992 lines
109 KiB
XML
992 lines
109 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4764829" ID-GBIF-Dataset="8b5afb3d-92bd-4e36-8c86-675537e88d70" ID-GBIF-Taxon="182137364" ID-ISSN="1854-0392" ID-Zenodo-Dep="4764829" checkinTime="1620922906122" checkinUser="felipe" docAuthor="Nelson, C. Riley & Baumann, Richard W." docDate="2009" docId="2B2C6D48DD2BFFCEFC0D2014FEEAFE25" docLanguage="en" docName="Illiesia.5.18.188-194.pdf" docOrigin="Illiesia 5 (18)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Capnia shasta Nelson & Baumann 2009, sp. n." docType="treatment" docVersion="4" lastPageId="6" lastPageNumber="194" masterDocId="D7151530DD2AFFC8FFFA2549FF9AFF94" masterDocTitle="Capnia Shasta, A New Species In The Californica Group From Northwestern California (Plecoptera, Capniidae)" masterLastPageNumber="194" masterPageNumber="188" pageId="1" pageNumber="189" updateTime="1636507396062" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Capnia Shasta, A New Species In The Californica Group From Northwestern California (Plecoptera, Capniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nelson, C. Riley</mods:namePart>
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<mods:affiliation>Department of Biology, Brigham Young University, Provo, Utah 84602 E-mail: rileynelson @ byu. edu</mods:affiliation>
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<mods:nameIdentifier type="email">rileynelson@byu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baumann, Richard W.</mods:namePart>
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<mods:affiliation>Department of Biology, Brigham Young University, Provo, Utah 84602 E-mail: rileynelson @ byu. edu & Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
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<mods:nameIdentifier type="email">rileynelson@byu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>Illiesia</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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||
</mods:detail>
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<mods:detail type="issue">
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<mods:number>18</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>188</mods:start>
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<mods:end>194</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758162</mods:identifier>
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<mods:identifier type="GBIF-Dataset">8b5afb3d-92bd-4e36-8c86-675537e88d70</mods:identifier>
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<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4758162</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4764829" ID-GBIF-Taxon="182137364" ID-Zenodo-Dep="4764829" LSID="urn:lsid:plazi:treatment:2B2C6D48DD2BFFCEFC0D2014FEEAFE25" httpUri="http://treatment.plazi.org/id/2B2C6D48DD2BFFCEFC0D2014FEEAFE25" lastPageId="6" lastPageNumber="194" pageId="1" pageNumber="189">
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<subSubSection box="[1015,1264,1372,1397]" pageId="1" pageNumber="189" type="nomenclature">
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||
<paragraph blockId="1.[1015,1264,1372,1397]" box="[1015,1264,1372,1397]" pageId="1" pageNumber="189">
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||
<heading bold="true" box="[1015,1264,1372,1397]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="189" reason="2">
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||
<emphasis bold="true" box="[1015,1264,1372,1397]" pageId="1" pageNumber="189">
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<taxonomicName authority="Nelson & Baumann, 2009" authorityName="Nelson & Baumann" authorityYear="2009" box="[1015,1185,1372,1397]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="189" phylum="Arthropoda" rank="species" species="shasta" status="sp. n.">
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<emphasis bold="true" box="[1015,1185,1372,1397]" italics="true" pageId="1" pageNumber="189">Capnia shasta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1199,1264,1373,1397]" pageId="1" pageNumber="189" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[1081,1197,1409,1433]" pageId="1" pageNumber="189" type="description">
|
||
<paragraph blockId="1.[1081,1197,1409,1433]" box="[1081,1197,1409,1433]" pageId="1" pageNumber="189">
|
||
(
|
||
<figureCitation box="[1089,1189,1409,1433]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="2.[189,238,1759,1783]" captionStartId-1="3.[189,230,1808,1832]" captionTargetBox-0="[287,1323,258,1668]" captionTargetBox-1="[229,724,1075,1766]" captionTargetId-0="figure-53@2.[287,1345,258,1668]" captionTargetId-1="figure-246@3.[229,758,1075,1766]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." captionText-1="Fig. 9. Dissecting stereomicrograph of Capnia shasta, Allotype. Ventral view of female terminalia, posterior end up." figureDoi-0="http://doi.org/10.5281/zenodo.4758165" figureDoi-1="http://doi.org/10.5281/zenodo.4758167" httpUri-0="https://zenodo.org/record/4758165/files/figure.png" httpUri-1="https://zenodo.org/record/4758167/files/figure.png" pageId="1" pageNumber="189">Figs. 1-9</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="189" type="reference_group">
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<paragraph blockId="1.[873,1443,1480,1530]" pageId="1" pageNumber="189">
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<treatmentCitationGroup pageId="1" pageNumber="189">
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<treatmentCitation author="Nelson, C. R. & R. W. Baumann" box="[873,1328,1480,1502]" page="306" pageId="1" pageNumber="189" year="1989">
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<taxonomicName authority=", Nelson & Baumann 1989: 306" authorityName="Nelson & Baumann" authorityPageNumber="306" authorityYear="1989" box="[873,1328,1480,1502]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="189" phylum="Arthropoda" rank="species" species="umpqua">
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<emphasis box="[873,1028,1480,1502]" italics="true" pageId="1" pageNumber="189">Capnia umpqua</emphasis>
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,
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<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[1039,1328,1480,1502]" pageId="1" pageNumber="189" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann 1989:306</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, Figs. 177
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<emphasis box="[1431,1443,1483,1501]" italics="true" pageId="1" pageNumber="189">–</emphasis>
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180, 252, map Fig. 263 (Not
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<taxonomicName authority="Frison 1942" authorityName="Frison" authorityYear="1942" box="[1154,1434,1508,1530]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="189" phylum="Arthropoda" rank="species" species="umpqua">
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<emphasis box="[1154,1309,1508,1530]" italics="true" pageId="1" pageNumber="189">Capnia umpqua</emphasis>
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<bibRefCitation author="Frison, T. H." box="[1316,1434,1508,1530]" pageId="1" pageNumber="189" pagination="61 - 73" refId="ref3882" refString="Frison, T. H. 1942. Descriptions, records and systematic notes concerning western North American stoneflies (Plecoptera). Pan-Pacific Entomologist, 18: 61 - 73." type="journal article" year="1942">Frison 1942</bibRefCitation>
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</taxonomicName>
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)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="191" pageId="1" pageNumber="189" type="materials_examined">
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<paragraph blockId="1.[835,1443,1578,1923]" lastBlockId="3.[189,798,232,1038]" lastPageId="3" lastPageNumber="191" pageId="1" pageNumber="189">
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||
<emphasis bold="true" box="[835,1070,1579,1603]" pageId="1" pageNumber="189">Material examined.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3111871301" collectingDate="1985-02-16" collectingDateMax="2009-06-17" collectingDateMin="1985-02-16" collectorName="R. W. Baumann & C. R. Nelson" country="United States of America" county="Shasta Co." elevation="708" latitude="41.15587" location="Castle Crags State Park" longLatPrecision="1" longitude="-122.36285" municipality="Sulphur Creek" pageId="1" pageNumber="189" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="California" typeStatus="holotype">
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<specimenCount box="[1075,1095,1578,1603]" pageId="1" pageNumber="189" type="male">♂</specimenCount>
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<typeStatus box="[1099,1203,1579,1603]" pageId="1" pageNumber="189">holotype</typeStatus>
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, female allotype, and
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<specimenCount box="[835,918,1615,1639]" pageId="1" pageNumber="189" type="male">2 male</specimenCount>
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and
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<specimenCount box="[989,1095,1615,1639]" pageId="1" pageNumber="189" type="female">4 female</specimenCount>
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paratypes,
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<collectingCountry box="[1243,1301,1615,1639]" name="United States of America" pageId="1" pageNumber="189">USA</collectingCountry>
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,
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<collectingRegion box="[1317,1438,1615,1639]" country="United States of America" name="California" pageId="1" pageNumber="189">California</collectingRegion>
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,
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<collectingCounty box="[835,960,1651,1675]" pageId="1" pageNumber="189">Shasta Co.</collectingCounty>
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,
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<collectingMunicipality box="[973,1148,1651,1675]" pageId="1" pageNumber="189">Sulphur Creek</collectingMunicipality>
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||
,
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<location LSID="urn:lsid:plazi:treatment:2B2C6D48DD2BFFCEFC0D2014FEEAFE25:A65A8A85DD2BFFC9FB72233AFA04F91F" box="[1160,1438,1651,1675]" country="United States of America" county="Shasta Co." latitude="41.15587" longLatPrecision="1" longitude="-122.36285" municipality="Sulphur Creek" name="Castle Crags State Park" pageId="1" pageNumber="189" stateProvince="California">Castle Crags State Park</location>
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,
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<date box="[835,1035,1686,1710]" pageId="1" pageNumber="189" value="1985-02-16">
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||
<collectingDate box="[835,1035,1686,1710]" pageId="1" pageNumber="189" value="1985-02-16">16 February 1985</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[1045,1220,1687,1710]" pageId="1" pageNumber="189">R.W. Baumann</collectorName>
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&
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<collectorName box="[1251,1391,1686,1710]" pageId="1" pageNumber="189">C.R. Nelson</collectorName>
|
||
. The locality was checked with Google Earth internet software on
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<date box="[979,1125,1757,1781]" pageId="1" pageNumber="189" value="2009-06-17">
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||
<collectingDate box="[979,1125,1757,1781]" pageId="1" pageNumber="189" value="2009-06-17">17 June 2009</collectingDate>
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</date>
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and found the location just outside of the park at:
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<geoCoordinate box="[1092,1227,1792,1816]" degrees="41.15587" direction="north" orientation="latitude" pageId="1" pageNumber="189" precision="1" value="41.15587">N 41.15587°</geoCoordinate>
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||
<geoCoordinate box="[1232,1385,1792,1816]" degrees="122.36285" direction="west" orientation="longitude" pageId="1" pageNumber="189" precision="1" value="-122.36285">W 122.36285°</geoCoordinate>
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||
elev.
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||
<quantity box="[835,907,1828,1852]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="1" pageNumber="189" unit="m" value="708.0">
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||
<elevation box="[835,907,1828,1852]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="1" pageNumber="189" unit="m" value="708.0">708 m</elevation>
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</quantity>
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||
</materialsCitation>
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||
. Each primary type specimen has been placed in a separate vial, in 70% ethanol. One
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<typeStatus box="[1277,1382,1864,1888]" pageId="1" pageNumber="189">paratype</typeStatus>
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male and the
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<typeStatus box="[961,1057,1899,1923]" pageId="1" pageNumber="189">allotype</typeStatus>
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female were the specimens illustrated in
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<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[344,644,232,256]" pageId="3" pageNumber="191" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann (1989)</bibRefCitation>
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as
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<taxonomicName authorityName="Frison" authorityYear="1942" box="[677,791,232,256]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="umpqua">
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<emphasis box="[677,791,232,256]" italics="true" pageId="3" pageNumber="191">C. umpqua</emphasis>
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</taxonomicName>
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.
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<collectorName box="[189,219,267,291]" pageId="3" pageNumber="191">Of</collectorName>
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the paraptypes,
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<specimenCount box="[411,516,267,291]" pageId="3" pageNumber="191" type="male">one male</specimenCount>
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and
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<specimenCount box="[571,727,267,291]" pageId="3" pageNumber="191" type="female">three females</specimenCount>
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are in 70% ethanol and the terminalia of
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<specimenCount box="[602,692,303,327]" pageId="3" pageNumber="191" type="male">2 males</specimenCount>
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and
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<specimenCount pageId="3" pageNumber="191" type="female">one female</specimenCount>
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are dried and on SEM stubs, including the specimens used for
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<figureCitation box="[418,514,374,398]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs. 1-8</figureCitation>
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(the remainders of these specimens are in 70% ethanol vials).
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<collectingCounty box="[638,682,409,433]" pageId="3" pageNumber="191">The</collectingCounty>
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<typeStatus box="[692,797,409,433]" pageId="3" pageNumber="191">holotype</typeStatus>
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and
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<typeStatus box="[252,348,445,469]" pageId="3" pageNumber="191">allotype</typeStatus>
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||
are deposited at the
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<collectingMunicipality pageId="3" pageNumber="191">Smithsonian Institution</collectingMunicipality>
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,
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<collectingCountry box="[348,526,480,505]" name="United States of America" pageId="3" pageNumber="191">United States</collectingCountry>
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||
<location LSID="urn:lsid:plazi:treatment:2B2C6D48DD2BFFCEFC0D2014FEEAFE25:A65A8A85DD29FFCBFDD024A9FC82FE6C" box="[554,792,480,504]" country="United States of America" county="The" municipality="Smithsonian Institution" name="National Museum" pageId="3" pageNumber="191" stateProvince="Washington">National Museum</location>
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,
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<collectingRegion box="[189,334,516,540]" country="United States of America" name="Washington" pageId="3" pageNumber="191">Washington</collectingRegion>
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, D.C. (
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<collectionCode box="[410,497,516,540]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="191" type="Museum">USNM</collectionCode>
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||
).
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||
<location LSID="urn:lsid:plazi:treatment:2B2C6D48DD2BFFCEFC0D2014FEEAFE25:A65A8A85DD29FFCBFDFB274DFDBFFD88" box="[513,549,516,540]" country="United States of America" county="The" municipality="Smithsonian Institution" name="All" pageId="3" pageNumber="191" stateProvince="Washington">All</location>
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||
<typeStatus box="[554,667,516,540]" pageId="3" pageNumber="191">paratypes</typeStatus>
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||
are housed at the
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||
<collectionCode box="[278,299,552,575]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="191">R</collectionCode>
|
||
.
|
||
<collectorName box="[302,792,551,575]" pageId="3" pageNumber="191">W. Baumann Aquatic Insect Collection</collectorName>
|
||
,
|
||
<collectorName box="[189,607,587,611]" pageId="3" pageNumber="191">Monte L. Bean Life Science Museum</collectorName>
|
||
, Brigham Young University, Provo,
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||
<collectingRegion box="[411,469,622,646]" country="United States of America" name="Utah" pageId="3" pageNumber="191">Utah</collectingRegion>
|
||
(
|
||
<collectionCode box="[484,564,623,647]" pageId="3" pageNumber="191">BYUC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758165" ID-Zenodo-Dep="4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="2" pageNumber="190" startId="2.[189,238,1759,1783]" targetBox="[287,1323,258,1668]" targetPageId="2">
|
||
<paragraph blockId="2.[189,1444,1759,1926]" pageId="2" pageNumber="190">
|
||
Figs. 1-8.
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[291,432,1759,1783]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="190" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[291,432,1759,1783]" italics="true" pageId="2" pageNumber="190">Capnia shasta</emphasis>
|
||
</taxonomicName>
|
||
, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale
|
||
<quantity box="[785,865,1795,1819]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="190" unit="mm" value="100.0">100µm</quantity>
|
||
. 3. male epiproct, dorsal, anterior to right, 387x, scale
|
||
<quantity box="[189,256,1830,1855]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="190" unit="mm" value="50.0">50µm</quantity>
|
||
. 4. male epiproct, lateral, anterior to right, 389x, scale
|
||
<quantity box="[861,929,1830,1855]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="190" unit="mm" value="50.0">50µm</quantity>
|
||
. 5. details of male epiproct, dorsal, anterior to right, 868x, scale
|
||
<quantity box="[379,446,1866,1890]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="190" unit="mm" value="20.0">20µm</quantity>
|
||
. 6. details of male epiproct, lateral, anterior to right,
|
||
<date box="[1025,1091,1866,1890]" pageId="2" pageNumber="190" value="1959-10">1959x</date>
|
||
, scale
|
||
<quantity box="[1160,1228,1866,1890]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="190" unit="mm" value="10.0">10µm</quantity>
|
||
. 7. male terminalia, dorsal, anterior down, 157x, scale
|
||
<quantity box="[567,648,1902,1926]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="190" unit="mm" value="100.0">100µm</quantity>
|
||
. 8. female terminalia, ventral, anterior down, 108x, scale
|
||
<quantity box="[1277,1358,1902,1926]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="190" unit="mm" value="200.0">200µm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="3" pageNumber="191" type="description">
|
||
<paragraph blockId="3.[189,798,232,1038]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[189,258,658,682]" pageId="3" pageNumber="191">Male.</emphasis>
|
||
Body length
|
||
<quantity box="[410,512,658,682]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="191" unit="mm" value="5.1">5.10 mm</quantity>
|
||
; interocular distance
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="3" pageNumber="191" unit="mm" value="0.57">0.57 mm</quantity>
|
||
; wings macropterous, forewing length
|
||
<quantity box="[703,792,694,718]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="191" unit="mm" value="5.9">5.9 mm</quantity>
|
||
; tergum 9 divided along one-half its length by medial membranous area, anterior margin heavily sclerotized. Well-developed tergal knobs present along inner margin of the divided tergum nine (
|
||
<figureCitation box="[198,292,872,896]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs.1-4</figureCitation>
|
||
), no tergal knobs on segment eight; epiproct broad, length
|
||
<quantity box="[377,464,907,931]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="3" pageNumber="191" unit="m" value="285.0">285 m</quantity>
|
||
, epiproct width
|
||
<quantity box="[693,792,907,931]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" pageId="3" pageNumber="191" unit="m" value="132.0">132 m</quantity>
|
||
, measured in dorsal view (
|
||
<figureCitation box="[517,631,943,967]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs. 1, 3</figureCitation>
|
||
) with only a single process well-developed (
|
||
<figureCitation box="[572,684,978,1002]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs. 2, 4</figureCitation>
|
||
); apex of epiproct unforked (
|
||
<figureCitation box="[418,550,1013,1038]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs. 1, 3, 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[835,1443,232,1536]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[835,929,232,256]" pageId="3" pageNumber="191">Female.</emphasis>
|
||
Body length
|
||
<quantity box="[1091,1195,232,256]" metricMagnitude="-3" metricUnit="m" metricValue="6.48" pageId="3" pageNumber="191" unit="mm" value="6.48">6.48 mm</quantity>
|
||
; interocular distance
|
||
<quantity box="[835,937,268,292]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="191" unit="mm" value="0.62">0.62 mm</quantity>
|
||
; wings macropterous, forewing length 6.80 mm; subgenital plate triangular with apex directed anteriorly (
|
||
<figureCitation box="[964,1041,339,363]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="3" pageNumber="191">Figs. 8</figureCitation>
|
||
,
|
||
<figureCitation box="[1051,1066,339,363]" captionStart="Fig" captionStartId="3.[189,230,1808,1832]" captionTargetBox="[229,724,1075,1766]" captionTargetId="figure-246@3.[229,758,1075,1766]" captionTargetPageId="3" captionText="Fig. 9. Dissecting stereomicrograph of Capnia shasta, Allotype. Ventral view of female terminalia, posterior end up." figureDoi="http://doi.org/10.5281/zenodo.4758167" httpUri="https://zenodo.org/record/4758167/files/figure.png" pageId="3" pageNumber="191">9</figureCitation>
|
||
); subgenital plate width
|
||
<quantity box="[1346,1443,339,363]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="191" unit="mm" value="0.46">0.46 mm</quantity>
|
||
on hind margin; hind margin straight with margin thickened and not overlapping sternite 9; subgenital plate with paired lateral light thinnings of sclerotization near distal corners, immediately anterior to the hind margin, darker lines (probably internal) running much of the length of subgenital plate near medial line (
|
||
<figureCitation box="[1106,1208,587,611]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="2.[189,238,1759,1783]" captionStartId-1="3.[189,230,1808,1832]" captionTargetBox-0="[287,1323,258,1668]" captionTargetBox-1="[229,724,1075,1766]" captionTargetId-0="figure-53@2.[287,1345,258,1668]" captionTargetId-1="figure-246@3.[229,758,1075,1766]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." captionText-1="Fig. 9. Dissecting stereomicrograph of Capnia shasta, Allotype. Ventral view of female terminalia, posterior end up." figureDoi-0="http://doi.org/10.5281/zenodo.4758165" figureDoi-1="http://doi.org/10.5281/zenodo.4758167" httpUri-0="https://zenodo.org/record/4758165/files/figure.png" httpUri-1="https://zenodo.org/record/4758167/files/figure.png" pageId="3" pageNumber="191">Figs. 8-9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="191" type="etymology">
|
||
<paragraph blockId="3.[835,1443,232,1536]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[835,973,623,647]" pageId="3" pageNumber="191">Etymology.</emphasis>
|
||
This species is named for prominent Mount Shasta of the Cascade Range in northern
|
||
<collectingRegion box="[835,954,694,718]" country="United States of America" name="California" pageId="3" pageNumber="191">California</collectingRegion>
|
||
as a noun, without gender, in apposition. It shares this name with a common, somewhat generic brand of soda water common in the western
|
||
<collectingCountry box="[835,996,801,825]" name="United States of America" pageId="3" pageNumber="191">United States</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="191" type="diagnosis">
|
||
<paragraph blockId="3.[835,1443,232,1536]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[835,967,836,860]" pageId="3" pageNumber="191">Diagnosis.</emphasis>
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[977,1132,836,860]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[977,1132,836,860]" italics="true" pageId="3" pageNumber="191">Capnia shasta</emphasis>
|
||
</taxonomicName>
|
||
is unique in that it is the only member of the Californica Group with tergal knobs only on segment 9 and no distinctively divided process on the epiproct. It lacks the distinct upper process of the epiproct found in the Ventura Subgroup. It differs from the Californica Subgroup in lacking knobs on tergum 8. Phylogenetic relationships of
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1032,1133,1085,1109]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1032,1133,1085,1109]" italics="true" pageId="3" pageNumber="191">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
with all known members of the Californica Group are summarized in
|
||
<figureCitation box="[1359,1438,1121,1145]" captionStart="Fig" captionStartId="4.[189,230,1866,1890]" captionTargetBox="[354,1279,1041,1807]" captionTargetId="figure-53@4.[325,1308,1004,1844]" captionTargetPageId="4" captionText="Fig. 10. Cladogram summarizing phylogenetic relationships among members of the Californica Group and two outgroups." figureDoi="http://doi.org/10.5281/zenodo.4758169" httpUri="https://zenodo.org/record/4758169/files/figure.png" pageId="3" pageNumber="191">Fig. 10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="191" type="discussion">
|
||
<paragraph blockId="3.[835,1443,232,1536]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[835,980,1156,1180]" pageId="3" pageNumber="191">Discussion.</emphasis>
|
||
<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[990,1316,1156,1181]" pageId="3" pageNumber="191" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann (1989)</bibRefCitation>
|
||
predicted that the area in
|
||
<collectingRegion box="[1032,1154,1192,1216]" country="United States of America" name="California" pageId="3" pageNumber="191">California</collectingRegion>
|
||
between Santa Barbara and Carmel would probably be a site for a new species or two yet to be discovered in the Californica Group. To this date, no one has reported any new records of
|
||
<taxonomicName box="[1200,1374,1334,1358]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[1200,1278,1335,1358]" italics="true" pageId="3" pageNumber="191">Capnia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from this area but in this paper we describe
|
||
<emphasis box="[1332,1442,1370,1394]" italics="true" pageId="3" pageNumber="191">
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1332,1439,1370,1394]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="shasta">C. shasta</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from a more northerly thought-to-be-well-collected and central portion of the overall group range. The challenge remains to find coastal new members of the group in northern
|
||
<collectingRegion box="[1104,1229,1512,1536]" country="United States of America" name="California" pageId="3" pageNumber="191">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="191" type="multiple">
|
||
<paragraph blockId="3.[835,1443,1583,1891]" pageId="3" pageNumber="191">
|
||
<emphasis bold="true" box="[835,1006,1583,1607]" pageId="3" pageNumber="191">Key to males.</emphasis>
|
||
All males of the previously known species in the Californica Group except true
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis italics="true" pageId="3" pageNumber="191">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
can be identified using the key (p. 304) of
|
||
<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[835,1164,1690,1714]" pageId="3" pageNumber="191" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann (1989)</bibRefCitation>
|
||
. The following key is a modification of the 1989 key written to accommodate both true
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1140,1266,1761,1785]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[1140,1266,1761,1785]" italics="true" pageId="3" pageNumber="191">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1330,1434,1761,1785]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1330,1434,1761,1785]" italics="true" pageId="3" pageNumber="191">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
. Couplet numbering follows that key, with couplet 8 added and figures from various sources noted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758167" ID-Zenodo-Dep="4758167" httpUri="https://zenodo.org/record/4758167/files/figure.png" pageId="3" pageNumber="191" startId="3.[189,230,1808,1832]" targetBox="[229,724,1075,1766]" targetPageId="3">
|
||
<paragraph blockId="3.[189,797,1808,1903]" pageId="3" pageNumber="191">
|
||
Fig. 9. Dissecting stereomicrograph of
|
||
<taxonomicName authority=", Allotype. Ventral" authorityName="Allotype. Ventral" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[641,797,1808,1832]" italics="true" pageId="3" pageNumber="191">Capnia shasta,</emphasis>
|
||
Allotype. Ventral
|
||
</taxonomicName>
|
||
view of female terminalia, posterior end up.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="4" pageNumber="192" type="key">
|
||
<key pageId="4" pageNumber="192">
|
||
<keyStep pageId="4" pageNumber="192">
|
||
<paragraph blockId="4.[189,798,232,646]" pageId="4" pageNumber="192">
|
||
<keyLead pageId="4" pageNumber="192">
|
||
7(4) Terga 5 and 6 each bearing paired knobs (Figs. 69 and 70 of
|
||
<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[461,789,267,291]" pageId="4" pageNumber="192" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann 1989</bibRefCitation>
|
||
) ……………………………………
|
||
<taxonomicName authority="Frison" authorityName="Frison" authorityYear="1942" box="[643,797,303,327]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[643,715,304,327]" italics="true" pageId="4" pageNumber="192">jewetti</emphasis>
|
||
Frison
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[189,798,232,646]" pageId="4" pageNumber="192">
|
||
<keyLead pageId="4" pageNumber="192">
|
||
7’ Terga 8 or 9 bearing paired knobs (
|
||
<figureCitation captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="4" pageNumber="192">
|
||
Figs. 1-4 and
|
||
<quantity box="[317,330,374,397]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="192" unit="in" value="7.0">7</quantity>
|
||
</figureCitation>
|
||
in this publication and Figs. 177-178 of
|
||
<bibRefCitation author="Nelson, C. R. & R. W. Baumann" box="[265,553,409,434]" pageId="4" pageNumber="192" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann 1989</bibRefCitation>
|
||
) ………………….... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="192">
|
||
<paragraph blockId="4.[189,798,232,646]" pageId="4" pageNumber="192">
|
||
<keyLead pageId="4" pageNumber="192">
|
||
8(7) Terga 8 bearing paired knobs (
|
||
<figureCitation box="[656,760,445,469]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="4" pageNumber="192">
|
||
Figs.
|
||
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|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation author="Baumann, R. W. & K. W. Stewart" box="[265,558,481,505]" pageId="4" pageNumber="192" pagination="34 - 39" refId="ref3841" refString="Baumann, R. W. & K. W. Stewart. 2009. What is Capnia umpqua Frison? (Plecoptera: Capniidae), distribution and variation of terminalia. Illiesia, 5: 34 - 39." type="journal article" year="2009">Baumann & Stewart 2009</bibRefCitation>
|
||
) …….
|
||
<taxonomicName authority="Frison" authorityName="Frison" authorityYear="1942" box="[634,798,480,504]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[634,720,481,504]" italics="true" pageId="4" pageNumber="192">umpqua</emphasis>
|
||
Frison
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[189,798,232,646]" pageId="4" pageNumber="192">
|
||
<keyLead pageId="4" pageNumber="192">
|
||
8’ Terga 9 bearing paired knobs (
|
||
<figureCitation box="[620,718,516,540]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="4" pageNumber="192">
|
||
Figs.
|
||
<quantity box="[682,718,516,540]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="4" pageNumber="192" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1-4</quantity>
|
||
</figureCitation>
|
||
in this publication; Figs.
|
||
<quantity box="[508,644,551,576]" metricMagnitude="0" metricUnit="m" metricValue="4.5339" metricValueMax="4.572" metricValueMin="4.4958" pageId="4" pageNumber="192" unit="in" value="178.5" valueMax="180.0" valueMin="177.0">177-180 in</quantity>
|
||
<bibRefCitation author="Nelson, C. R. & R. W. Baumann" pageId="4" pageNumber="192" pagination="289 - 363" refId="ref4002" refString="Nelson, C. R. & R. W. Baumann. 1989. Systematics and zoogeography of the winter stonefly genus Capnia (Plecoptera: Capniidae) in North America. Great Basin Naturalist, 49: 289 - 363 + index." type="journal article" year="1989">Nelson & Baumann 1989</bibRefCitation>
|
||
; and
|
||
<figureCitation box="[509,609,587,611]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="4" pageNumber="192">
|
||
Figs.
|
||
<quantity box="[573,609,587,611]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="4" pageNumber="192" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3-4</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation author="Nelson, C. R." box="[651,797,587,611]" pageId="4" pageNumber="192" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
……………………………..
|
||
<taxonomicName authority="Nelson & Baumann, 2009" authorityName="Nelson & Baumann" authorityYear="2009" box="[577,643,622,646]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta" status="new species">
|
||
<emphasis box="[577,643,622,646]" italics="true" pageId="4" pageNumber="192">shasta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[657,797,622,646]" pageId="4" pageNumber="192" rank="species">new species</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="194" pageId="4" pageNumber="192" type="discussion">
|
||
<paragraph blockId="4.[189,798,694,967]" lastBlockId="4.[835,1444,232,966]" pageId="4" pageNumber="192">
|
||
<emphasis bold="true" box="[189,320,694,718]" pageId="4" pageNumber="192">Phylogeny.</emphasis>
|
||
A purported phylogeny for the entire Californica Group was produced in (
|
||
<bibRefCitation author="Nelson, C. R." box="[611,755,729,754]" pageId="4" pageNumber="192" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
). In the 2004 paper, the species illustrated (and used for the phylogenetic analysis) as
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[472,584,801,824]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[472,584,801,824]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
was the species we herein describe as
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[408,504,836,860]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[408,504,836,860]" italics="true" pageId="4" pageNumber="192">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<tableCitation box="[521,606,836,860]" captionStart="Table 1" captionStartId="5.[189,254,232,256]" captionTargetBox="[227,1428,872,1334]" captionTargetPageId="5" captionText="Table 1. Matrix of characters used in the parsimony analysis of the Californica Group, with Capnia vernalis and C. nana as outgroups. Character states updated from Nelson (2004), with the correction of data related to C. umpqua and addition of data for C. shasta. State of character 14 is corrected for C. regilla from 0 in Nelson (2004) to 1. All character state texts given are for the “1” state. Question marks (?) indicate missing data. Characters: 1. Knobs present on tergite 8; 2. Knobs present on tergite 9; 3. Lower process of epiproct forked. 4. Female subgenital plate overlapping posterior margin of sternum 8. 5. Upper process of epiproct absent (or very greatly reduced) apically. 6. Upper process of epiproct divided in lateral view. 7. Declivity between processes of epiproct vertical. 8. Lower process of epiproct longer than upper process. 9. Lower process of epiproct much longer than upper process. 10. Knobs present on tergites 5 and 6. 11. Both processes of epiproct of about the same length. 12. Lower process of epiproct widely forked in dorsal view. 13. Declivity between processes of epiproct medially carinate. 14. Declivity between upper and lower processes of epiproct with an extra medial process. 15. Knobs on tergite 8 angulate. 16. Notch between knobs of tergite 8 broad in dorsal view. 17. Notch between upper and lower processes deeply incised in lateral view. 18. Lower process of epiproct narrow in dorsal view. 19. Upper and lower processes of epiproct forming a 90 degree angle in lateral view. 20. Lower process of epiproct with lower surface bent at mid length in lateral view. 21. Lower process of epiproct constricted near apex in dorsal view. 22. Lower process of epiproct evenly tapered, not inflated at midlength in dorsal view." httpUri="http://table.plazi.org/id/F7FA8CD6DD2FFFCDFF4725A1FE77FCAC" pageId="4" pageNumber="192" tableUuid="F7FA8CD6DD2FFFCDFF4725A1FE77FCAC">Table 1</tableCitation>
|
||
summarizes the morphological characters used in the 2004 paper with addition of character states observed from the
|
||
<typeStatus box="[697,797,907,931]" pageId="4" pageNumber="192">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[218,332,943,966]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[218,332,943,966]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
, a minor reinterpretation of Character 14, and a correction of species name for the Sulphur Creek, Castle Crags State Park specimens (
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1281,1406,268,291]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[1281,1406,268,291]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<bibRefCitation author="Nelson, C. R." box="[835,979,303,327]" pageId="4" pageNumber="192" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
) as
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1022,1117,303,327]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1022,1117,303,327]" italics="true" pageId="4" pageNumber="192">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
. We ran a branch and bound analysis using this corrected matrix (
|
||
<tableCitation box="[1279,1367,338,362]" captionStart="Table 1" captionStartId="5.[189,254,232,256]" captionTargetBox="[227,1428,872,1334]" captionTargetPageId="5" captionText="Table 1. Matrix of characters used in the parsimony analysis of the Californica Group, with Capnia vernalis and C. nana as outgroups. Character states updated from Nelson (2004), with the correction of data related to C. umpqua and addition of data for C. shasta. State of character 14 is corrected for C. regilla from 0 in Nelson (2004) to 1. All character state texts given are for the “1” state. Question marks (?) indicate missing data. Characters: 1. Knobs present on tergite 8; 2. Knobs present on tergite 9; 3. Lower process of epiproct forked. 4. Female subgenital plate overlapping posterior margin of sternum 8. 5. Upper process of epiproct absent (or very greatly reduced) apically. 6. Upper process of epiproct divided in lateral view. 7. Declivity between processes of epiproct vertical. 8. Lower process of epiproct longer than upper process. 9. Lower process of epiproct much longer than upper process. 10. Knobs present on tergites 5 and 6. 11. Both processes of epiproct of about the same length. 12. Lower process of epiproct widely forked in dorsal view. 13. Declivity between processes of epiproct medially carinate. 14. Declivity between upper and lower processes of epiproct with an extra medial process. 15. Knobs on tergite 8 angulate. 16. Notch between knobs of tergite 8 broad in dorsal view. 17. Notch between upper and lower processes deeply incised in lateral view. 18. Lower process of epiproct narrow in dorsal view. 19. Upper and lower processes of epiproct forming a 90 degree angle in lateral view. 20. Lower process of epiproct with lower surface bent at mid length in lateral view. 21. Lower process of epiproct constricted near apex in dorsal view. 22. Lower process of epiproct evenly tapered, not inflated at midlength in dorsal view." httpUri="http://table.plazi.org/id/F7FA8CD6DD2FFFCDFF4725A1FE77FCAC" pageId="4" pageNumber="192" tableUuid="F7FA8CD6DD2FFFCDFF4725A1FE77FCAC">Table 1</tableCitation>
|
||
). This analysis yielded six equally parsimonious trees. In all of these trees, the topology conformed largely to that of the tree in
|
||
<bibRefCitation author="Nelson, C. R." box="[993,1155,445,469]" pageId="4" pageNumber="192" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson (2004)</bibRefCitation>
|
||
except that now
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[1359,1443,445,469]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis box="[1359,1443,445,469]" italics="true" pageId="4" pageNumber="192">C. nana</emphasis>
|
||
</taxonomicName>
|
||
falls into a polytomy with
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1176,1278,480,504]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1176,1278,480,504]" italics="true" pageId="4" pageNumber="192">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
and the two subgroups (
|
||
<figureCitation box="[973,1053,516,540]" captionStart="Fig" captionStartId="4.[189,230,1866,1890]" captionTargetBox="[354,1279,1041,1807]" captionTargetId="figure-53@4.[325,1308,1004,1844]" captionTargetPageId="4" captionText="Fig. 10. Cladogram summarizing phylogenetic relationships among members of the Californica Group and two outgroups." figureDoi="http://doi.org/10.5281/zenodo.4758169" httpUri="https://zenodo.org/record/4758169/files/figure.png" pageId="4" pageNumber="192">Fig. 10</figureCitation>
|
||
). Still
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1123,1218,516,540]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1123,1218,516,540]" italics="true" pageId="4" pageNumber="192">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
is basal to
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1338,1437,516,540]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[1338,1437,516,540]" italics="true" pageId="4" pageNumber="192">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
. In three of these trees
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1096,1193,551,575]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1096,1193,551,575]" italics="true" pageId="4" pageNumber="192">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
is immediately basal to
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[864,966,587,611]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[864,966,587,611]" italics="true" pageId="4" pageNumber="192">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
and in the remaining three it is also basal to
|
||
<emphasis box="[866,973,622,646]" italics="true" pageId="4" pageNumber="192">
|
||
<taxonomicName box="[866,969,622,646]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="regilla">C. regilla</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Thus in the consensus of the six trees it appears as a polytomy with the two major clades in the Californica Group (
|
||
<figureCitation box="[1103,1181,694,718]" captionStart="Fig" captionStartId="4.[189,230,1866,1890]" captionTargetBox="[354,1279,1041,1807]" captionTargetId="figure-53@4.[325,1308,1004,1844]" captionTargetPageId="4" captionText="Fig. 10. Cladogram summarizing phylogenetic relationships among members of the Californica Group and two outgroups." figureDoi="http://doi.org/10.5281/zenodo.4758169" httpUri="https://zenodo.org/record/4758169/files/figure.png" pageId="4" pageNumber="192">Fig. 10</figureCitation>
|
||
) and
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[1243,1324,694,717]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis box="[1243,1324,694,717]" italics="true" pageId="4" pageNumber="192">C. nana</emphasis>
|
||
</taxonomicName>
|
||
. The more important question then becomes, “Where does true
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[835,958,765,788]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[835,958,765,788]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
fit on the tree?” In five of the six most parsimonious trees,
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1086,1212,801,824]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[1086,1212,801,824]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
fits in a pectinate fashion up the tree from
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1122,1224,836,859]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[1122,1224,836,859]" italics="true" pageId="4" pageNumber="192">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
and down from
|
||
<taxonomicName class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="ophiona">
|
||
<emphasis italics="true" pageId="4" pageNumber="192">C. ophiona</emphasis>
|
||
</taxonomicName>
|
||
. In the sixth tree
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1140,1263,872,895]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[1140,1263,872,895]" italics="true" pageId="4" pageNumber="192">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
is still up tree from
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[898,1000,907,930]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[898,1000,907,930]" italics="true" pageId="4" pageNumber="192">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
but in a polytomy with
|
||
<taxonomicName box="[1279,1392,907,931]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="ophiona">
|
||
<emphasis box="[1279,1392,907,931]" italics="true" pageId="4" pageNumber="192">C. ophiona</emphasis>
|
||
</taxonomicName>
|
||
and (
|
||
<emphasis box="[844,1204,942,966]" italics="true" pageId="4" pageNumber="192">
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[844,987,942,966]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="californica">C. californica</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Hitchcock" authorityYear="1958" box="[1013,1204,942,966]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="quadrituberosa">C. quadrituberosa</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758169" ID-Zenodo-Dep="4758169" httpUri="https://zenodo.org/record/4758169/files/figure.png" pageId="4" pageNumber="192" startId="4.[189,230,1866,1890]" targetBox="[354,1279,1041,1807]" targetPageId="4">
|
||
<paragraph blockId="4.[189,1443,1866,1926]" pageId="4" pageNumber="192">Fig. 10. Cladogram summarizing phylogenetic relationships among members of the Californica Group and two outgroups.</paragraph>
|
||
</caption>
|
||
<caption ID-Table-UUID="F7FA8CD6DD2FFFCDFF4725A1FE77FCAC" httpUri="http://table.plazi.org/id/F7FA8CD6DD2FFFCDFF4725A1FE77FCAC" pageId="5" pageNumber="193" startId="5.[189,254,232,256]" targetBox="[227,1428,872,1334]" targetIsTable="true" targetPageId="5">
|
||
<paragraph blockId="5.[189,1443,232,824]" pageId="5" pageNumber="193">
|
||
Table 1. Matrix of characters used in the parsimony analysis of the Californica Group, with
|
||
<taxonomicName authorityName="Newport" authorityYear="1848" box="[1232,1395,232,256]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis box="[1232,1395,232,256]" italics="true" pageId="5" pageNumber="193">Capnia vernalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[189,273,268,291]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis box="[189,273,268,291]" italics="true" pageId="5" pageNumber="193">C. nana</emphasis>
|
||
</taxonomicName>
|
||
as outgroups. Character states updated from
|
||
<bibRefCitation author="Nelson, C. R." box="[812,977,267,291]" pageId="5" pageNumber="193" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson (2004)</bibRefCitation>
|
||
, with the correction of data related to
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis italics="true" pageId="5" pageNumber="193">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
and addition of data for
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[579,677,303,327]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[579,677,303,327]" italics="true" pageId="5" pageNumber="193">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
. State of character 14 is corrected for
|
||
<taxonomicName box="[1125,1233,303,327]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="regilla">C. regilla</taxonomicName>
|
||
from 0 in
|
||
<bibRefCitation author="Nelson, C. R." pageId="5" pageNumber="193" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson (2004)</bibRefCitation>
|
||
to 1. All character state texts given are for the “1” state. Question marks (?) indicate missing data. Characters: 1. Knobs present on tergite 8; 2. Knobs present on tergite 9; 3. Lower process of epiproct forked. 4. Female subgenital plate overlapping posterior margin of sternum 8. 5. Upper process of epiproct absent (or very greatly reduced) apically. 6. Upper process of epiproct divided in lateral view. 7. Declivity between processes of epiproct vertical. 8. Lower process of epiproct longer than upper process. 9. Lower process of epiproct much longer than upper process. 10. Knobs present on tergites 5 and 6. 11. Both processes of epiproct of about the same length. 12. Lower process of epiproct widely forked in dorsal view. 13. Declivity between processes of epiproct medially carinate. 14. Declivity between upper and lower processes of epiproct with an extra medial process. 15. Knobs on tergite 8 angulate. 16. Notch between knobs of tergite 8 broad in dorsal view. 17. Notch between upper and lower processes deeply incised in lateral view. 18. Lower process of epiproct narrow in dorsal view. 19. Upper and lower processes of epiproct forming a 90 degree angle in lateral view. 20. Lower process of epiproct with lower surface bent at mid length in lateral view. 21. Lower process of epiproct constricted near apex in dorsal view. 22. Lower process of epiproct evenly tapered, not inflated at midlength in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="5" pageNumber="193">
|
||
<table box="[227,1428,872,1334]" gridcols="23" gridrows="14" pageId="5" pageNumber="193">
|
||
<tr box="[227,1428,872,898]" gridrow="0" pageId="5" pageNumber="193" rowspan-1="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[227,505,872,898]" gridcol="0" gridrow="0" pageId="5" pageNumber="193">Taxon</th>
|
||
<th box="[897,1052,872,898]" colspan="4" colspanRight="3" gridcol="10" gridrow="0" pageId="5" pageNumber="193">
|
||
<emphasis bold="true" box="[911,1052,872,898]" pageId="5" pageNumber="193">Characters</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[227,1428,935,959]" gridrow="1" pageId="5" pageNumber="193" rowspan-0="1">
|
||
<td box="[536,549,935,959]" gridcol="1" gridrow="1" pageId="5" pageNumber="193">1</td>
|
||
<td box="[572,585,935,959]" gridcol="2" gridrow="1" pageId="5" pageNumber="193">2</td>
|
||
<td box="[611,624,935,959]" gridcol="3" gridrow="1" pageId="5" pageNumber="193">3</td>
|
||
<td box="[653,666,935,959]" gridcol="4" gridrow="1" pageId="5" pageNumber="193">4</td>
|
||
<td box="[696,709,935,959]" gridcol="5" gridrow="1" pageId="5" pageNumber="193">5</td>
|
||
<td box="[739,752,935,959]" gridcol="6" gridrow="1" pageId="5" pageNumber="193">6</td>
|
||
<td box="[779,792,935,959]" gridcol="7" gridrow="1" pageId="5" pageNumber="193">7</td>
|
||
<td box="[821,834,935,959]" gridcol="8" gridrow="1" pageId="5" pageNumber="193">8</td>
|
||
<td box="[864,877,935,959]" gridcol="9" gridrow="1" pageId="5" pageNumber="193">9</td>
|
||
<td box="[897,924,935,959]" gridcol="10" gridrow="1" pageId="5" pageNumber="193">10</td>
|
||
<td box="[944,971,935,959]" gridcol="11" gridrow="1" pageId="5" pageNumber="193">11</td>
|
||
<td box="[984,1011,935,959]" gridcol="12" gridrow="1" pageId="5" pageNumber="193">12</td>
|
||
<td box="[1025,1052,935,959]" gridcol="13" gridrow="1" pageId="5" pageNumber="193">13</td>
|
||
<td box="[1065,1092,935,959]" gridcol="14" gridrow="1" pageId="5" pageNumber="193">14</td>
|
||
<td box="[1105,1132,935,959]" gridcol="15" gridrow="1" pageId="5" pageNumber="193">15</td>
|
||
<td box="[1152,1179,935,959]" gridcol="16" gridrow="1" pageId="5" pageNumber="193">16</td>
|
||
<td box="[1192,1219,935,959]" gridcol="17" gridrow="1" pageId="5" pageNumber="193">17</td>
|
||
<td box="[1239,1266,935,959]" gridcol="18" gridrow="1" pageId="5" pageNumber="193">18</td>
|
||
<td box="[1279,1306,935,959]" gridcol="19" gridrow="1" pageId="5" pageNumber="193">19</td>
|
||
<td box="[1320,1347,935,959]" gridcol="20" gridrow="1" pageId="5" pageNumber="193">
|
||
<uuid box="[1320,1347,935,959]" pageId="5" pageNumber="193">20</uuid>
|
||
</td>
|
||
<td box="[1368,1428,935,959]" colspan="2" colspanRight="1" gridcol="21" gridrow="1" pageId="5" pageNumber="193">
|
||
<heading allCaps="true" bold="true" box="[1368,1428,935,959]" fontSize="10" level="1" pageId="5" pageNumber="193" reason="1">
|
||
<emphasis bold="true" box="[1368,1428,935,959]" pageId="5" pageNumber="193">21 22</emphasis>
|
||
</heading>
|
||
</td>
|
||
</tr>
|
||
<tr box="[227,1428,965,989]" gridrow="2" pageId="5" pageNumber="193">
|
||
<th box="[227,505,965,989]" gridcol="0" gridrow="2" pageId="5" pageNumber="193">
|
||
<taxonomicName authorityName="Newport" authorityYear="1848" box="[227,398,965,989]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis box="[227,398,965,989]" italics="true" pageId="5" pageNumber="193">Capnia vernalis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[536,549,965,989]" gridcol="1" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[572,585,965,989]" gridcol="2" gridrow="2" pageId="5" pageNumber="193">0</td>
|
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|
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|
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|
||
<td box="[739,752,965,989]" gridcol="6" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[779,792,965,989]" gridcol="7" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[821,834,965,989]" gridcol="8" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[864,877,965,989]" gridcol="9" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[897,924,965,989]" gridcol="10" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[944,971,965,989]" gridcol="11" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[984,1011,965,989]" gridcol="12" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1025,1052,965,989]" gridcol="13" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1065,1092,965,989]" gridcol="14" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1105,1132,965,989]" gridcol="15" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1152,1179,965,989]" gridcol="16" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1192,1219,965,989]" gridcol="17" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1239,1266,965,989]" gridcol="18" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1279,1306,965,989]" gridcol="19" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1320,1347,965,989]" gridcol="20" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1368,1384,965,989]" gridcol="21" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1411,1428,965,989]" gridcol="22" gridrow="2" pageId="5" pageNumber="193">0</td>
|
||
</tr>
|
||
<tr box="[227,1428,996,1020]" gridrow="3" pageId="5" pageNumber="193">
|
||
<th box="[227,505,996,1020]" gridcol="0" gridrow="3" pageId="5" pageNumber="193">
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[227,366,997,1020]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis box="[227,366,997,1020]" italics="true" pageId="5" pageNumber="193">Capnia nana</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[536,549,996,1020]" gridcol="1" gridrow="3" pageId="5" pageNumber="193">1</td>
|
||
<td box="[572,585,996,1020]" gridcol="2" gridrow="3" pageId="5" pageNumber="193">0</td>
|
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<td box="[611,624,996,1020]" gridcol="3" gridrow="3" pageId="5" pageNumber="193">0</td>
|
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<td box="[653,666,996,1020]" gridcol="4" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[696,709,996,1020]" gridcol="5" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[739,752,996,1020]" gridcol="6" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[779,792,996,1020]" gridcol="7" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[821,834,996,1020]" gridcol="8" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[864,877,996,1020]" gridcol="9" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[897,924,996,1020]" gridcol="10" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[944,971,996,1020]" gridcol="11" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[984,1011,996,1020]" gridcol="12" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1025,1052,996,1020]" gridcol="13" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1065,1092,996,1020]" gridcol="14" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1105,1132,996,1020]" gridcol="15" gridrow="3" pageId="5" pageNumber="193">0</td>
|
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<td box="[1152,1179,996,1020]" gridcol="16" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1192,1219,996,1020]" gridcol="17" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1239,1266,996,1020]" gridcol="18" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1279,1306,996,1020]" gridcol="19" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1320,1347,996,1020]" gridcol="20" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1368,1384,996,1020]" gridcol="21" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1411,1428,996,1020]" gridcol="22" gridrow="3" pageId="5" pageNumber="193">0</td>
|
||
</tr>
|
||
<tr box="[227,1428,1028,1052]" gridrow="4" pageId="5" pageNumber="193">
|
||
<th box="[227,505,1028,1052]" gridcol="0" gridrow="4" pageId="5" pageNumber="193">
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[227,429,1028,1052]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="californica">
|
||
<emphasis box="[227,429,1028,1052]" italics="true" pageId="5" pageNumber="193">Capnia californica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[536,549,1028,1052]" gridcol="1" gridrow="4" pageId="5" pageNumber="193">1</td>
|
||
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|
||
<td box="[611,624,1028,1052]" gridcol="3" gridrow="4" pageId="5" pageNumber="193">1</td>
|
||
<td box="[653,666,1028,1052]" gridcol="4" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[696,709,1028,1052]" gridcol="5" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[739,752,1028,1052]" gridcol="6" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[779,792,1028,1052]" gridcol="7" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[821,834,1028,1052]" gridcol="8" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[864,877,1028,1052]" gridcol="9" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[897,924,1028,1052]" gridcol="10" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[944,971,1028,1052]" gridcol="11" gridrow="4" pageId="5" pageNumber="193">1</td>
|
||
<td box="[984,1011,1028,1052]" gridcol="12" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1025,1052,1028,1052]" gridcol="13" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1065,1092,1028,1052]" gridcol="14" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1105,1132,1028,1052]" gridcol="15" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1152,1179,1028,1052]" gridcol="16" gridrow="4" pageId="5" pageNumber="193">1</td>
|
||
<td box="[1192,1219,1028,1052]" gridcol="17" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1239,1266,1028,1052]" gridcol="18" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1279,1306,1028,1052]" gridcol="19" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1320,1347,1028,1052]" gridcol="20" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1368,1384,1028,1052]" gridcol="21" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1411,1428,1028,1052]" gridcol="22" gridrow="4" pageId="5" pageNumber="193">0</td>
|
||
</tr>
|
||
<tr box="[227,1428,1059,1083]" gridrow="5" pageId="5" pageNumber="193">
|
||
<th box="[227,505,1059,1083]" gridcol="0" gridrow="5" pageId="5" pageNumber="193">
|
||
<taxonomicName authorityName="Hitchcock" authorityYear="1958" box="[227,476,1059,1083]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="quadrituberosa">
|
||
<emphasis box="[227,476,1059,1083]" italics="true" pageId="5" pageNumber="193">Capnia quadrituberosa</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[536,549,1059,1083]" gridcol="1" gridrow="5" pageId="5" pageNumber="193">1</td>
|
||
<td box="[572,585,1059,1083]" gridcol="2" gridrow="5" pageId="5" pageNumber="193">1</td>
|
||
<td box="[611,624,1059,1083]" gridcol="3" gridrow="5" pageId="5" pageNumber="193">1</td>
|
||
<td box="[653,666,1059,1083]" gridcol="4" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[696,709,1059,1083]" gridcol="5" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[739,752,1059,1083]" gridcol="6" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[779,792,1059,1083]" gridcol="7" gridrow="5" pageId="5" pageNumber="193">1</td>
|
||
<td box="[821,834,1059,1083]" gridcol="8" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[864,877,1059,1083]" gridcol="9" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[897,924,1059,1083]" gridcol="10" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[944,971,1059,1083]" gridcol="11" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[984,1011,1059,1083]" gridcol="12" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
<td box="[1025,1052,1059,1083]" gridcol="13" gridrow="5" pageId="5" pageNumber="193">0</td>
|
||
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<taxonomicName box="[227,400,1091,1115]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="ophiona">
|
||
<emphasis box="[227,400,1091,1115]" italics="true" pageId="5" pageNumber="193">Capnia ophiona</emphasis>
|
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</taxonomicName>
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</th>
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||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[227,383,1123,1146]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[227,383,1123,1146]" italics="true" pageId="5" pageNumber="193">Capnia jewetti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
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</tr>
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||
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||
<taxonomicName box="[227,383,1153,1177]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="regilla">
|
||
<emphasis box="[227,383,1153,1177]" italics="true" pageId="5" pageNumber="193">Capnia regilla</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
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||
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||
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||
</tr>
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||
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||
<th box="[227,505,1185,1209]" gridcol="0" gridrow="9" pageId="5" pageNumber="193">
|
||
<taxonomicName box="[227,407,1185,1209]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="saratoga">
|
||
<emphasis box="[227,407,1185,1209]" italics="true" pageId="5" pageNumber="193">Capnia saratoga</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
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||
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||
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||
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||
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||
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||
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||
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||
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|
||
</tr>
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||
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||
<th box="[227,505,1216,1240]" gridcol="0" gridrow="10" pageId="5" pageNumber="193">
|
||
<taxonomicName box="[227,371,1216,1240]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="kersti">
|
||
<emphasis box="[227,371,1216,1240]" italics="true" pageId="5" pageNumber="193">Capnia kersti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
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||
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||
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|
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||
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|
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|
||
</tr>
|
||
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||
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|
||
<taxonomicName box="[227,393,1248,1271]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="ventura">
|
||
<emphasis box="[227,393,1248,1271]" italics="true" pageId="5" pageNumber="193">Capnia ventura</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
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<th box="[227,505,1279,1303]" gridcol="0" gridrow="12" pageId="5" pageNumber="193">
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<emphasis box="[227,405,1279,1303]" italics="true" pageId="5" pageNumber="193">
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<taxonomicName authorityName="Frison" authorityYear="1942" box="[227,401,1279,1303]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="umpqua">Capnia umpqua</taxonomicName>
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,
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</emphasis>
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<typeStatus box="[412,505,1279,1303]" pageId="5" pageNumber="193">holotype</typeStatus>
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<td box="[653,666,1279,1303]" gridcol="4" gridrow="12" pageId="5" pageNumber="193">?</td>
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<taxonomicName authority="Nelson & Baumann, 2009" authorityName="Nelson & Baumann" authorityYear="2009" box="[227,380,1310,1334]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="shasta" status="n. sp.">
|
||
<emphasis box="[227,380,1310,1334]" italics="true" pageId="5" pageNumber="193">Capnia shasta</emphasis>
|
||
</taxonomicName>
|
||
,
|
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<taxonomicNameLabel box="[394,451,1311,1334]" pageId="5" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
|
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</th>
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</tr>
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||
</table>
|
||
</paragraph>
|
||
<subSection lastPageId="6" lastPageNumber="194" pageId="5" pageNumber="193" type="multiple">
|
||
<paragraph blockId="5.[189,797,1416,1902]" pageId="5" pageNumber="193">
|
||
<emphasis bold="true" box="[189,371,1416,1440]" pageId="5" pageNumber="193">Zoogeography.</emphasis>
|
||
With the consensus cladogram of purported relationships (
|
||
<figureCitation box="[499,585,1451,1476]" captionStart="Fig" captionStartId="4.[189,230,1866,1890]" captionTargetBox="[354,1279,1041,1807]" captionTargetId="figure-53@4.[325,1308,1004,1844]" captionTargetPageId="4" captionText="Fig. 10. Cladogram summarizing phylogenetic relationships among members of the Californica Group and two outgroups." figureDoi="http://doi.org/10.5281/zenodo.4758169" httpUri="https://zenodo.org/record/4758169/files/figure.png" pageId="5" pageNumber="193">Fig. 10</figureCitation>
|
||
) and the overall distribution of the group’s species in hand (
|
||
<bibRefCitation author="Nelson, C. R." pageId="5" pageNumber="193" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
) one is tempted to hypothesize the origins, dispersal, and speciation sequence for members of the group. The zoogeography for the group as previously presented (
|
||
<bibRefCitation author="Nelson, C. R." box="[473,629,1629,1653]" pageId="5" pageNumber="193" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
) stands. The Californica Group, taken as a whole (
|
||
<bibRefCitation author="Nelson, C. R." box="[642,792,1665,1689]" pageId="5" pageNumber="193" refId="ref3951" refString="Nelson, C. R. 2004. Systematics of the Capnia californica Species Group, including a morphological phylogeny, zoogeography, and description of Capnia kersti, new species (Plecoptera: Capniidae). Annals of the" type="book" year="2004">Nelson 2004</bibRefCitation>
|
||
,
|
||
<figureCitation box="[189,254,1700,1724]" captionStart="Figs" captionStartId="2.[189,238,1759,1783]" captionTargetBox="[287,1323,258,1668]" captionTargetId="figure-53@2.[287,1345,258,1668]" captionTargetPageId="2" captionText="Figs. 1-8. Capnia shasta, scanning electron micrographs: 1. male terminalia, dorsal, anterior down, 152x, scale 200µm. 2. male terminalia, lateral, anterior to right, 249x, scale 100µm. 3. male epiproct, dorsal, anterior to right, 387x, scale 50µm. 4. male epiproct, lateral, anterior to right, 389x, scale 50µm. 5. details of male epiproct, dorsal, anterior to right, 868x, scale 20µm. 6. details of male epiproct, lateral, anterior to right, 1959x, scale 10µm. 7. male terminalia, dorsal, anterior down, 157x, scale 100µm. 8. female terminalia, ventral, anterior down, 108x, scale 200µm." figureDoi="http://doi.org/10.5281/zenodo.4758165" httpUri="https://zenodo.org/record/4758165/files/figure.png" pageId="5" pageNumber="193">Fig. 3</figureCitation>
|
||
) divides into two sister taxa basally (Ventura Subgroup) and (Californica Subgroup +
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[660,778,1736,1759]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[660,778,1736,1759]" italics="true" pageId="5" pageNumber="193">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[189,300,1772,1795]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[189,300,1772,1795]" italics="true" pageId="5" pageNumber="193">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
). But our new species comes out in a polytomy with these two subgroups (and
|
||
<taxonomicName authorityName="Claassen" authorityYear="1924" box="[701,788,1807,1830]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis box="[701,788,1807,1830]" italics="true" pageId="5" pageNumber="193">C. nana</emphasis>
|
||
</taxonomicName>
|
||
) and thus leads to no strong zoogeographic hypotheses.
|
||
</paragraph>
|
||
<paragraph blockId="5.[835,1443,1416,1901]" lastBlockId="6.[189,798,232,967]" lastPageId="6" lastPageNumber="194" pageId="5" pageNumber="193">
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[873,1026,1416,1440]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[873,1026,1416,1440]" italics="true" pageId="5" pageNumber="193">Capnia shasta</emphasis>
|
||
</taxonomicName>
|
||
is currently known from a single locality on a single date. It is currently known only from far down the flanks of Mount Shasta in a small tributary of the Sacramento River at the northern head of California’s Central Valley. We suggest that further careful winter collections of many sites in the area are warranted. We examined a range of specimens of
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[999,1119,1665,1688]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[999,1119,1665,1688]" italics="true" pageId="5" pageNumber="193">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
from throughout its broad
|
||
<collectingRegion box="[835,924,1700,1724]" country="United States of America" name="Oregon" pageId="5" pageNumber="193">Oregon</collectingRegion>
|
||
– southern
|
||
<collectingRegion box="[1057,1176,1700,1724]" country="United States of America" name="California" pageId="5" pageNumber="193">California</collectingRegion>
|
||
range. The shape of the epiproct in these specimens varied widely in relative width both within series from a single collection and across its range. Using this information we tentatively propose that
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[1129,1228,1842,1866]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[1129,1228,1842,1866]" italics="true" pageId="5" pageNumber="193">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[1245,1350,1842,1866]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="jewetti">
|
||
<emphasis box="[1245,1350,1842,1866]" italics="true" pageId="5" pageNumber="193">C. jewetti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="ophiona">
|
||
<emphasis italics="true" pageId="5" pageNumber="193">C. ophiona</emphasis>
|
||
</taxonomicName>
|
||
arose as independent peripheral isolates from the widespread
|
||
<taxonomicName authorityName="Frison" authorityYear="1942" box="[410,545,232,256]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="194" phylum="Arthropoda" rank="species" species="umpqua">
|
||
<emphasis box="[410,545,232,256]" italics="true" pageId="6" pageNumber="194">C. umpqua</emphasis>
|
||
</taxonomicName>
|
||
-like ancestor. This speculative hypothesis could best be tested using fast evolving sequences of DNA. We have little hope that further morphological study of this question will resolve these polytomies, and hope that the sequences will.
|
||
</paragraph>
|
||
<paragraph blockId="6.[189,798,232,967]" pageId="6" pageNumber="194">
|
||
<emphasis bold="true" box="[189,347,445,469]" pageId="6" pageNumber="194">Conclusions.</emphasis>
|
||
This new, interesting species shows that small, relictual populations and species probably remain to be discovered in western North America. Careful, fine scale collecting of the numerous streams in northern
|
||
<collectingRegion box="[326,444,587,611]" country="United States of America" name="California" pageId="6" pageNumber="194">California</collectingRegion>
|
||
continues to yield new species. The limited distribution of
|
||
<taxonomicName authorityName="Nelson & Baumann" authorityYear="2009" box="[508,605,623,647]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="194" phylum="Arthropoda" rank="species" species="shasta">
|
||
<emphasis box="[508,605,623,647]" italics="true" pageId="6" pageNumber="194">C. shasta</emphasis>
|
||
</taxonomicName>
|
||
points to a need for careful conservation of the tributaries of the Sacramento River. The Californica Group remains an important and virtually untapped resource to learn more about the paleogeography of this part of western North America. The use of sequence data on a phylogeographic scale remains a next best step in sorting out the range relationships of these populations and species in understanding this perplexingly complex group of stoneflies.
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |