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<document id="EC4593D410A9C0953DC1DD43776EB4B6" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629576139757" checkinUser="felipe" docAuthor="McAlpine, DK" docDate="2007" docId="03FA02406A69650637C64A0DB817D3B0" docLanguage="en" docName="RecAustMus.59.3.143-219.pdf" docOrigin="Records of the Australian Museum 59" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Borboroides parva McAlpine 2007, n.sp." docType="treatment" docVersion="2" lastPageNumber="190" masterDocId="FFC37A386A456529342A4C0DBD22D171" masterDocTitle="Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with Reconsideration of the Status of Families Heleomyzidae and Sphaeroceridae, and Descriptions of Femoral Gland-baskets" masterLastPageNumber="219" masterPageNumber="143" pageNumber="187" updateTime="1699078874016" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C0D5E6E53E11AD594C7FEFA157465D46">Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with Reconsideration of the Status of Families Heleomyzidae and Sphaeroceridae, and Descriptions of Femoral Gland-baskets</mods:title>
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<mods:namePart id="432533E5897BD6EA568BA6DB8044CEE1">McAlpine, DK</mods:namePart>
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<mods:title id="B72A53C1B4623631409B026983D72D27">Records of the Australian Museum</mods:title>
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<mods:date id="C4BA499CF83E0A4A364B21245DD05588">2007</mods:date>
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<treatment id="03FA02406A69650637C64A0DB817D3B0" LSID="urn:lsid:plazi:treatment:03FA02406A69650637C64A0DB817D3B0" httpUri="http://treatment.plazi.org/id/03FA02406A69650637C64A0DB817D3B0" lastPageId="47" lastPageNumber="190" pageId="44" pageNumber="187">
<subSubSection id="C349E0DD6A69650537C64A0DB837D76B" box="[1004,1301,1536,1562]" pageId="44" pageNumber="187" type="nomenclature">
<paragraph id="8BECB3566A69650537C64A0DB837D76B" blockId="44.[1004,1301,1536,1562]" box="[1004,1301,1536,1562]" pageId="44" pageNumber="187">
<heading id="D0A4043A6A69650537C64A0DB837D76B" box="[1004,1301,1536,1562]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="187" reason="2">
<emphasis id="B9276F446A69650537C64A0DB837D76B" bold="true" box="[1004,1301,1536,1562]" pageId="44" pageNumber="187">
<taxonomicName id="4C53C8D56A69650537C64A0DB9F0D76B" ID-CoL="MJF4" authority="McAlpine, 2007" authorityName="McAlpine" authorityYear="2007" box="[1004,1234,1536,1562]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="44" pageNumber="187" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="B9276F446A69650537C64A0DB9F0D76B" bold="true" box="[1004,1234,1536,1562]" italics="true" pageId="44" pageNumber="187">Borboroides parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A214D23F6A69650530F34A0DB837D76B" box="[1241,1301,1536,1562]" pageId="44" pageNumber="187" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349E0DD6A696505303B4A23B9D3D739" box="[1041,1265,1582,1608]" pageId="44" pageNumber="187" type="description">
<paragraph id="8BECB3566A696505303B4A23B9D3D739" blockId="44.[839,1467,1582,1849]" box="[1041,1265,1582,1608]" pageId="44" pageNumber="187">
<figureCitation id="1368AFD36A696505303B4A23B9BBD739" box="[1041,1177,1582,1608]" captionStart="Figs 1114" captionStartId="7.[121,162,994,1015]" captionTargetBox="[121,1421,151,977]" captionTargetId="figure-13@7.[121,1421,151,980]" captionTargetPageId="7" captionText="Figs 1114. (11) Borboroides fimbria, part of left hind femur, showing posteroventral tract of microtrichia covering glandular groove; scale = 10 µm. (12) Detail of glandular groove of same; scale = 2 µm. (13) B. parva, part of ventral surface of left hind femur showing many microtrichia and larger macrotrichia; scale = 10 µm. (14) Detail of same showing microtrichia and posteroventral macrotrichia, apparently with some adhering secretion; scale = 3 µm." pageId="44" pageNumber="187">Figs 13, 14</figureCitation>
,
<figureCitation id="1368AFD36A696505308C4A23B9D3D739" box="[1190,1265,1582,1608]" captionStart-0="Figs 8891" captionStart-1="Figs 9296" captionStartId-0="45.[121,162,1068,1089]" captionStartId-1="46.[166,207,1877,1898]" captionTargetBox-0="[121,1421,151,1054]" captionTargetBox-1="[183,1458,157,1835]" captionTargetId-0="figure-13@45.[121,1421,151,1058]" captionTargetId-1="figure-136@46.[166,1466,151,1863]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="Figs 8891. (88) Borboroides parva, head. (89) Dorsal view of thorax of same, postalar bristle indicated. (90) Right propleural and mesopleural region of thorax of same. (91) Part of right mid leg of male of same, anterior view." captionText-1="Figs 9296. (92) Borboroides parva, wing. (93) Right antenna of same; scale = 0.05 mm. (94) Male postabdomen of same, left lateral view; scale = 0.05 mm.(95) Hypandrial structures and aedeagus of same, left lateral view; scale = 0.05 mm.(96) Epandrial and hypandrial structures of same, anteroventral view, gonites and aedeagus cut short; scale = 0.05 mm. aa, aedeagal apodeme; avb, anteroventral bridge of epandrium; bp, basiphallus; c, cercus; co, conus; dp, distiphallus; e, epandrium; g, gonite; hy, hypandrial sclerite; pm, prehypandrial membrane; pr, proctiger; sc, sacculus; ss, surstylus; s6, s8, sternites 6 and 8; t6, tergite 6." pageId="44" pageNumber="187">8896</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349E0DD6A696505376D4A58B856D648" pageId="44" pageNumber="187" type="materials_examined">
<paragraph id="8BECB3566A696505376D4A58B856D648" blockId="44.[839,1467,1582,1849]" pageId="44" pageNumber="187">
<emphasis id="B9276F446A696505376D4A58B93BD71C" bold="true" box="[839,1049,1621,1645]" pageId="44" pageNumber="187">Material examined</emphasis>
.
<materialsCitation id="3B3BB90B6A696505300E4A58B996D7B5" collectingDate="2002-05-31" collectionCode="AM" collectorName="DK McAlpine &amp; Mounted" elevation="220" latitude="-33.183334" location="Putty Road" longLatPrecision="1206" longitude="150.68333" pageId="44" pageNumber="187" specimenCode="K219752" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<typeStatus id="54E80DF46A696505300E4A58B9B4D71D" box="[1060,1174,1621,1644]" pageId="44" pageNumber="187" type="holotype">HOLOTYPE</typeStatus>
.?,
<collectingRegion id="49977DB46A69650530EB4A58B859D71D" box="[1217,1403,1621,1644]" country="Australia" name="New South Wales" pageId="44" pageNumber="187">New South Wales</collectingRegion>
:
<location id="8E8CE58D6A69650531A94A58BEA0D7F8" LSID="urn:lsid:plazi:treatment:03FA02406A69650637C64A0DB817D3B0:8E8CE58D6A69650531A94A58BEA0D7F8" latitude="-33.183334" longLatPrecision="1206" longitude="150.68333" name="Putty Road" pageId="44" pageNumber="187" stateProvince="New South Wales">Putty Road</location>
,
<locationDeviation id="9A5EAA746A69650537BA4A7FB9B7D7F8" box="[912,1173,1650,1673]" location="Colo River" pageId="44" pageNumber="187">
<quantity id="4CAB1EB36A69650537BA4A7FBEF4D7F8" box="[912,982,1650,1673]" metricMagnitude="4" metricUnit="m" metricValue="4.1" pageId="44" pageNumber="187" unit="km" value="41.0">41 km</quantity>
N of Colo River
</locationDeviation>
bridge,
<geoCoordinate id="EE67D5916A69650530DE4A7FB865D7F8" box="[1268,1351,1650,1673]" degrees="33" direction="south" minutes="11" orientation="latitude" pageId="44" pageNumber="187" precision="925" value="-33.183334">33°11'S</geoCoordinate>
<geoCoordinate id="EE67D5916A696505317B4A7FB894D7F8" box="[1361,1462,1650,1673]" degrees="150" direction="east" minutes="41" orientation="longitude" pageId="44" pageNumber="187" precision="925" value="150.68333">150°41'E</geoCoordinate>
, c.
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<elevation id="007E54656A69650537484A9DBE8AD7D6" box="[866,936,1680,1703]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="44" pageNumber="187" unit="m" value="220.0">220 m</elevation>
</quantity>
[Yengo National Park],
<date id="FFED95966A69650530994A9DB83DD7D6" box="[1203,1311,1680,1703]" pageId="44" pageNumber="187" value="2002-05-31">
<collectingDate id="EFA96C7E6A69650530994A9DB83DD7D6" box="[1203,1311,1680,1703]" pageId="44" pageNumber="187" value="2002-05-31">31.v.2002</collectingDate>
</date>
,
<collectorName id="26A6D6806A69650531064A9DB85CD7D6" box="[1324,1406,1680,1703]" name="DK McAlpine" pageId="44" pageNumber="187">D.K.M.</collectorName>
(
<collectionCode id="ED422B936A69650531A44A9DB899D7D6" box="[1422,1467,1680,1703]" country="Australia" name="Australian Museum" pageId="44" pageNumber="187" type="Museum">AM</collectionCode>
<specimenCode id="DBF51B2D6A696505376D4AA0BE8DD7B5" box="[839,943,1709,1732]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="44" pageNumber="187" type="Museum">K219752</specimenCode>
).
<collectorName id="26A6D6806A69650537EA4AA0B902D7B5" box="[960,1056,1709,1732]" pageId="44" pageNumber="187">Mounted</collectorName>
on card point
</materialsCitation>
.
<materialsCitation id="3B3BB90B6A69650530974AA0B852D648" collectingDate="2002-05-31" collectingDateMax="2002-09" collectingDateMin="2002-05-31" collectionCode="AM, ANIC, BM, CNC, MNHG, USNM, AMST" collectorName="DK McAlpine" county="May" elevation="220" latitude="-33.166668" location="Putty Road" longLatPrecision="1206" longitude="150.7" municipality="June" pageId="44" pageNumber="187" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<typeStatus id="54E80DF46A69650530974AA0B81AD7B5" box="[1213,1336,1709,1732]" pageId="44" pageNumber="187" type="paratype">PARATYPES</typeStatus>
.
<collectingRegion id="49977DB46A696505316F4AA0BEA8D790" country="Australia" name="New South Wales" pageId="44" pageNumber="187">New South Wales</collectingRegion>
: 139??, 32!!, same locality as holotype,
<collectingCounty id="628DCBDA6A69650531664AC7B85ED790" box="[1356,1404,1738,1761]" pageId="44" pageNumber="187">May</collectingCounty>
,
<collectingMunicipality id="6B88292C6A69650531AF4AC7B895D790" box="[1413,1463,1738,1761]" pageId="44" pageNumber="187">June</collectingMunicipality>
,
<date id="FFED95966A696505376D4AE5BE9FD78E" box="[839,957,1768,1791]" pageId="44" pageNumber="187" value="2002-09">
<collectingDate id="EFA96C7E6A696505376D4AE5BE9FD78E" box="[839,957,1768,1791]" pageId="44" pageNumber="187" value="2002-09">Sept. 2002</collectingDate>
</date>
2004,
<collectorName id="26A6D6806A69650530254AE5B940D78E" box="[1039,1122,1768,1791]" name="DK McAlpine" pageId="44" pageNumber="187">D.K.M.</collectorName>
(
<collectionCode id="ED422B936A69650530594AE5B980D78E" box="[1139,1186,1768,1791]" country="Australia" name="Australian Museum" pageId="44" pageNumber="187" type="Museum">AM</collectionCode>
,
<collectionCode id="ED422B936A69650530844AE5B9D0D78E" box="[1198,1266,1768,1791]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="44" pageNumber="187">ANIC</collectionCode>
,
<collectionCode id="ED422B936A696505312A4AE5B80ED78E" box="[1280,1324,1768,1791]" country="United Kingdom" name="Bristol Museum" pageId="44" pageNumber="187">BM</collectionCode>
, BPB,
<collectionCode id="ED422B936A69650531574AE5B894D78E" box="[1405,1462,1768,1791]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="44" pageNumber="187">CNC</collectionCode>
,
<collectionCode id="ED422B936A696505376D4B08BEB8D66D" box="[839,922,1797,1820]" pageId="44" pageNumber="187">MNHG</collectionCode>
,
<collectionCode id="ED422B936A69650537894B08BED1D66D" box="[931,1011,1797,1820]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="44" pageNumber="187" type="Museum">USNM</collectionCode>
); 22??, 16!!,
<location id="8E8CE58D6A696505308C4B08B83BD66D" LSID="urn:lsid:plazi:treatment:03FA02406A69650637C64A0DB817D3B0:8E8CE58D6A696505308C4B08B83BD66D" box="[1190,1305,1797,1820]" county="May" latitude="-33.166668" longLatPrecision="1206" longitude="150.7" municipality="June" name="Putty Road" pageId="44" pageNumber="187" stateProvince="New South Wales">Putty Road</location>
at
<location id="8E8CE58D6A696505311E4B08B894D66D" LSID="urn:lsid:plazi:treatment:03FA02406A69650637C64A0DB817D3B0:8E8CE58D6A696505311E4B08B894D66D" box="[1332,1462,1797,1820]" county="May" latitude="-33.166668" longLatPrecision="1206" longitude="150.7" municipality="June" name="Tinda Creek" pageId="44" pageNumber="187" stateProvince="New South Wales">Tinda Creek</location>
,
<geoCoordinate id="EE67D5916A696505376D4B2FBEBBD648" box="[839,921,1826,1849]" degrees="33" direction="south" minutes="10" orientation="latitude" pageId="44" pageNumber="187" precision="925" value="-33.166668">33°10'S</geoCoordinate>
<geoCoordinate id="EE67D5916A696505378A4B2FB926D648" box="[928,1028,1826,1849]" degrees="150" direction="east" minutes="42" orientation="longitude" pageId="44" pageNumber="187" precision="925" value="150.7">150°42'E</geoCoordinate>
,
<date id="FFED95966A69650530254B2FB95AD648" box="[1039,1144,1826,1849]" pageId="44" pageNumber="187" value="2002-05-31">
<collectingDate id="EFA96C7E6A69650530254B2FB95AD648" box="[1039,1144,1826,1849]" pageId="44" pageNumber="187" value="2002-05-31">31.v.2002</collectingDate>
</date>
,
<collectorName id="26A6D6806A69650530A94B2FB9F7D648" box="[1155,1237,1826,1849]" name="DK McAlpine" pageId="44" pageNumber="187">D.K.M.</collectorName>
(
<collectionCode id="ED422B936A69650530C94B2FB833D648" box="[1251,1297,1826,1849]" country="Australia" name="Australian Museum" pageId="44" pageNumber="187" type="Museum">AM</collectionCode>
,
<collectionCode id="ED422B936A69650531304B2FB84AD648" box="[1306,1384,1826,1849]" pageId="44" pageNumber="187">AMST</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349E0DD6A696506376D4B50BF5CD4CD" lastPageId="47" lastPageNumber="190" pageId="44" pageNumber="187" type="description">
<paragraph id="8BECB3566A696505376D4B50B916D6E0" blockId="44.[839,1467,1885,1967]" pageId="44" pageNumber="187">
<emphasis id="B9276F446A696505376D4B50BEF5D604" bold="true" box="[839,983,1885,1909]" pageId="44" pageNumber="187">Description</emphasis>
(male, female). Minute, blackish, stout sphaerocerine-like fly.
</paragraph>
<paragraph id="8BECB3566A696504374F4B95BF02D76E" blockId="44.[839,1467,1885,1967]" lastBlockId="45.[121,749,1163,2007]" lastPageId="45" lastPageNumber="188" pageId="44" pageNumber="187">
<emphasis id="B9276F446A696505374F4B95BEFCD6DE" box="[869,990,1944,1967]" italics="true" pageId="44" pageNumber="187">Coloration</emphasis>
. Head, thorax, legs and abdomen almost uniformly black to very dark brown. Postfrons entirely dull-pruinescent, usually with yellowish brown suffusion anteriorly; lower margin of cheek grey-pruinescent. Antenna blackish. Palpus dark grey-brown. Mesoscutum and scutellum grey-pruinescent; mesopleuron shining blackish centrally and anteroventrally, extensively grey-pruinescent anterodorsally and posterodorsally; sternopleuron largely subshining, with little pruinescence; pteropleuron and pleurotergite relatively densely grey-pruinescent. Legs almost uniformly dark brown; fore coxa tending to yellowish brown. Wing without markings; membrane tinged with smoky brown. Halter with yellowish brown pedicel and darker greyish brown capitellum. Abdominal tergites extensively grey- to brown-pruinescent.
</paragraph>
<caption id="DF2CE3DE6A68650434534821B92ED52D" pageId="45" pageNumber="188" startId="45.[121,162,1068,1089]" targetBox="[121,1421,151,1054]" targetPageId="45">
<paragraph id="8BECB3566A68650434534821B92ED52D" blockId="45.[121,1421,1068,1116]" pageId="45" pageNumber="188">
Figs 8891. (
<emphasis id="B9276F446A68650434D74821BC37D530" box="[253,277,1068,1089]" italics="true" pageId="45" pageNumber="188">88</emphasis>
)
<taxonomicName id="4C53C8D56A686504350F4821BCFED530" authorityName="McAlpine" authorityYear="2007" box="[293,476,1068,1089]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="188" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B9276F446A686504350F4821BCFED530" box="[293,476,1068,1089]" italics="true" pageId="45" pageNumber="188">Borboroides parva</emphasis>
</taxonomicName>
, head. (
<emphasis id="B9276F446A68650436044821BF64D530" box="[558,582,1068,1089]" italics="true" pageId="45" pageNumber="188">89</emphasis>
) Dorsal view of thorax of same, postalar bristle indicated. (
<emphasis id="B9276F446A68650430B04821B990D530" box="[1178,1202,1068,1089]" italics="true" pageId="45" pageNumber="188">90</emphasis>
) Right propleural and mesopleural region of thorax of same. (
<emphasis id="B9276F446A68650435DE484ABF2ED52D" box="[500,524,1095,1116]" italics="true" pageId="45" pageNumber="188">91</emphasis>
) Part of right mid leg of male of same, anterior view.
</paragraph>
</caption>
<paragraph id="8BECB3566A68650434BD4A2BB87ED422" blockId="45.[121,749,1163,2007]" lastBlockId="45.[794,1422,1164,2009]" pageId="45" pageNumber="188">
<emphasis id="B9276F446A68650434BD4A2BBDF3D74C" box="[151,209,1574,1597]" italics="true" pageId="45" pageNumber="188">Head</emphasis>
higher than long; eye subcircular, with moderately sparse but obvious ommatrichia; postfrons broad and convex, with few, mostly erect anterior setulae; face slightly concave in profile, but with upper median part elevated, i.e. more distinctly carinate than in
<taxonomicName id="4C53C8D56A68650435E24A96BF32D7C3" box="[456,528,1691,1714]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="188" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A68650435E24A96BF32D7C3" box="[456,528,1691,1714]" italics="true" pageId="45" pageNumber="188">B. atra</emphasis>
</taxonomicName>
etc.; height of cheek 0.330.44 of height of eye; postvertical bristle moderately developed, smaller than in
<taxonomicName id="4C53C8D56A68650435B24ADBBCC0D79C" box="[408,482,1750,1773]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="188" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A68650435B24ADBBCC0D79C" box="[408,482,1750,1773]" italics="true" pageId="45" pageNumber="188">B. atra</emphasis>
</taxonomicName>
; ocellar bristle large, but not potentially reaching ptilinal suture; posterior frontoorbital bristle reclinate and slightly sloped outwards; anterior fronto-orbital about as long as or slightly longer than posterior one, strongly directed outwards. Antenna more markedly porrect than in
<emphasis id="B9276F446A68650435CA4B65BF91D60E" box="[480,691,1896,1919]" italics="true" pageId="45" pageNumber="188">
<taxonomicName id="4C53C8D56A68650435CA4B65BF77D60E" authorityName="McAlpine" authorityYear="2007" box="[480,597,1896,1919]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="188" phylum="Arthropoda" rank="species" species="fimbria">B. fimbria</taxonomicName>
,
<taxonomicName id="4C53C8D56A68650436494B65BF91D60E" box="[611,691,1896,1919]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="188" phylum="Arthropoda" rank="species" species="atra">B. atra</taxonomicName>
</emphasis>
etc.; segment 3 almost hemispherical, with doughnut-shaped proximal surface containing large central cavity, with both aristal insertion and sacculus remarkably close to basal rim; arista c. 1.62.0× as long as maximum diameter of eye, with numerous hairs/pubescence on whole length, those on distal half slightly longer than diameter of segment 5, those on basal part shorter. Prelabrum well sclerotized but not prominent, slightly but significantly larger in female than in male; palpus stout and moderately short; proboscis moderately small, but labella not particularly reduced.
</paragraph>
<paragraph id="8BECB3566A68650637124957BCCDD010" blockId="45.[794,1422,1164,2009]" lastBlockId="47.[121,749,154,1469]" lastPageId="47" lastPageNumber="190" pageId="45" pageNumber="188">
<emphasis id="B9276F446A68650437124957BEAAD400" box="[824,904,1370,1393]" italics="true" pageId="45" pageNumber="188">Thorax</emphasis>
stout; intradorsocentral setulae in four rows (two pairs) anteriorly and centrally; usually a pair of prescutellar acrostichal setulae scarcely larger than other setulae; scutellum without setulae; mesopleuron with few coarse posterior setulae; pteropleuron without setulae; the following thoracic bristles present: humeral, presutural, 1+1 notopleurals, postalar (displaced, mesad of postalar callus), 0+1 dorsocentrals; supra-alar and additional postsutural dorsocentral sometimes weakly differentiated; posterior intra-alar represented only by small setula; two unequal pairs of scutellars; one sternopleural. Fore femur with two to four posteroventral bristles and mostly shorter posterodorsal bristles; mid femur with one differentiated anterior bristle slightly beyond mid length; hind femur without differentiated bristles; fore and hind tibiae with some rather long setulae but no differentiated bristles or spurs; mid tibia with two large preapical dorsal bristles at slightly different levels, long apical ventral spur, and smaller apical anterior spur, otherwise without anterior bristles. Wing: costa with one well differentiated anterodorsal bristle just before humeral break (rarely duplicated or located just beyond humeral break); subcosta weakened opposite humeral break of costa, but sclerotized for short distance beyond break; anterior crossvein meeting vein 4 near or slightly basad of mid-length of discal cell; apical section of vein 4 c. 2.42.9× as long as penultimate section; basal crossvein oblique and aligned with strongly curved section of vein 5 bordering discal cell; apical section of vein 5 straight or almost so; anal crossvein and apical section of vein 6 absent.
</paragraph>
<caption id="DF2CE3DE6A6B6507348C4B58BED9D6A4" pageId="46" pageNumber="189" startId="46.[166,207,1877,1898]" targetBox="[183,1458,157,1835]" targetPageId="46">
<paragraph id="8BECB3566A6B6507348C4B58BED9D6A4" blockId="46.[166,1467,1877,2005]" pageId="46" pageNumber="189">
Figs 9296. (
<emphasis id="B9276F446A6B6507350D4B5BBC1DD61A" box="[295,319,1878,1899]" italics="true" pageId="46" pageNumber="189">92</emphasis>
)
<taxonomicName id="4C53C8D56A6B650735644B5BBF26D61A" authorityName="McAlpine" authorityYear="2007" box="[334,516,1878,1899]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="189" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B9276F446A6B650735644B5BBF26D61A" box="[334,516,1878,1899]" italics="true" pageId="46" pageNumber="189">Borboroides parva</emphasis>
</taxonomicName>
, wing. (
<emphasis id="B9276F446A6B6507367F4B5BBF4FD61A" box="[597,621,1878,1899]" italics="true" pageId="46" pageNumber="189">93</emphasis>
) Right antenna of same; scale = 0.05 mm. (
<emphasis id="B9276F446A6B650730334B5BB913D61A" box="[1049,1073,1878,1899]" italics="true" pageId="46" pageNumber="189">94</emphasis>
) Male postabdomen of same, left lateral view; scale = 0.05 mm. (
<emphasis id="B9276F446A6B650735A64B7DBC86D6F4" box="[396,420,1904,1925]" italics="true" pageId="46" pageNumber="189">95</emphasis>
) Hypandrial structures and aedeagus of same, left lateral view; scale = 0.05 mm. (
<emphasis id="B9276F446A6B6507308C4B7DB99CD6F4" box="[1190,1214,1904,1925]" italics="true" pageId="46" pageNumber="189">96</emphasis>
) Epandrial and hypandrial structures of same, anteroventral view, gonites and aedeagus cut short; scale = 0.05 mm.
<emphasis id="B9276F446A6B650737C44B86B924D6D1" box="[1006,1030,1931,1952]" italics="true" pageId="46" pageNumber="189">aa</emphasis>
, aedeagal apodeme;
<emphasis id="B9276F446A6B650730E64B86B9CDD6D1" box="[1228,1263,1931,1952]" italics="true" pageId="46" pageNumber="189">avb</emphasis>
, anteroventral bridge of epandrium;
<emphasis id="B9276F446A6B6507351F4BABBC6FD6CA" box="[309,333,1958,1979]" italics="true" pageId="46" pageNumber="189">bp</emphasis>
, basiphallus;
<emphasis id="B9276F446A6B650735F84BABBCFFD6CA" box="[466,477,1958,1979]" italics="true" pageId="46" pageNumber="189">c</emphasis>
, cercus;
<emphasis id="B9276F446A6B6507361E4BABBF69D6CA" box="[564,587,1958,1979]" italics="true" pageId="46" pageNumber="189">co</emphasis>
, conus;
<emphasis id="B9276F446A6B650736B64BABBF96D6CA" box="[668,692,1958,1979]" italics="true" pageId="46" pageNumber="189">dp</emphasis>
, distiphallus;
<emphasis id="B9276F446A6B650737164BABBE65D6CA" box="[828,839,1958,1979]" italics="true" pageId="46" pageNumber="189">e</emphasis>
, epandrium;
<emphasis id="B9276F446A6B650737ED4BABBEF1D6CA" box="[967,979,1958,1979]" italics="true" pageId="46" pageNumber="189">g</emphasis>
, gonite;
<emphasis id="B9276F446A6B650730034BABB962D6CA" box="[1065,1088,1958,1979]" italics="true" pageId="46" pageNumber="189">hy</emphasis>
, hypandrial sclerite;
<emphasis id="B9276F446A6B650731264BABB80BD6CA" box="[1292,1321,1958,1979]" italics="true" pageId="46" pageNumber="189">pm</emphasis>
, prehypandrial membrane;
<emphasis id="B9276F446A6B650735324BCDBC0FD6A4" box="[280,301,1984,2005]" italics="true" pageId="46" pageNumber="189">pr</emphasis>
, proctiger;
<emphasis id="B9276F446A6B650735B74BCDBC93D6A4" box="[413,433,1984,2005]" italics="true" pageId="46" pageNumber="189">sc</emphasis>
, sacculus;
<emphasis id="B9276F446A6B650736314BCDBF0CD6A4" box="[539,558,1984,2005]" italics="true" pageId="46" pageNumber="189">ss</emphasis>
, surstylus;
<emphasis id="B9276F446A6B650736B64BCDBFF0D6A4" box="[668,722,1984,2005]" italics="true" pageId="46" pageNumber="189">s6, s8</emphasis>
, sternites 6 and 8;
<emphasis id="B9276F446A6B650737A24BCDBEB8D6A4" box="[904,922,1984,2005]" italics="true" pageId="46" pageNumber="189">t6</emphasis>
, tergite 6.
</paragraph>
</caption>
<paragraph id="8BECB3566A6A650634BD4D65BCD6D415" blockId="47.[121,749,154,1469]" pageId="47" pageNumber="190">
<emphasis id="B9276F446A6A650634BD4D65BC22D00E" box="[151,256,360,383]" italics="true" pageId="47" pageNumber="190">Abdomen</emphasis>
. Sternite 1 very short, sclerotized only at sides; tergite 5 slightly shorter than tergite 4 (both sexes). Male postabdomen: tergite 6 rather small, undivided; each spiracle
<quantity id="4CAB1EB36A6A650634F24DCDBC27D0A6" box="[216,261,448,471]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="47" pageNumber="190" unit="in" value="6.0">6 in</quantity>
membrane below lateral margin of tergite; spiracle 7 apparently absent; protandrial region posteriorly attenuated to form very short hypopygial petiole; sternite 6 reduced to long, darkly sclerotized strip on left side only, dorsally connected to sternite 8; sternite 7 distinct on left side, extending round ventral surface, weakly sclerotized; sternite 8 well sclerotized dorsally with two or three pairs of bristles or large setulae and no or few smaller setulae, with sclerotized ventral bridge anteriorly, and behind this with submarginal heavily sclerotized strip on each side terminating on posterior condyle; epandrium more elongate than in
<taxonomicName id="4C53C8D56A6A650634EE4F0EBC1DD26B" authorityName="D. McAlpine" authorityYear="1985" box="[196,319,771,794]" class="Insecta" family="Heleomyzidae" genus="Heleomicra" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B9276F446A6A650634EE4F0EBC1DD26B" box="[196,319,771,794]" italics="true" pageId="47" pageNumber="190">Heleomicra</emphasis>
</taxonomicName>
but less so than in other
<taxonomicName id="4C53C8D56A6A6506361F4F0EBFCBD268" box="[565,745,770,794]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9276F446A6A6506361F4F0EBF94D26B" box="[565,694,771,794]" italics="true" pageId="47" pageNumber="190">Borboroides</emphasis>
spp.
</taxonomicName>
, with several stout setulae, and anteroventral bridge sclerotized but much attenuated; prehypandrial membrane smooth and glassy, without microtrichia; surstylus moderately elongate, articulated basally, with numerous microtrichia and few larger setulae on inner surface, with few microtrichia only on outer surface; hypandrium sclerotized anteriorly in front of aedeagal base, with lateral extension on each side articulating with epandrium but bearing no setulae; gonite long, curved, its apex posteriorly directed, with few small, mostly blunt setulae distally; basiphallus elongate, curved, with apex posteriorly directed; distiphallus small, membranous, largely recessed in subapical part of basiphallus; phallapodeme short, broadly bilaterally compressed, with no sclerotized connection to hypandrium; cerci joined by membranous proctiger for their full length, lacking ventral connecting sclerotized bar at bases and definite articulation with epandrium, each with fine microtrichia, few setulae, and one very long terminal bristle. Female postabdomen simple, tapered on c. posterior half; cerci somewhat elongate, free, with fine setulae of various lengths.
</paragraph>
<paragraph id="8BECB3566A6A650634BD4966BF5CD4CD" blockId="47.[121,749,154,1469]" pageId="47" pageNumber="190">
<emphasis id="B9276F446A6A650634BD4966BC37D4F3" box="[151,277,1387,1410]" italics="true" pageId="47" pageNumber="190">Dimensions</emphasis>
. Total length,?
<quantity id="4CAB1EB36A6A650635F04967BF4CD4F0" box="[474,622,1386,1409]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="12.0" metricValueMin="7.6" pageId="47" pageNumber="190" unit="mm" value="0.98" valueMax="1.2" valueMin="0.76">0.761.2 mm</quantity>
,! 1.0
<quantity id="4CAB1EB36A6A650636E64967BD84D4EE" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="47" pageNumber="190" unit="mm" value="1.3">1.3 mm</quantity>
; length of thorax,?
<quantity id="4CAB1EB36A6A650635AF4985BF00D4EE" box="[389,546,1416,1439]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="5.0" metricValueMin="3.3" pageId="47" pageNumber="190" unit="mm" value="0.415" valueMax="0.5" valueMin="0.33">0.330.50 mm</quantity>
,!
<quantity id="4CAB1EB36A6A650636604985BFCBD4EE" box="[586,745,1416,1439]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.2" metricValueMin="4.2" pageId="47" pageNumber="190" unit="mm" value="0.47" valueMax="0.52" valueMin="0.42">0.420.52 mm</quantity>
; length of wing,?
<quantity id="4CAB1EB36A6A6506351549A8BCEDD4CD" box="[319,463,1445,1468]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.4" metricValueMin="0.94" pageId="47" pageNumber="190" unit="mm" value="1.17" valueMax="1.4" valueMin="0.94">0.941.4 mm</quantity>
,!
<quantity id="4CAB1EB36A6A650635DD49A8BF58D4CD" box="[503,634,1445,1468]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="47" pageNumber="190" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C349E0DD6A6A6506345349D2B817D3B0" pageId="47" pageNumber="190" type="distribution">
<paragraph id="8BECB3566A6A6506345349D2BC5DD71D" blockId="47.[121,749,1503,1644]" pageId="47" pageNumber="190">
<emphasis id="B9276F446A6A6506345349D2BC26D486" bold="true" box="[121,260,1503,1527]" pageId="47" pageNumber="190">Distribution</emphasis>
.
<collectingRegion id="49977DB46A6A6506353F49EDBCFFD486" box="[277,477,1504,1527]" country="Australia" name="New South Wales" pageId="47" pageNumber="190">New South Wales</collectingRegion>
: lowlands of Colo River district, between Blue Mountains and Hunter Valley. The known distribution is very restricted, including only two spot localities c.
<quantity id="4CAB1EB36A6A650634D34A35BC0CD73E" box="[249,302,1592,1615]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="190" unit="km" value="3.0">3 km</quantity>
apart. This may change with wider use of the collection technique.
</paragraph>
<paragraph id="8BECB3566A6A650635A54A9BBCF4D7C1" blockId="47.[399,470,1686,1712]" box="[399,470,1686,1712]" pageId="47" pageNumber="190">
<emphasis id="B9276F446A6A650635A54A9BBCF4D7C1" bold="true" box="[399,470,1686,1712]" pageId="47" pageNumber="190">Notes</emphasis>
</paragraph>
<paragraph id="8BECB3566A6A650634534ADEB812D360" blockId="47.[121,749,1746,2004]" lastBlockId="47.[794,1422,154,705]" pageId="47" pageNumber="190">
<taxonomicName id="4C53C8D56A6A650634534ADEBC63D79B" authorityName="McAlpine" authorityYear="2007" box="[121,321,1747,1770]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B9276F446A6A650634534ADEBC63D79B" box="[121,321,1747,1770]" italics="true" pageId="47" pageNumber="190">Borboroides parva</emphasis>
</taxonomicName>
shares some features with species of the
<taxonomicName id="4C53C8D56A6A650634534AFDBD87D676" box="[121,165,1776,1799]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A6A650634534AFDBD87D676" box="[121,165,1776,1799]" italics="true" pageId="47" pageNumber="190">atra</emphasis>
</taxonomicName>
group, notably the long, outwardly directed anterior fronto-orbital bristle, the presence of only one dorsocentral bristle, and the strongly bowed section of vein 5 bordering the discal cell; the dull, dark coloration and the distribution of pruinescence on the head and thorax are similar conditions to those of
<taxonomicName id="4C53C8D56A6A650634FB4B8EBC0FD6EB" authorityName="McAlpine" authorityYear="2007" box="[209,301,1923,1946]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="shippi">
<emphasis id="B9276F446A6A650634FB4B8EBC0FD6EB" box="[209,301,1923,1946]" italics="true" pageId="47" pageNumber="190">B. shippi</emphasis>
</taxonomicName>
of that group. It differs from the
<taxonomicName id="4C53C8D56A6A650636544B8EBF88D6EB" box="[638,682,1923,1946]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A6A650636544B8EBF88D6EB" box="[638,682,1923,1946]" italics="true" pageId="47" pageNumber="190">atra</emphasis>
</taxonomicName>
group in the longer arista and divided abdominal sternite 1, but the large costal bristle near the humeral break, the distally open anal cell, the single anterior bristle on the mid femur, the absence of any non-terminal anterior bristle on the mid-tibia, the relatively short protandrium, the continuous sclerotization of the hypandrial plate in front of the aedeagal base, and the unusual structure of the aedeagus are unique in the genus. Some of these conditions would have been considered to represent generic characters in pre-Hennigian taxonomy, but careful consideration suggests that the species is probably phylogenetically nested among the taxa of
<taxonomicName id="4C53C8D56A6A6506312A4D88B8A4D0ED" authorityName="Malloch" authorityYear="1925" box="[1280,1414,389,412]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B9276F446A6A6506312A4D88B8A4D0ED" box="[1280,1414,389,412]" italics="true" pageId="47" pageNumber="190">Borboroides</emphasis>
</taxonomicName>
, particularly those with the distinctive few dorsally directed mesopleural setulae and curved penultimate section of vein 5 (
<emphasis id="B9276F446A6A6506371F4DD0BED4D085" box="[821,1014,477,500]" italics="true" pageId="47" pageNumber="190">
<taxonomicName id="4C53C8D56A6A6506371F4DD0BEBBD085" authorityName="McAlpine" authorityYear="2007" box="[821,921,477,500]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="staniochi">staniochi</taxonomicName>
,
<taxonomicName id="4C53C8D56A6A650637894DD0BED4D085" authorityName="McAlpine" authorityYear="2007" box="[931,1014,477,500]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="helenae">helenae</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C53C8D56A6A650630044DD0B978D085" box="[1070,1114,477,500]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A6A650630044DD0B978D085" box="[1070,1114,477,500]" italics="true" pageId="47" pageNumber="190">atra</emphasis>
</taxonomicName>
groups). Its unique character states are interpreted as probable autapomorphies.
</paragraph>
<paragraph id="8BECB3566A6A650637124E15BE9BD318" blockId="47.[794,1422,154,705]" pageId="47" pageNumber="190">
<taxonomicName id="4C53C8D56A6A650637124E15B922D35E" authorityName="McAlpine" authorityYear="2007" box="[824,1024,536,559]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B9276F446A6A650637124E15B922D35E" box="[824,1024,536,559]" italics="true" pageId="47" pageNumber="190">Borboroides parva</emphasis>
</taxonomicName>
is, on the average, the smallest insect in the genus, but there is size overlap with some species of the
<taxonomicName id="4C53C8D56A6A6506376B4E5EBE4FD31B" box="[833,877,595,618]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B9276F446A6A6506376B4E5EBE4FD31B" box="[833,877,595,618]" italics="true" pageId="47" pageNumber="190">atra</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BECB3566A6A650637124E7DB9EFD3D5" blockId="47.[794,1422,154,705]" pageId="47" pageNumber="190">All material was collected around wombat dung bait in an area with numerous wombat burrows.</paragraph>
<paragraph id="8BECB3566A6A650637124EA7B817D3B0" blockId="47.[794,1422,154,705]" box="[824,1333,682,705]" pageId="47" pageNumber="190">The specific epithet is a Latin adjective—small.</paragraph>
</subSubSection>
</treatment>
</document>