcus</emphasis>
</taxonomicName>
-group. The affinities of
<taxonomicName id="64DE88A8341EFFBADC4BFAAA677206E2" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBADC4BFAAA677206E2" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
with other related genera are discussed below.
</paragraph>
<paragraph id="A361F32B341EFFBAD9C8FA5367780536" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD9C8FA5367F306C6" box="[197,391,1444,1468]" class="Insecta" family="Pseudococcidae" genus="Asphodelococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBAD9C8FA5367F306C6" box="[197,391,1444,1468]" italics="true" pageId="4" pageNumber="53">Asphodelococcus</emphasis>
</taxonomicName>
can be readily separated from
<taxonomicName id="64DE88A8341EFFBADBF5FA5365C006C6" box="[760,948,1444,1468]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBADBF5FA5365C006C6" box="[760,948,1444,1468]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
by the following combination of features (features of
<taxonomicName id="64DE88A8341EFFBAD82EFA3F67AB069A" box="[291,479,1480,1504]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBAD82EFA3F67AB069A" box="[291,479,1480,1504]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
in parenthesis): (i) antennae 6 segmented (antennae 9 segmented), (ii) multilocular pores of unusual shape and often quadrate, and with heavily sclerotised hexagonal pores (pores normal), and (iii) cerarii numbering 4 pairs and restricted to abdomen (cerarii numbering 5 or more pairs, present on both abdomen and head).
</paragraph>
</subSubSection>
<subSubSection id="EBC4A0A0341EFFBBD9C8F9AF67F602AA" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53" type="reference_group">
<paragraph id="A361F32B341EFFBAD9C8F9AF6738045E" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD9C8F9AF6716050A" ID-CoL="WMS" box="[197,354,1624,1648]" class="Insecta" family="Pseudococcidae" genus="Annulococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBAD9C8F9AF6716050A" box="[197,354,1624,1648]" italics="true" pageId="4" pageNumber="53">Annulococcus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="64DE88A8341EFFBADB12F9AF64AF050A" box="[543,731,1624,1648]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBADB12F9AF64AF050A" box="[543,731,1624,1648]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
in the following combination of features (features of
<taxonomicName id="64DE88A8341EFFBAD99AF98B672705EE" box="[151,339,1660,1684]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBAD99AF98B672705EE" box="[151,339,1660,1684]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
in parenthesis): (i) cerarii poorly developed, present along the body margin (cerarii numbering 5 or more pairs, present on both abdomen and head), (ii) quinquelocular pores larger than other multilocular pores (quinquelocular pores smaller than other multilocular pores), (iii) oral-collar tubular ducts very short and stout (oral-collar tubular ducts elongate), and (iv) trilocular pores absent (trilocular pores present) (
<bibRefCitation id="C74F8EDA341EFFBADDCFF91F6320047A" author="Miller" box="[1218,1364,1768,1792]" pageId="4" pageNumber="53" refString="Miller, D. R. (1975) A revision of the genus Heterococcus Ferris with a diagnosis of Brevennia Goux (Homoptera: Coccoidea: Pseudococcidae). United States Department of Agriculture Technical Bulletin, 1497, 1 - 61." type="journal article" year="1975">Miller, 1975</bibRefCitation>
;
<bibRefCitation id="C74F8EDA341EFFBADC6CF91F674B045E" author="Foldi" pageId="4" pageNumber="53" refString="Foldi, I. &amp; Cox, J. M. (1989) A new genus and species of mealybug from the Venezuelan Andes, with a guide to the identification of genera in the grass-feeding Heterococcus - group (Homoptera: Pseudococcidae). Journal of Natural History, 23, 1119 - 1131." type="journal article" year="1989">Foldi and Cox, 1989</bibRefCitation>
).
</paragraph>
<paragraph id="A361F32B341EFFBAD9C8F8C6621B04EA" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD9C8F8C667080432" ID-CoL="3D2T" authority="Goux" authorityName="Goux" box="[197,380,1840,1864]" class="Insecta" family="Pseudococcidae" genus="Brevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBAD9C8F8C667430432" box="[197,311,1841,1864]" italics="true" pageId="4" pageNumber="53">Brevennia</emphasis>
Goux
</taxonomicName>
shares many morphological features with
<taxonomicName id="64DE88A8341EFFBADA56F8C762630432" box="[859,1047,1840,1864]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBADA56F8C762630432" box="[859,1047,1840,1864]" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
, namely, the presence of trilocular pores on the dorsum and venter and the abundance of quinquelocular pores on both dorsum and venter, but the two genera differ in the absence of claw denticles and
<taxonomicName id="64DE88A8341EFFBADBCAF88E654D04EA" ID-CoL="3D2T" box="[711,825,1913,1936]" class="Insecta" family="Pseudococcidae" genus="Brevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBADBCAF88E654D04EA" box="[711,825,1913,1936]" italics="true" pageId="4" pageNumber="53">Brevennia</emphasis>
</taxonomicName>
also lacks anterior ostioles.
</paragraph>
<paragraph id="A361F32B341EFFBAD9C8F86A626F04A3" blockId="4.[151,1437,541,2009]" pageId="4" pageNumber="53">
<taxonomicName id="64DE88A8341EFFBAD9C8F86A67D804CE" ID-CoL="3BKG" authority="Danzig" authorityName="Danzig" box="[197,428,1948,1972]" class="Insecta" family="Pseudococcidae" genus="Boreococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBAD9C8F86A672104CE" box="[197,341,1949,1972]" italics="true" pageId="4" pageNumber="53">Boreococcus</emphasis>
Danzig
</taxonomicName>
,
<taxonomicName id="64DE88A8341EFFBAD8B4F86A649004CE" ID-CoL="4WCJ" authority="Borchsenius" authorityName="Borchsenius" box="[441,740,1948,1972]" class="Insecta" family="Pseudococcidae" genus="Heterococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBAD8B4F86A642604CE" box="[441,594,1949,1972]" italics="true" pageId="4" pageNumber="53">Heterococcus</emphasis>
Borchsenius
</taxonomicName>
and
<taxonomicName id="64DE88A8341EFFBADA14F86B65A604CE" ID-CoL="73D4" box="[793,978,1948,1972]" class="Insecta" family="Pseudococcidae" genus="Pseudorhodania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341EFFBADA14F86B65A604CE" box="[793,978,1948,1972]" italics="true" pageId="4" pageNumber="53">Pseudorhodania</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="64DE88A8341EFFBADC5EF86A676004A2" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341EFFBADC5EF86A676004A2" italics="true" pageId="4" pageNumber="53">Heterobrevennia</emphasis>
</taxonomicName>
due to the complete absence of trilocular pores in these three genera.
</paragraph>
<paragraph id="A361F32B341FFFBBD9C8FF6E63310266" blockId="5.[151,1436,151,535]" pageId="5" pageNumber="54">
<emphasis id="91AA2F39341FFFBBD9C8FF6E671503CA" box="[197,353,153,176]" italics="true" pageId="5" pageNumber="54">Laingococcus</emphasis>
Williams differs from
<taxonomicName id="64DE88A8341FFFBBDB53FF6F656E03CA" box="[606,794,152,176]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341FFFBBDB53FF6F656E03CA" box="[606,794,152,176]" italics="true" pageId="5" pageNumber="54">Heterobrevennia</emphasis>
</taxonomicName>
in having the following combination of features (features of
<taxonomicName id="64DE88A8341FFFBBD9B7FF4B670203AE" box="[186,374,188,212]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341FFFBBD9B7FF4B670203AE" box="[186,374,188,212]" italics="true" pageId="5" pageNumber="54">Heterobrevennia</emphasis>
</taxonomicName>
in parenthesis): (i) ostioles absent (ostioles present), (ii) cerarii absent (5 or more cerarii present on both abdomen and head); (iii) a circulus present (absent); (iv) dorsal body setae similar in size to those on venter (dorsal body setae spinose, ventral setose); and (v) in having a rotund body (body elongate oval).
</paragraph>
<paragraph id="A361F32B341FFFBBD9C8FEDF6530021E" blockId="5.[151,1436,151,535]" pageId="5" pageNumber="54">
In having trilocular pores on the dorsum and venter,
<taxonomicName id="64DE88A8341FFFBBDA29FEDF6594023A" box="[804,992,296,320]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341FFFBBDA29FEDF6594023A" box="[804,992,296,320]" italics="true" pageId="5" pageNumber="54">Heterobrevennia</emphasis>
</taxonomicName>
is somewhat similar to
<emphasis id="91AA2F39341FFFBBDDF8FEDE63E8023A" box="[1269,1436,297,320]" italics="true" pageId="5" pageNumber="54">Paramacoccus</emphasis>
Foldi &amp; Cox but
<emphasis id="91AA2F39341FFFBBD85BFEBA6788021E" box="[342,508,333,356]" italics="true" pageId="5" pageNumber="54">Paramacoccus</emphasis>
is unique in lacking ostioles.
</paragraph>
<paragraph id="A361F32B341FFFBBD9C8FE9867F602AA" blockId="5.[151,1436,151,535]" pageId="5" pageNumber="54">
On the other hand, in having trilocular pores on the dorsum and venter, and spine-like setae on dorsum,
<taxonomicName id="64DE88A8341FFFBBDC5EFE86676002D6" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341FFFBBDC5EFE86676002D6" italics="true" pageId="5" pageNumber="54">Heterobrevennia</emphasis>
</taxonomicName>
is somewhat similar to
<taxonomicName id="64DE88A8341FFFBBDB2AFE63654502D6" ID-CoL="6N5N" authority="Cockerell" authorityName="Cockerell" box="[551,817,404,428]" class="Insecta" family="Pseudococcidae" genus="Phenacoccus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341FFFBBDB2AFE6364CF02D6" box="[551,699,404,428]" italics="true" pageId="5" pageNumber="54">Phenacoccus</emphasis>
Cockerell
</taxonomicName>
but new genus differs in having many quinquelocular pores on the dorsum.
</paragraph>
</subSubSection>
<subSubSection id="EBC4A0A0341FFFBBD9CAFE2B6730016D" pageId="5" pageNumber="54" type="etymology">
<paragraph id="A361F32B341FFFBBD9CAFE2B6730016D" blockId="5.[151,1436,151,535]" pageId="5" pageNumber="54">
<emphasis id="91AA2F39341FFFBBD9CAFE2B673A028E" bold="true" box="[199,334,476,500]" pageId="5" pageNumber="54">Etymology.</emphasis>
The name
<taxonomicName id="64DE88A8341FFFBBD8DAFE2B64E7028E" box="[471,659,476,500]" class="Insecta" family="Pseudococcidae" genus="Heterobrevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="91AA2F39341FFFBBD8DAFE2B64E7028E" box="[471,659,476,500]" italics="true" pageId="5" pageNumber="54">Heterobrevennia</emphasis>
</taxonomicName>
refers to its similarity to the genera
<taxonomicName id="64DE88A8341FFFBBDD46FE2A6290028E" box="[1099,1252,477,500]" class="Insecta" family="Pseudococcidae" genus="Heterococcus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341FFFBBDD46FE2A6290028E" box="[1099,1252,477,500]" italics="true" pageId="5" pageNumber="54">Heterococcus</emphasis>
</taxonomicName>
and
<taxonomicName id="64DE88A8341FFFBBDC2EFE2A63E1028E" box="[1315,1429,477,500]" class="Insecta" family="Pseudococcidae" genus="Brevennia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="91AA2F39341FFFBBDC2EFE2A63E1028E" box="[1315,1429,477,500]" italics="true" pageId="5" pageNumber="54">Brevennia</emphasis>
</taxonomicName>
. Gender female.
</paragraph>
</subSubSection>
</treatment>
</document>
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