155 lines
17 KiB
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155 lines
17 KiB
XML
<document id="9F97A78ED65C5C5C0420A060BE686200" ID-DOI="10.11646/zootaxa.3869.1.3" ID-GBIF-Dataset="9e471c93-c970-4791-b652-1946027fb74c" ID-ISSN="1175-5326" ID-Zenodo-Dep="287204" ID-ZooBank="A01710FC-7462-4F22-B92E-08E39555F71B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460922332829" checkinUser="plazi" docAuthor="Wedmann, Sonja, Pouillon, Jean-Marc & Nel, André" docDate="2014" docId="313CB827FF87FFEDFF176859FD62FE86" docLanguage="en" docName="zt03869p043.pdf" docOrigin="Zootaxa 3869 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Urocerus ligniticus Piton 1940" docType="treatment" docVersion="6" lastPageNumber="37" masterDocId="CD05C05FFF83FFE8FF806D0FFFD9FFD1" masterDocTitle="New Palaeogene horntail wasps (Hymenoptera, Siricidae) and a discussion of their fossil record" masterLastPageNumber="43" masterPageNumber="33" pageNumber="37" updateTime="1698362126742" updateUser="plazi">
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<mods:titleInfo id="0E15FBA32451FACA1E63EB23A9BF5EEB">
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<mods:title id="254B094042FA860BF5805FFEC0E62E30">New Palaeogene horntail wasps (Hymenoptera, Siricidae) and a discussion of their fossil record</mods:title>
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</mods:titleInfo>
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<mods:name id="15E621345C25DBB5EFE30BD9F7D87E72" type="personal">
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<mods:roleTerm id="553E97535FA5961AAF7D1E141D834B45">Author</mods:roleTerm>
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<mods:namePart id="D11B2DD0FCEF3F324D65760D81E2AE01">Wedmann, Sonja</mods:namePart>
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<mods:namePart id="1BB628DBBC94EDD185D8A9E70F6110A6">Pouillon, Jean-Marc</mods:namePart>
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<mods:name id="AF91FD8424A7F77FC1802ACA74630E23" type="personal">
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<mods:namePart id="3C553C2FC8C7819799D8FAD5073965A2">Nel, André</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="73964E30E63E2EA1296D221FB922486C">text</mods:typeOfResource>
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<mods:title id="E0F8685059D2A265412E48410EF78E36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B34D6E4FFCE33978DC1AD0FE6DDDB9A6">
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<mods:date id="DB8DA5FB37F867AC8B18F7BA7F3758C2">2014</mods:date>
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<mods:detail id="B9C861802C0948965D897A239A358820" type="volume">
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<mods:number id="49DDBB64A2758588098E5DCE2389E4BA">3869</mods:number>
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</mods:detail>
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<mods:classification id="642D8294D4200F20396F0EB90845267F">journal article</mods:classification>
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<mods:identifier id="A7646FAFA6F49761F30372FDC6A70F43" type="DOI">10.11646/zootaxa.3869.1.3</mods:identifier>
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<mods:identifier id="4C8253EEDB0B29CF8B668C4444C1BDD1" type="GBIF-Dataset">9e471c93-c970-4791-b652-1946027fb74c</mods:identifier>
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<treatment id="313CB827FF87FFEDFF176859FD62FE86" ID-DOI="http://doi.org/10.5281/zenodo.5672358" ID-GBIF-Taxon="127670194" ID-Zenodo-Dep="5672358" LSID="urn:lsid:plazi:treatment:313CB827FF87FFEDFF176859FD62FE86" httpUri="http://treatment.plazi.org/id/313CB827FF87FFEDFF176859FD62FE86" lastPageId="5" lastPageNumber="37" pageId="4" pageNumber="37">
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<subSubSection id="F18F5ABAFF87FFECFF176859FECBFA41" pageId="4" pageNumber="37" type="nomenclature">
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<paragraph id="B92A0931FF87FFECFF176859FDCDFABF" blockId="4.[151,532,1365,1424]" box="[151,532,1365,1390]" pageId="4" pageNumber="37">
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<heading id="E262BE5DFF87FFECFF176859FDCDFABF" bold="true" box="[151,532,1365,1390]" fontSize="10" level="2" pageId="4" pageNumber="37" reason="2">
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<taxonomicName id="7E9572B2FF87FFECFF176859FDCDFABF" authority="Piton, 1940" authorityName="Piton" authorityYear="1940" box="[151,532,1365,1390]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ligniticus">
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<emphasis id="8BE1D523FF87FFECFF176859FDCDFABF" bold="true" box="[151,532,1365,1390]" pageId="4" pageNumber="37">
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<emphasis id="8BE1D523FF87FFECFF176859FEACFABF" bold="true" box="[151,373,1365,1390]" italics="true" pageId="4" pageNumber="37">Urocerus ligniticus</emphasis>
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(
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<bibRefCitation id="DD0474C0FF87FFECFE03685AFDD5FABF" author="Piton" box="[387,524,1365,1390]" pageId="4" pageNumber="37" refString="Piton, L. (1940) Paleontologie du gisement eocene de Menat (Puy-de-Dome), flore et faune. Memoire de la Societe d'Histoire Naturelle d'Auvergne, Clermont-Ferrand, 1, 1 - 303." type="journal article" year="1940">Piton, 1940</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="B92A0931FF87FFECFF176878FECBFA41" blockId="4.[151,532,1365,1424]" box="[151,274,1399,1424]" pageId="4" pageNumber="37">
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(
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<figureCitation id="21AE15B4FF87FFECFF1F6878FF37FA5E" box="[159,238,1399,1424]" captionStart="FIGURE 4" captionStartId="5.[151,249,1455,1477]" captionTargetBox="[151,1435,379,1433]" captionTargetId="figure@5.[151,1435,373,1433]" captionTargetPageId="5" captionText="FIGURE 4. Urocerus ligniticus, new additional specimen, MNHN coll. no. JMP-Menat- 801, photograph of whole fossil. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/287208/files/figure.png" pageId="4" pageNumber="37">Figs. 4</figureCitation>
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,
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<figureCitation id="21AE15B4FF87FFECFF7A6878FED1FA41" box="[250,264,1399,1424]" captionStart="FIGURE 5" captionStartId="5.[151,249,1976,1998]" captionTargetBox="[193,1405,1550,1956]" captionTargetId="figure@5.[190,1409,1546,1960]" captionTargetPageId="5" captionText="FIGURE 5. Urocerus ligniticus, new additional specimen, MNHN coll. no. JMP-Menat- 801, detail showing fore wing venation, drawing. Labelling of longitudinal veins in capital letters, cross-veins in small letters. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/287209/files/figure.png" pageId="4" pageNumber="37">5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F18F5ABAFF87FFECFF1768B5FAD4F9E7" pageId="4" pageNumber="37" type="materials_examined">
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<paragraph id="B92A0931FF87FFECFF1768B5FE9AF9C4" blockId="4.[151,1437,1466,2020]" pageId="4" pageNumber="37">
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<emphasis id="8BE1D523FF87FFECFF1768B5FEDFFA02" bold="true" box="[151,262,1466,1491]" pageId="4" pageNumber="37">Material.</emphasis>
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<typeStatus id="662EB793FF87FFECFE8D68B5FEACFA02" box="[269,373,1466,1491]" pageId="4" pageNumber="37" type="holotype">Holotype</typeStatus>
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R. 0 7004, other specimens R. 0 7722, B. 47308, all in collection of Palaeontology department,
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<collectionCode id="DF8491F4FF87FFECFF1768D3FF36FA22" LSID="urn:lsid:biocol.org:col:34988" box="[151,239,1500,1523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="37">MNHN</collectionCode>
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, Paris (isolated forewings). New specimen JMP-Menat-801 (stored in the Entomology department,
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<collectionCode id="DF8491F4FF87FFECFF1768F2FF36F9C5" LSID="urn:lsid:biocol.org:col:34988" box="[151,239,1533,1556]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="37">MNHN</collectionCode>
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, Paris).
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</paragraph>
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<paragraph id="B92A0931FF87FFECFF476B12FAD4F9E7" blockId="4.[151,1437,1466,2020]" box="[199,1293,1565,1590]" pageId="4" pageNumber="37">
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<emphasis id="8BE1D523FF87FFECFF476B12FE4CF9E7" bold="true" box="[199,405,1565,1590]" pageId="4" pageNumber="37">Age and outcrop.</emphasis>
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Palaeocene, spongo-diatomite maar Palaeolake, Menat, Puy-de-Dôme,
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<collectingCountry id="C18249A1FF87FFECFB396B10FAD0F9E7" box="[1209,1289,1567,1590]" name="France" pageId="4" pageNumber="37">France</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F18F5ABAFF87FFECFF476B4DFE33F84F" pageId="4" pageNumber="37" type="description">
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<paragraph id="B92A0931FF87FFECFF476B4DFEE8F917" blockId="4.[151,1437,1466,2020]" pageId="4" pageNumber="37">
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<emphasis id="8BE1D523FF87FFECFF476B4DFE88F98A" bold="true" box="[199,337,1602,1627]" pageId="4" pageNumber="37">Description</emphasis>
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of the new specimen JMP-Menat-801. Imprint and counterimprint of a nearly complete female wasp in ventral view. Body
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<quantity id="7E6DA4D4FF87FFECFE536B6AFDE1F9AF" box="[467,568,1637,1662]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="4" pageNumber="37" unit="mm" value="36.5">36.5 mm</quantity>
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long, Head
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<quantity id="7E6DA4D4FF87FFECFD426B6AFCC0F9AF" box="[706,793,1637,1662]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="37" unit="mm" value="4.5">4.5 mm</quantity>
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long, 6.0 mm wide; thorax 11.0 mm long, 9.0 mm wide; abdomen 21.0 mm long,
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<quantity id="7E6DA4D4FF87FFECFE2E6B85FDDDF973" box="[430,516,1674,1699]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="37" unit="mm" value="9.5">9.5 mm</quantity>
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wide; ovipositor 19.0 mm long; hindwing 20.0 mm long; forewing 28.0 mm long, 8.0 mm wide.
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</paragraph>
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<paragraph id="B92A0931FF87FFECFF476BDDFE33F84F" blockId="4.[151,1437,1466,2020]" pageId="4" pageNumber="37">
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The specimen is dorsoventrally embedded, its head is almost completely preserved, the gena has no or almost no pits in its central area. The rest of the body shows no distinct details, except for the ovipositor which is quite prominent. The semicircular rami valvularum at the base of the ovipositor are preserved, but its tip is broken, and no details are observable. The wings lie beside the body, the venation similar to those of the three specimens already attributed to
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<taxonomicName id="7E9572B2FF87FFECFE0C6A6CFCD7F8AB" authority="Piton, 1940" authorityName="Piton" authorityYear="1940" box="[396,782,1890,1915]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ligniticus">
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<emphasis id="8BE1D523FF87FFECFE0C6A6CFDB1F8AA" box="[396,616,1890,1915]" italics="true" pageId="4" pageNumber="37">Urocerus ligniticus</emphasis>
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(
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<bibRefCitation id="DD0474C0FF87FFECFDFB6A6DFCDFF8AB" author="Piton" box="[635,774,1890,1915]" pageId="4" pageNumber="37" refString="Piton, L. (1940) Paleontologie du gisement eocene de Menat (Puy-de-Dome), flore et faune. Memoire de la Societe d'Histoire Naturelle d'Auvergne, Clermont-Ferrand, 1, 1 - 303." type="journal article" year="1940">Piton, 1940</bibRefCitation>
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)
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</taxonomicName>
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, as described and figured in
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<bibRefCitation id="DD0474C0FF87FFECFBE86A6DFB30F8AB" author="Nel" box="[1128,1257,1890,1915]" pageId="4" pageNumber="37" refString="Nel, A. (1988) Redescription de Eosirex ligniticus Piton, 1940 (Hymenoptera, Symphyta, Siricidae). L'Entomologiste, 44, 287 - 292." type="journal article" year="1988">Nel (1988)</bibRefCitation>
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, with the wing membrane apically undulated.
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</paragraph>
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</subSubSection>
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<subSubSection id="F18F5ABAFF87FFEDFF476AA5FD62FE86" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37" type="discussion">
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<paragraph id="B92A0931FF87FFEDFF476AA5FD62FE86" blockId="4.[151,1437,1466,2020]" lastBlockId="5.[151,1437,151,343]" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37">
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<emphasis id="8BE1D523FF87FFECFF476AA5FE95F812" bold="true" box="[199,332,1962,1987]" pageId="4" pageNumber="37">Discussion.</emphasis>
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<collectingCountry id="C18249A1FF87FFECFED46AA5FEAAF813" box="[340,371,1962,1986]" name="American Samoa" pageId="4" pageNumber="37">As</collectingCountry>
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discussed by
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<bibRefCitation id="DD0474C0FF87FFECFD926AA5FD55F813" author="Nel" box="[530,652,1962,1987]" pageId="4" pageNumber="37" refString="Nel, A. (1988) Redescription de Eosirex ligniticus Piton, 1940 (Hymenoptera, Symphyta, Siricidae). L'Entomologiste, 44, 287 - 292." type="journal article" year="1988">Nel (1988)</bibRefCitation>
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and
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<bibRefCitation id="DD0474C0FF87FFECFD456AA5FC0EF813" author="Fidalgo" box="[709,983,1962,1987]" pageId="4" pageNumber="37" refString="Fidalgo, P. & Smith, D. R. (1987) A fossil Siricidae (Hymenoptera) from Argentinia. Entomological News, 98, 63 - 66." type="journal article" year="1987">Fidalgo & Smith (1987)</bibRefCitation>
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, the preserved character combination of the fore wing venation allows no differentiation from
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<taxonomicName id="7E9572B2FF87FFECFD746AC3FC84F832" box="[756,861,1996,2019]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="8BE1D523FF87FFECFD746AC3FC84F832" box="[756,861,1996,2019]" italics="true" pageId="4" pageNumber="37">Urocerus</emphasis>
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</taxonomicName>
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, crossvein 2r-m is present, veins Rs and M meet at the same point; crossvein 2m-cu meets M apical to where crossvein 2r-m meets M; basal stub of Cu1 absent; basal stub of 3A absent. Additionally, this fossil has one of the autapomorphies of
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<taxonomicName id="7E9572B2FF86FFEDFC706DB5FB80FF00" box="[1008,1113,186,209]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="8BE1D523FF86FFEDFC706DB5FB80FF00" box="[1008,1113,186,209]" italics="true" pageId="5" pageNumber="38">Urocerus</emphasis>
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</taxonomicName>
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given in
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<bibRefCitation id="DD0474C0FF86FFEDFB416DB6FA4EFF00" box="[1217,1431,185,210]" pageId="5" pageNumber="38" refString="Schiff, N. M., Goulet, H., Smith, D. R., Boudreault, C., Wilson, A. D. & Scheffler, B. E. (2012) Siricidae (Hymenoptera, Symphyta, Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification, 21, 1 - 305." type="journal article">
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Schiff
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<emphasis id="8BE1D523FF86FFEDFA8C6DB5FAE5FF03" box="[1292,1340,185,210]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
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. (2012)
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</bibRefCitation>
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, i.e. a gena without or almost without pits in its central area. Thus the new fossil specimen supports the attribution of
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<taxonomicName id="7E9572B2FF86FFEDFF366DF2FE98FEC4" box="[182,321,252,277]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="ligniticus">
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<emphasis id="8BE1D523FF86FFEDFF366DF2FE98FEC4" box="[182,321,252,277]" italics="true" pageId="5" pageNumber="38">U. ligniticus</emphasis>
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</taxonomicName>
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to the genus
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<taxonomicName id="7E9572B2FF86FFEDFE5C6DF2FD9CFEC5" box="[476,581,253,276]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="8BE1D523FF86FFEDFE5C6DF2FD9CFEC5" box="[476,581,253,276]" italics="true" pageId="5" pageNumber="38">Urocerus</emphasis>
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</taxonomicName>
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. All the specimens of
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<taxonomicName id="7E9572B2FF86FFEDFCC66DF2FBC6FEC4" box="[838,1055,252,277]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="ligniticus">
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<emphasis id="8BE1D523FF86FFEDFCC66DF2FBC6FEC4" box="[838,1055,252,277]" italics="true" pageId="5" pageNumber="38">Urocerus ligniticus</emphasis>
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</taxonomicName>
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have the same shape of the basal part of Rs, curved and nearly perpendicular to the radial stem and aligned with the basal part of M. Modern
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<taxonomicName id="7E9572B2FF86FFEDFF176C30FED9FE87" box="[151,256,319,342]" class="Insecta" family="Siricidae" genus="Urocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="8BE1D523FF86FFEDFF176C30FED9FE87" box="[151,256,319,342]" italics="true" pageId="5" pageNumber="38">Urocerus</emphasis>
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</taxonomicName>
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spp. have the same vein more reclined.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |