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<mods:title id="D1AC259571DEDDEB35C726644039E5C1">Taxonomy and distribution of the genus Trichomyrmex Mayr, 1865 (Hymenoptera: Formicidae) in the Arabian Peninsula, with the description of two new species</mods:title>
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<mods:namePart id="6259207E481BC1DE206E7363293B87A8">Mostafa R. Sharaf</mods:namePart>
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<mods:namePart id="CE76C89E181ABBD9DE6FF75704112B99">Shehzad Salman</mods:namePart>
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<treatment id="03F0F7622C6D974FFDE252928A07F967" ID-DOI="http://doi.org/10.5281/zenodo.6060812" ID-GBIF-Taxon="124955330" ID-Zenodo-Dep="6060812" LSID="urn:lsid:plazi:treatment:03F0F7622C6D974FFDE252928A07F967" httpUri="http://treatment.plazi.org/id/03F0F7622C6D974FFDE252928A07F967" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20">
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<paragraph id="8BE646742C6D974BFDE252928CB0FBD8" blockId="19.[586,1001,1014,1076]" box="[586,1001,1014,1041]" pageId="19" pageNumber="20">
<heading id="D0AEF1182C6D974BFDE252928CB0FBD8" box="[586,1001,1014,1041]" centered="true" fontSize="11" level="4" pageId="19" pageNumber="20" reason="4">
<taxonomicName id="4C593DF72C6D974BFDE252928CB0FBD8" authority="Forel, 1902" authorityName="Forel" authorityYear="1902" box="[586,1001,1014,1041]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B92D9A662C6D974BFDE252928C12FBD8" bold="true" box="[586,843,1014,1041]" italics="true" pageId="19" pageNumber="20">Trichomyrmex mayri</emphasis>
(
<bibRefCitation id="EFC83B852C6D974BFCF252938CB8FBD8" author="Forel A." box="[858,993,1015,1041]" journalOrPublisher="A. Diehl. Annales de la Societe Entomologique de Belgique" pageId="19" pageNumber="20" pagination="147 - 158" part="46" refString="Forel A. 1902 a. Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr. A. Diehl. Annales de la Societe Entomologique de Belgique 46: 147 - 158." title="Les fourmis du Sahara algerien recoltees par M. le Professeur A. Lameere et le Dr" type="journal article" year="1902">Forel, 1902</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974BFD7E557E8C04FBFD" box="[726,861,1050,1076]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BE646742C6D974BFD7E557E8C04FBFD" blockId="19.[586,1001,1014,1076]" box="[726,861,1050,1076]" pageId="19" pageNumber="20">
<figureCitation id="13625AF12C6D974BFD7E557E8C7CFBFD" box="[726,805,1050,1076]" captionStart="Fig. 10" captionStartId="22.[189,231,1960,1986]" captionTargetBox="[407,1180,337,1920]" captionTargetId="figure@22.[407,1180,337,1920]" captionTargetPageId="22" captionText="Fig. 10. Trichomyrmex mayri (Forel, 1902), lectotype, worker (CASENT 0249904). A. Body in profile. B. Body in dorsal view. C. Head in full-face view (photos: Shannon Hartman)." httpUri="https://zenodo.org/record/193055/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
AC
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974BFF1555388C5FFA61" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="8BE646742C6D974BFF1555388BE0FB50" blockId="19.[189,1399,1115,1177]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF1555388B45FBBC" ID-CoL="4487N" authority="Forel, 1902b: 209" authorityName="Forel" authorityPageNumber="209" authorityYear="1902" box="[189,1052,1115,1142]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="variety" species="gracillimum" subGenus="Parholcomyrmex" variety="mayri">
<emphasis id="B92D9A662C6D974BFF1555388D8CFBBC" box="[189,725,1115,1141]" italics="true" pageId="19" pageNumber="20">Monomorium (Parholcomyrmex) gracillimum</emphasis>
var.
<emphasis id="B92D9A662C6D974BFCA255388C16FBBC" box="[778,847,1116,1141]" italics="true" pageId="19" pageNumber="20">mayri</emphasis>
<bibRefCitation id="EFC83B852C6D974BFCFC553F8CB1FBBC" author="Forel A." box="[852,1000,1115,1141]" journalOrPublisher="Bulletin de la Societe Vaudoise des Sciences Naturelles" pageId="19" pageNumber="20" pagination="369 - 407" part="45" refString="Forel A. 1902 b. Myrmicinae nouveaux de l'Inde et de Ceylan. Revue Suisse de Zoologie 10: 165 - 249. Forel A. 1909. Etudes myrmecologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bulletin de la Societe Vaudoise des Sciences Naturelles 45: 369 - 407." title="Myrmicinae nouveaux de l'Inde et de Ceylan. Revue Suisse de Zoologie 10: 165 - 249. Forel A. 1909. Etudes myrmecologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis" type="journal article" year="1902" yearSuffix="b">Forel, 1902b</bibRefCitation>
: 209
</taxonomicName>
(workers). India, Indomalaya. Syntype worker [examined]
<emphasis id="B92D9A662C6D974BFDA4551B8D4FFB50" box="[524,534,1151,1177]" italics="true" pageId="19" pageNumber="20">(</emphasis>
lectotype here designated (CASENT0249904) (MHNG)).
</paragraph>
<paragraph id="8BE646742C6D974BFF1555A48D8FFB13" blockId="19.[189,726,1216,1242]" box="[189,726,1216,1242]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF1555A48D96FB13" authority="Bolton, 1987: 326" authorityName="Bolton" authorityPageNumber="326" authorityYear="1987" box="[189,719,1216,1242]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="karawajewi">
<emphasis id="B92D9A662C6D974BFF1555A48EA0FB13" box="[189,505,1216,1242]" italics="true" pageId="19" pageNumber="20">Trichomyrmex karawajewi</emphasis>
<bibRefCitation id="EFC83B852C6D974BFDA955A48DC3FB13" author="Bolton B." box="[513,666,1216,1242]" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology" pageId="19" pageNumber="20" pagination="263 - 452" part="54" publicationUrl="http://dx.doi.org/10.5281/zenodo.26850" refString="Bolton B. 1987. A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History), Entomology 54: 263 - 452. http: // dx. doi. org / 10.5281 / zenodo. 26850" title="A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae)" type="journal article" year="1987">Bolton, 1987</bibRefCitation>
: 326
</taxonomicName>
.
</paragraph>
<paragraph id="8BE646742C6D974BFF1554658DEBFAD2" blockId="19.[189,939,1281,1449]" box="[189,690,1281,1307]" pageId="19" pageNumber="20">
<emphasis id="B92D9A662C6D974BFF1554658D59FAD2" box="[189,512,1281,1307]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF1554658EB3FAD2" box="[189,490,1281,1307]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="destructor">Trichomyrmex destructor</taxonomicName>
</emphasis>
<bibRefCitation id="EFC83B852C6D974BFDAF54658DDAFAD2" author="Forel A." box="[519,643,1281,1307]" journalOrPublisher="Revue Suisse de Zoologie" pageId="19" pageNumber="20" pagination="23 - 62" part="19" refString="Forel A. 1911. Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke &amp; Waldo. Revue Suisse de Zoologie 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke &amp; Waldo" type="journal article" year="1911">Forel 1911</bibRefCitation>
: 24.
</paragraph>
<paragraph id="8BE646742C6D974BFF1554418DB9FAF6" blockId="19.[189,939,1281,1449]" box="[189,736,1316,1343]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF1554418EF3FAF7" ID-CoL="4487N" box="[189,426,1317,1342]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B92D9A662C6D974BFF1554418EF3FAF7" box="[189,426,1317,1342]" italics="true" pageId="19" pageNumber="20">Monomorium mayri</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6D974BFE6054408D36FAF6" author="Wheeler W. M." box="[456,623,1316,1343]" journalOrPublisher="P. Jacot. American Museum Novitates" pageId="19" pageNumber="20" pagination="1 - 6" part="69" refString="Wheeler W. M. 1923. Chinese ants collected by Professor S. F. Light and Professor A. P. Jacot. American Museum Novitates 69: 1 - 6." title="Chinese ants collected by Professor S. F. Light and Professor A" type="journal article" year="1923">Wheeler 1923</bibRefCitation>
: 3 (male).
</paragraph>
<paragraph id="8BE646742C6D974BFF15542D8C25FAAB" blockId="19.[189,939,1281,1449]" box="[189,892,1352,1378]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF15542D8DD6FAAB" ID-CoL="4487N" box="[189,655,1352,1378]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mayri" subGenus="Parholcomyrmex">
<emphasis id="B92D9A662C6D974BFF15542D8DD6FAAB" box="[189,655,1352,1378]" italics="true" pageId="19" pageNumber="20">Monomorium (Parholcomyrmex) mayri</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6D974BFD04542C8C19FAAB" author="Emery C." box="[684,832,1352,1378]" journalOrPublisher="Genera Insectorum" pageId="19" pageNumber="20" pagination="95 - 206" part="174" refString="Emery C. 1922.: Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae [part]. Genera Insectorum 174 B: 95 - 206, Brussels." title="Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae [part]" type="journal article" year="1922">Emery 1922</bibRefCitation>
: 180.
</paragraph>
<paragraph id="8BE646742C6D974BFF15540F8CF2FA4C" blockId="19.[189,939,1281,1449]" box="[189,939,1387,1414]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF15540F8EEDFA4C" box="[189,436,1387,1413]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B92D9A662C6D974BFF15540F8EEDFA4C" box="[189,436,1387,1413]" italics="true" pageId="19" pageNumber="20">Trichomyrmex mayri</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6D974BFE7A540F8C27FA4F" author="Ward P. S. &amp; Brady S. G. &amp; Fisher B. L. &amp; Schultz T. R." box="[466,894,1387,1414]" journalOrPublisher="Systematic Entomology" pageId="19" pageNumber="20" pagination="61 - 81" part="40" publicationUrl="http://dx.doi.org/10.1111/syen.12090" refString="Ward P. S., Brady S. G., Fisher B. L. &amp; Schultz T. R. 2015. The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology 40: 61 - 81. http: // dx. doi. org / 10.1111 / syen. 12090" title="The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae)" type="journal article" year="2015">Ward, Brady, Fisher &amp; Schultz 2015</bibRefCitation>
: 16.
</paragraph>
<paragraph id="8BE646742C6D974BFF1554EA8C5FFA61" blockId="19.[189,939,1281,1449]" box="[189,774,1422,1449]" pageId="19" pageNumber="20">
<taxonomicName id="4C593DF72C6D974BFF1554EA8EB3FA61" box="[189,490,1422,1448]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="destructor">
<emphasis id="B92D9A662C6D974BFF1554EA8EB3FA61" box="[189,490,1422,1448]" italics="true" pageId="19" pageNumber="20">Trichomyrmex destructor</emphasis>
</taxonomicName>
<bibRefCitation id="EFC83B852C6D974BFDAF54EA8D90FA61" author="Viehmeyer H." box="[519,713,1422,1448]" journalOrPublisher="Archiv fur Naturgeschichte (A" pageId="19" pageNumber="20" pagination="108 - 168" part="81" refString="Viehmeyer H. 1916. Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte (A) 81 (8): 108 - 168." title="Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck" type="journal article" year="1916">Viehmeyer 1916</bibRefCitation>
: 132.
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C6D974FFF1554AB8DB1FD08" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BE646742C6D974BFF1554AB8EF2FA20" blockId="19.[189,427,1487,1513]" box="[189,427,1487,1513]" pageId="19" pageNumber="20">
<heading id="D0AEF1182C6D974BFF1554AB8EF2FA20" bold="true" box="[189,427,1487,1513]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="B92D9A662C6D974BFF1554AB8EF2FA20" bold="true" box="[189,427,1487,1513]" pageId="19" pageNumber="20">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C6D974BFF1557758A2EF9E2" blockId="19.[189,1399,1553,1580]" box="[189,1399,1553,1580]" pageId="19" pageNumber="20">QATAR: 1 w, Doha, Ben Mahmoud, 25.28˚ N, 51.52˚ E, 14 Mar. 2005, M.S. Abdel-Dayem leg. (KSMA).</paragraph>
<paragraph id="8BE646742C6D974DFF1557368DF9F8C6" blockId="19.[189,1399,1618,1997]" lastBlockId="21.[189,1399,267,1807]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
SAUDI ARABIA: 8 w, Al Bahah Province, Raghadan forest, 20.00˚ N, 41.44˚ E, alt. 2222 m, 13 May 2010, M.R. Sharaf leg.; 17 w, Park, Beljurashi, 19.80˚ N, 41.71˚ E, alt. 2222 m, 17 May 2010, M.R. Sharaf leg.; 12 w, Elqama Park, Beljurashi, 19.80˚ N, 41.71˚ E, alt. 1950 m, 21 Sep. 2011, M.R. Sharaf leg.; 10 w, Amadan forest, Al Mandaq, 20.20˚ N, 41.23˚ E, alt. 1881 m, 19 May 2010, M.R. Sharaf leg.; 2 w, Wadi Elzaraeb (w. Kheir), 20.06˚ N, 41.38˚ E, alt. 2123 m, 15 May 2010, M.R. Sharaf leg.; 5 w, Wadi Elzaraeb (w. Kheir), 20.07˚ N, 41.38˚ E, alt. 2086 m, 9 May 2011, M.R. Sharaf leg.; 20 w, Shohba forest, 20.04˚ N, 41.47˚ E, alt. 2324 m, 14 May 2010, M.R. Sharaf leg.; 2 w, Wadi Turabah, 20.24˚ N, 41.26˚ E, alt. 1751 m, 19 Sep. 2011, M.R. Sharaf leg. — 2 w, Asir Province, Raydah, 18.19˚ N, 42.39˚ E, alt. 1897 m, 21 Feb. 2014, Aldhafer
<emphasis id="B92D9A662C6D974BFED956098EF5F84F" box="[369,428,1900,1926]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
leg.; 2 w, Raydah, 18.19˚ N, 42.39˚ E, alt. 1772 m, 29 Jan. 2015, Aldhafer
<emphasis id="B92D9A662C6D974BFAAC56098A67F84F" box="[1284,1342,1900,1926]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
leg.; 4 w, Raydah, 18.19˚ N, 42.40˚ E, alt. 2285 m, 29 Jan. 2015, Aldhafer
<emphasis id="B92D9A662C6D974BFC4456F48B71F860" box="[1004,1064,1935,1961]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
leg.; 2 w, Raydah, 18.19˚ N, 42.39˚ E, alt. 1897 m, 29 Jan. 2015, Aldhafer
<emphasis id="B92D9A662C6D974BFD5D56D78C6DF805" box="[757,820,1970,1996]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
leg.; 1 w, Almajardah, wadi Bagara, 18.79˚ N, 4 2.01˚ E, alt. 641 m, 10 Nov. 2012, M.R. Sharaf leg.; Dalaghan, 18.10˚ N, 42.70˚ E, alt. 2100 m, 17 Apr. 2008, M.R. Sharaf leg. — 2 w, Eastern Province, Al Qatif, 26.50˚ N, 50.01˚ E, alt. 3 m, 21 Dec. 2013, M.R. Sharaf leg. — 5 w, Jazan Province, Abu Arish, 17.01˚ N, 42.80˚ E, alt. 90 m, 10 Apr. 2012, M.R. Sharaf leg.; 3 w, Jazan, 16.97˚ N, 42.61˚ E, alt. 38 m, 12 Apr. 2012, M.R. Sharaf leg.; 1 w, Jazan, Sabia, 17.10˚ N, 42.65˚ E, alt. 43 m, 9 Apr. 2012, M.R. Sharaf leg. — 20 w, Makkah Province, Wadi Gonouna, 19.42˚ N, 41.60˚ E, alt. 353 m, 14 May 2010, M.R. Sharaf leg.; 11 w, Allaith gov., Adam, wadi Elarj, 20.45˚ N, 40.81˚ E, alt. 450 m, 9 Nov. 2012, M.R. Sharaf leg. — 4 w, Riyadh Province, Al Dawadmi, 24.46˚ N, 44.34˚ E, alt. 1038 m, 18 Apr. 2014, S. Salman leg.; 12 w, Al Ammariyah, 24.81˚ N, 46.44˚ E, alt. 696 m, 12 Oct. 2013, S. Salman leg.; 7 w, Al Ghat, 26.06660˚ N, 44.91957˚ E, alt. 675 m, 7 Feb. 2015, S. Salman leg.; 1 w, Al Kharaj, 24.15˚ N, 47.23˚ E, alt. 463 m, 19 Jan. 2014, S. Salman leg.; 2 w, Kharaj, 24.29˚ N, 47.15˚ E, alt. 453 m, 8 Sep. 2014, S. Salman leg.; 5 w, Dhurma, 24.69˚ N, 46.12˚ E, alt. 649 m, 30 Jan. 2015, S. Salman leg.; 17 w, Al Majmaah, 26.00˚ N, 45.01˚ E, alt. 594 m, 13 Sep. 2014, S. Salman leg.; 1 w, Al Majmaah, 25.93˚ N, 45.29˚ E, alt. 736 m, 5 Jun. 2014, S. Salman leg.; 13 w, Rawdhat Khorim, 25.38˚ N, 47.27˚ E, alt. 551 m, 16 Jan. 2015, S. Salman leg.; 1 w, Rawdhat Khorim, 25.38˚N, 47.28˚ E, alt. 618 m, 2 Jun. 2013, S. Salman leg.; 26 w, Rawdhat Khorim, 25.38˚ N, 47.27˚ E, alt. 618 m, 2 Jun.2013, S. Salman leg.; 6 w, Layla, 20.49˚ N, 44.79˚ E, alt. 721 m, 19 Feb. 2015, S. Salman leg.; 3 w, Layla, 22.28˚ N, 46.74˚ E, alt. 558 m, 19 Feb. 2015, S. Salman leg.; 7 w, Ergah, 24.67˚ N, 46.59˚ E, 625 m, 19 Jan. 2015, S. Salman leg.; 6 w, Ergah, 24.67˚ N, 46.59˚ E, 625 m, 1 Apr. 2015, S. Salman leg.; 1 w, Diriyah, 24.76˚ N, 47.55˚ E, alt. 673 m, 9 Oct. 2013, S. Salman leg.; 1 w, Dirab, 24.41˚ N, 46.65˚ E, alt. 604 m, 18 Sep. 2014, S. Salman leg.; 2 w, Hawtet Bani Tamim, 23.45˚ N, 46.81˚ E, alt. 582 m, 13 Dec. 2014, S. Salman leg.; 2 w, Huraymila, 25.13˚ N, 46.13˚ E, alt. 783 m, 15 Sep. 2014, S. Salman leg.; 8 w, Kharrarah, 24.39˚ N, 46.24˚ E, alt. 726 m, 8 Apr. 2014, S. Salman leg.; 9 w, Mezahmiyah, 24.47˚ N, 46.23˚ E, alt. 633 m, 25 Jan. 2014, S. Salman leg.; 1 w, Mezahmiyah, 24.46˚ N, 46.22˚ E, alt. 641 m, 25 Jan. 2014, S. Salman leg.; 6 w, Mezahmiyah, 24.48˚ N, 46.26˚ E, alt. 648 m, 25 Jan. 2014, S. Salman leg.; 1 w, Naajan, 24.06˚ N, 47.17˚ E, alt. 513 m, 13 Dec. 2014, S. Salman leg.; 3 w, Naajan, 24.02˚ N, 47.13˚ E, alt. 467 m, 13 Dec. 2014, S. Salman leg.; 5 w, Sajir, 25.06˚ N, 44.69˚ E, alt. 751 m, 23 Jan. 2015, S. Salman leg.; 1 w, Salboukh, 25.07˚ N, 46.34˚ E, alt. 716 m, 26 Dec. 2014, S. Salman leg.; 12 w, Shagra, 25.26˚ N, 45.26˚ E, alt. 728 m, 30 May 2014, S. Salman leg.; 5 w, Shagra, Al Qarain, 25.23˚ N, 45.31˚ E, alt. 703 m, 24 Jan. 2015, S. Salman leg.; 5 w, Buraydah, 26.32˚ N, 44.02˚ E, alt. 643 m, 19 Oct. 2013, S. Salman leg.; 15 w, Thadiq, 25.29˚ N, 45.87˚ E, alt. 735 m, 26 Apr. 2014, S. Salman leg.; 5 w, Wadi Al Dawasir, 20.49˚ N, 44.76˚ E, alt. 706 m, 22 Jan. 2014, S. Salman leg.; 3 w, Wadi Al Dawasir, 20.45˚ N, 44.86˚ E, alt. 680 m, 20 Feb. 2015, S. Salman leg.; 3 w, Wadi Al Dawasir, 20.49˚ N, 44.79˚ E, alt. 721 m, 22 Feb. 2015, S. Salman leg.; 11 w, Wadi Hanifa, 25.77˚ N, 47.52˚ E, alt. 690 m, 9 Jan. 2013, S. Salman leg.; 1 w, Wadi Hanifa, 24.74˚ N, 46.56˚ E, alt. 673 m, 18 Sep. 2014, S. Salman leg.; 2 w, Wadi Hanifa, 24.67˚ N, 46.58˚ E, alt. 654 m, 14 Feb. 2014, S. Salman leg.; 2 w, Wadi Hanifa, 24.86˚ N, 46.45˚ E, alt. 695 m, 21 Jan. 2015, S. Salman leg.; 6 w, Wadi Hanifa, 24.54˚ N, 46.74˚ E, alt. 647 m, 11 Apr. 2014, S. Salman leg.; 1 w, Deryia, 24.76˚ N, 46.55˚ E, alt. 673 m, 9 Oct. 2013, S. Salman leg.; 1 w, Dirab, 24.41˚ N, 46.65˚ E, alt. 604 m, 18 Sep. 2014, S. Salman leg.; 18 w, Oyaina, 24.90˚ N, 46.38˚ E, alt. 749 m, 28 Apr. 2010, M.R. Sharaf leg.; 11 w, Wadi Hanifa, 24.67˚ N, 46.59˚ E, alt. 641 m, 11 Apr. 2013, M.R. Sharaf leg.; 9 w, Al Ammariyah, 24.81˚ N, 46.44˚ E, alt. 630 m, 21 Jan. 2010, M.R. Sharaf leg.; 6 w, Al Ammariyah, 24.81˚ N, 46.44˚ E, alt. 696 m, 12 Oct. 2013, M.R. Sharaf leg.; 10 w, Salboukh, 25.07˚ N, 46.34˚ E, alt. 689 m, 5 Nov. 2009, M.R. Sharaf leg.; 4 w, Kharaj, 24.19˚ N, 47.23˚ E, alt. 465 m, 19 Jan. 2014, M.R. Sharaf leg.; 1 w, King Saud University Campus, 24.71˚ N, 46.62˚ E, alt. 660 m, 9 Jul. 2009, M.R. Sharaf leg. (all material in KSMA).
</paragraph>
<caption id="DF2616FC2C6A974CFF1556CE8B0CF821" httpUri="https://zenodo.org/record/193054/files/figure.png" pageId="20" pageNumber="21" targetBox="[433,1155,340,1923]" targetPageId="20">
<paragraph id="8BE646742C6A974CFF1556CE8B0CF821" blockId="20.[189,1399,1962,2024]" pageId="20" pageNumber="21">
<emphasis id="B92D9A662C6A974CFF1556CE8E51F80D" bold="true" box="[189,264,1962,1988]" pageId="20" pageNumber="21">Fig. 9.</emphasis>
<taxonomicName id="4C593DF72C6A974CFEBA56CE8D9FF80C" authority="Forel, 1902" authorityName="Forel" authorityYear="1902" box="[274,710,1962,1989]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lameerei">
<emphasis id="B92D9A662C6A974CFEBA56CE8D7EF80D" box="[274,551,1962,1988]" italics="true" pageId="20" pageNumber="21">Trichomyrmex lameerei</emphasis>
(Forel, 1902)
</taxonomicName>
, lectotype, worker (CASENT0249871).
<emphasis id="B92D9A662C6A974CFB3256CE8BF6F80D" bold="true" box="[1178,1199,1962,1988]" pageId="20" pageNumber="21">A</emphasis>
. Body in profile.
<emphasis id="B92D9A662C6A974CFF1556AA8F89F821" bold="true" box="[189,208,1998,2024]" pageId="20" pageNumber="21">B</emphasis>
. Body in dorsal view.
<emphasis id="B92D9A662C6A974CFE7B56A98EB1F82E" bold="true" box="[467,488,1997,2023]" pageId="20" pageNumber="21">C</emphasis>
. Head in full-face view (photos: Shannon Hartman).
</paragraph>
</caption>
<paragraph id="8BE646742C6B974DFF1556538DFAF872" blockId="21.[189,1398,1846,1979]" pageId="21" pageNumber="22">
UNITED ARAB EMIRATES: 1 w, Wadi Wurayah farm, 25.23˚ N, 56.19˚ E, 2 Jun.16 Jul. 2009, M. Hauser
<emphasis id="B92D9A662C6B974DFEEE563E8EDDF8BD" box="[326,388,1882,1908]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
leg. (CASENT0265232); 1 w, Jebel Hafit, 24.03˚ N, 55.46˚ E, 27 Feb.3 Mar. 2011, M. Hauser
<emphasis id="B92D9A662C6B974DFEE1561A8ED1F85E" box="[329,392,1917,1943]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
leg. (CASENT0261837); 1 w, Wadi Safad, 25.13˚N, 56.19˚E, 4 May15 Jul. 2006, M. Hauser
<emphasis id="B92D9A662C6B974DFEE856C58E25F873" box="[320,380,1952,1978]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
leg. (CASENT0264937).
</paragraph>
<caption id="DF2616FC2C68974EFF1556CC8B0CF82F" httpUri="https://zenodo.org/record/193055/files/figure.png" pageId="22" pageNumber="23" targetBox="[407,1180,337,1920]" targetPageId="22">
<paragraph id="8BE646742C68974EFF1556CC8B0CF82F" blockId="22.[189,1399,1960,2022]" pageId="22" pageNumber="23">
<emphasis id="B92D9A662C68974EFF1556CC8E41F80B" bold="true" box="[189,280,1960,1986]" pageId="22" pageNumber="23">Fig. 10.</emphasis>
<taxonomicName id="4C593DF72C68974EFE8B56CC8DE3F80A" authority="Forel, 1902" authorityName="Forel" authorityYear="1902" box="[291,698,1960,1987]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B92D9A662C68974EFE8B56CC8D41F80B" box="[291,536,1960,1986]" italics="true" pageId="22" pageNumber="23">Trichomyrmex mayri</emphasis>
(Forel, 1902)
</taxonomicName>
, lectotype, worker (CASENT0249904).
<emphasis id="B92D9A662C68974EFB3F56CC8BF5F80B" bold="true" box="[1175,1196,1960,1986]" pageId="22" pageNumber="23">A</emphasis>
. Body in profile.
<emphasis id="B92D9A662C68974EFF1556A88F89F82F" bold="true" box="[189,208,1996,2022]" pageId="22" pageNumber="23">B</emphasis>
. Body in dorsal view.
<emphasis id="B92D9A662C68974EFE7B56AF8EB1F82C" bold="true" box="[467,488,1995,2021]" pageId="22" pageNumber="23">C</emphasis>
. Head in full-face view (photos: Shannon Hartman).
</paragraph>
</caption>
<paragraph id="8BE646742C69974FFF1550748B7EFEE0" blockId="23.[189,1063,271,298]" box="[189,1063,271,298]" pageId="23" pageNumber="24">YEMEN: 1 w, Taizz, 13.57˚ N, 44.02˚ E, 2628 Jul. 1999, A.V. Harten leg.</paragraph>
<paragraph id="8BE646742C69974FFF4250348EE5FEA2" blockId="23.[189,1399,336,504]" box="[234,444,336,363]" pageId="23" pageNumber="24">
<heading id="D0AEF1182C69974FFF4250348EE5FEA2" bold="true" box="[234,444,336,363]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="6">
<emphasis id="B92D9A662C69974FFF4250348EE5FEA2" bold="true" box="[234,444,336,363]" pageId="23" pageNumber="24">Previous records</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C69974FFF1550108A51FE31" blockId="23.[189,1399,336,504]" pageId="23" pageNumber="24">
SAUDI ARABIA: 1 w, Wadi Khumra, 21.15˚ N, 41.35˚ E, 10 Feb. 1978, C.A. Collingwood leg.; 1 w, Al Mandaq, 20.20˚ N, 41.23˚ E, 8 Apr. 1980, C.A. Collingwood leg.; 1 w, Al Tawala, 25.5˚ N, 37.28˚ E, 7 Apr. 1983, C.A. Collingwood leg.; 1 w, Bisha, 20˚ N, 42.6˚ E, 7 Apr. 1983, C.A. Collingwood leg.; 1 w, Al Qatif, 26.51˚ N, 49.96˚ E, alt. 30 m, 14 Apr. 1983, C.A. Collingwood leg. (
<bibRefCitation id="EFC83B852C69974FFBB550B98BA2FE31" author="Collingwood C. A." box="[1053,1275,477,504]" journalOrPublisher="Formicidae of Saudi Arabia. Fauna of Saudi Arabia" pageId="23" pageNumber="24" pagination="230 - 302" part="7" refString="Collingwood C. A. 1985. Hymenoptera: Fam. Formicidae of Saudi Arabia. Fauna of Saudi Arabia 7: 230 - 302." title="Hymenoptera: Fam" type="journal article" year="1985">Collingwood 1985</bibRefCitation>
).
</paragraph>
<paragraph id="8BE646742C69974FFF15537B8BA9FD95" blockId="23.[189,1399,542,605]" pageId="23" pageNumber="24">
UNITED ARAB EMIRATES: 1 w Al-Jazira Al-Hamra, 25.70˚ N, 55.79˚ E, 9 Sep. 2004, A.V. Harten leg.; 1 w, Ruwais, 24.15˚ N, 52.73˚ E, Mar. 1995, A.V. Harten leg. (
<bibRefCitation id="EFC83B852C69974FFC6F53268BBDFD95" author="Collingwood C. A. &amp; Agosti D. &amp; Sharaf M. R. &amp; Harten A." box="[967,1252,578,604]" journalOrPublisher="Arthropod Fauna of the United Arab Emirates" pageId="23" pageNumber="24" pagination="405 - 474" part="4" refString="Collingwood C. A., Agosti D., Sharaf M. R. &amp; Harten A. 2011. Order Hymenoptera, Family Formicidae. Arthropod Fauna of the United Arab Emirates 4: 405 - 474." title="Order Hymenoptera, Family Formicidae" type="journal article" year="2011">
Collingwood
<emphasis id="B92D9A662C69974FFBC153278BFDFD95" box="[1129,1188,578,604]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<paragraph id="8BE646742C69974FFF1553E08DB1FD08" blockId="23.[189,1398,643,705]" pageId="23" pageNumber="24">
YEMEN: 1 w, Sana a, 15.34˚ N, 44.20˚ E, Jan. 1991Nov. 1991; w, Aden, 12.78˚ N, 44.95˚ E, Dec. 1991, A.V. Harten leg. (
<bibRefCitation id="EFC83B852C69974FFE2153C38D82FD08" author="Collingwood C. A. &amp; Agosti D." box="[393,731,678,705]" journalOrPublisher="Fauna of Saudi Arabia" pageId="23" pageNumber="24" pagination="300 - 385" part="15" refString="Collingwood C. A. &amp; Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2). Fauna of Saudi Arabia 15: 300 - 385." title="Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2)" type="journal article" year="1996">Collingwood &amp; Agosti 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C69974FFF15538C8DD4FCF8" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BE646742C69974FFF15538C8E95FCCB" blockId="23.[189,460,744,770]" box="[189,460,744,770]" pageId="23" pageNumber="24">
<heading id="D0AEF1182C69974FFF15538C8E95FCCB" bold="true" box="[189,460,744,770]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<emphasis id="B92D9A662C69974FFF15538C8E95FCCB" bold="true" box="[189,460,744,770]" pageId="23" pageNumber="24">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C69974FFF1552738DD4FCF8" blockId="23.[189,653,791,817]" box="[189,653,791,817]" pageId="23" pageNumber="24">
Separation is given under
<taxonomicName id="4C593DF72C69974FFE5952738DDEFCF8" box="[497,647,791,817]" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="destructor">
<emphasis id="B92D9A662C69974FFE5952738DDEFCF8" box="[497,647,791,817]" italics="true" pageId="23" pageNumber="24">T. destructor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C69974FFF15523C8B7EFA47" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph id="8BE646742C69974FFF15523C8D67FCBB" blockId="23.[189,574,856,882]" box="[189,574,856,882]" pageId="23" pageNumber="24">
<heading id="D0AEF1182C69974FFF15523C8D67FCBB" bold="true" box="[189,574,856,882]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<emphasis id="B92D9A662C69974FFF15523C8D67FCBB" bold="true" box="[189,574,856,882]" pageId="23" pageNumber="24">Ecological and biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C69974FFF1552E38B7EFA47" blockId="23.[189,1399,903,1422]" pageId="23" pageNumber="24">
A nest series was found under a
<taxonomicName id="4C593DF72C69974FFDE152E38B12FC68" authority="(Aiton) W.T.Aiton 1811" authorityName="(Aiton) W.T.Aiton" authorityYear="1811" box="[585,1099,903,930]" class="Magnoliopsida" family="Apocynaceae" genus="Calotropis" kingdom="Plantae" order="Gentianales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="procera">
<emphasis id="B92D9A662C69974FFDE152E38C77FC69" box="[585,814,903,929]" italics="true" pageId="23" pageNumber="24">Calotropis procera</emphasis>
(Aiton) W.T.Aiton 1811
</taxonomicName>
tree and coexisting with several ant species including:
<taxonomicName id="4C593DF72C69974FFDBD52CE8BD1FC0D" authority="Collingwood &amp; Van Harten, 2001" authorityName="Collingwood &amp; Van Harten" authorityYear="2001" box="[533,1160,938,964]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="B92D9A662C69974FFDBD52CE8DB0FC0D" box="[533,745,938,964]" italics="true" pageId="23" pageNumber="24">Carebara arabica</emphasis>
(Collingwood &amp; Van Harten, 2001)
</taxonomicName>
,
<taxonomicName id="4C593DF72C69974FFB3A52CF8EDAFC2E" authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="melanocephalum">
<emphasis id="B92D9A662C69974FFB3A52CF8A2EFC0D" box="[1170,1399,938,964]" italics="true" pageId="23" pageNumber="24">T. melanocephalum</emphasis>
(Fabricius, 1793)
</taxonomicName>
,
<taxonomicName id="4C593DF72C69974FFE2652AA8DAEFC2E" authority="Mayr, 1904" authorityName="Mayr" authorityYear="1904" box="[398,759,973,999]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="jaegerskioeldi">
<emphasis id="B92D9A662C69974FFE2652AA8D0FFC2E" box="[398,598,973,999]" italics="true" pageId="23" pageNumber="24">N. jaegerskioeldi</emphasis>
(Mayr, 1904)
</taxonomicName>
,
<taxonomicName id="4C593DF72C69974FFCAA52AA8CFAFC2E" box="[770,931,974,999]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B92D9A662C69974FFCAA52AA8CFAFC2E" box="[770,931,974,999]" italics="true" pageId="23" pageNumber="24">Monomorium</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="4C593DF72C69974FFBA952A98BEBFC2E" box="[1025,1202,973,999]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B92D9A662C69974FFBA952A98BEBFC2E" box="[1025,1202,973,999]" italics="true" pageId="23" pageNumber="24">Cardiocondyla</emphasis>
</taxonomicName>
sp. Another nest series was found under a stone next to a
<taxonomicName id="4C593DF72C69974FFD1752958C00FBC3" box="[703,857,1008,1035]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="limon">
<emphasis id="B92D9A662C69974FFD1752958C00FBC3" box="[703,857,1008,1035]" italics="true" pageId="23" pageNumber="24">Citrus limon</emphasis>
</taxonomicName>
(L.) Burm. F. (
<taxonomicName id="4C593DF72C69974FFBB252948BD0FBC3" box="[1050,1161,1008,1034]" class="Magnoliopsida" family="Rutaceae" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Rutaceae</taxonomicName>
) tree and a nest of
<taxonomicName id="4C593DF72C69974FFF1555778DEEFBE7" authority="Mayr, 1862" authorityName="Mayr" authorityYear="1862" box="[189,695,1043,1070]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sennaarensis">
<emphasis id="B92D9A662C69974FFF1555778D55FBE4" box="[189,524,1043,1069]" italics="true" pageId="23" pageNumber="24">Brachyponera sennaarensis</emphasis>
(Mayr, 1862)
</taxonomicName>
. Many workers were foraging on moist clay soil under a mango tree,
<taxonomicName id="4C593DF72C69974FFEFD55538D7CFB98" box="[341,549,1079,1105]" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="B92D9A662C69974FFEFD55538D7CFB98" box="[341,549,1079,1105]" italics="true" pageId="23" pageNumber="24">Mangifera indica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C593DF72C69974FFDF255538C54FB9B" box="[602,781,1079,1106]" class="Magnoliopsida" family="Anacardiaceae" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
). Some other workers were collected from dry leaf litter under a
<taxonomicName id="4C593DF72C69974FFEF1553F8C31FBBC" authority="R.Br." authorityName="R.Br." box="[345,872,1114,1141]" class="Magnoliopsida" family="Scrophulariaceae" genus="Myoporum" kingdom="Plantae" order="Lamiales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="insulare">
<emphasis id="B92D9A662C69974FFEF1553F8D1BFBBD" box="[345,578,1114,1140]" italics="true" pageId="23" pageNumber="24">Myoporum insulare</emphasis>
R.Br. (Scrophulariaceae)
</taxonomicName>
tree. A nest was found under a stone in moist soil and next to a
<taxonomicName id="4C593DF72C69974FFE22551A8D9DFB5E" authority="Hochst." authorityName="Hochst." box="[394,708,1149,1175]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="procera">
<emphasis id="B92D9A662C69974FFE22551A8D3AFB5E" box="[394,611,1150,1175]" italics="true" pageId="23" pageNumber="24">Juniperus procera</emphasis>
Hochst.
</taxonomicName>
ex. Endlicher (
<taxonomicName id="4C593DF72C69974FFCDC55198B4FFB5E" box="[884,1046,1149,1175]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
) in the Asir Mountains. Some workers were foraging in leaf litter under a
<taxonomicName id="4C593DF72C69974FFD7C55C48CC4FB73" box="[724,925,1184,1210]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="guajava">
<emphasis id="B92D9A662C69974FFD7C55C48CC4FB73" box="[724,925,1184,1210]" italics="true" pageId="23" pageNumber="24">Psidium guajava</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C593DF72C69974FFC7B55C48B0DFB73" box="[979,1108,1184,1210]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) tree. Another nest was found in moist soil, rich in decayed organic matter and the surrounding area of dense grasses. Many workers were collected by sifting the leaf litter under date palms. In Shada Al Ala, a nest was found under a discarded bag filled with soil rich in decaying organic matter under a coffee tree,
<taxonomicName id="4C593DF72C69974FFB6F546E8A2FFAED" box="[1223,1398,1290,1316]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="B92D9A662C69974FFB6F546E8A2FFAED" box="[1223,1398,1290,1316]" italics="true" pageId="23" pageNumber="24">Coffea arabica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C593DF72C69974FFF4C54498E3DFA81" box="[228,356,1325,1352]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
). Several workers were found under the bark of
<taxonomicName id="4C593DF72C69974FFC2D54498C8FFA8E" box="[901,982,1325,1351]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B92D9A662C69974FFC2D54498C8FFA8E" box="[901,982,1325,1351]" italics="true" pageId="23" pageNumber="24">Acacia</emphasis>
</taxonomicName>
sp. trees. A nest series was foraging under a pomegranate tree,
<taxonomicName id="4C593DF72C69974FFE5B54358D99FAA3" box="[499,704,1361,1386]" class="Magnoliopsida" family="Lythraceae" genus="Punica" kingdom="Plantae" order="Myrtales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="granatum">
<emphasis id="B92D9A662C69974FFE5B54358D99FAA3" box="[499,704,1361,1386]" italics="true" pageId="23" pageNumber="24">Punica granatum</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C593DF72C69974FFD4554348C2FFAA2" box="[749,886,1360,1387]" class="Magnoliopsida" family="Lythraceae" kingdom="Plantae" order="Myrtales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Lythraceae</taxonomicName>
). Some workers were collected under a ficus tree (
<taxonomicName id="4C593DF72C69974FFF5154108E29FA47" box="[249,368,1396,1422]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
) where soil was moist and rich in decaying organic matter.
</paragraph>
</subSubSection>
<subSubSection id="C34315FF2C69974FFF1554D18A07F967" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BE646742C69974FFF1554D18E0EFA06" blockId="23.[189,343,1461,1487]" box="[189,343,1461,1487]" pageId="23" pageNumber="24">
<heading id="D0AEF1182C69974FFF1554D18E0EFA06" bold="true" box="[189,343,1461,1487]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<emphasis id="B92D9A662C69974FFF1554D18E0EFA06" bold="true" box="[189,343,1461,1487]" pageId="23" pageNumber="24">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE646742C69974FFF1554878A07F967" blockId="23.[189,1399,1507,1710]" pageId="23" pageNumber="24">
This species has successfully invaded many regions of the tropics, with a speculated origin in the Indian subcontinent (
<bibRefCitation id="EFC83B852C69974FFECC57638EA2F9E8" author="Bolton B." box="[356,507,1543,1569]" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology" pageId="23" pageNumber="24" pagination="263 - 452" part="54" publicationUrl="http://dx.doi.org/10.5281/zenodo.26850" refString="Bolton B. 1987. A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History), Entomology 54: 263 - 452. http: // dx. doi. org / 10.5281 / zenodo. 26850" title="A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae)" type="journal article" year="1987">Bolton 1987</bibRefCitation>
). Geographically, this species has spread westward to the Middle East, the Arabian Peninsula (
<bibRefCitation id="EFC83B852C69974FFE00574E8DDFF98C" author="Collingwood C. A." box="[424,646,1578,1605]" journalOrPublisher="Formicidae of Saudi Arabia. Fauna of Saudi Arabia" pageId="23" pageNumber="24" pagination="230 - 302" part="7" refString="Collingwood C. A. 1985. Hymenoptera: Fam. Formicidae of Saudi Arabia. Fauna of Saudi Arabia 7: 230 - 302." title="Hymenoptera: Fam" type="journal article" year="1985">Collingwood 1985</bibRefCitation>
;
<bibRefCitation id="EFC83B852C69974FFD3B574E8CBFF98D" author="Collingwood C. A. &amp; Agosti D." box="[659,998,1578,1604]" journalOrPublisher="Fauna of Saudi Arabia" pageId="23" pageNumber="24" pagination="300 - 385" part="15" refString="Collingwood C. A. &amp; Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2). Fauna of Saudi Arabia 15: 300 - 385." title="Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2)" type="journal article" year="1996">Collingwood &amp; Agosti 1996</bibRefCitation>
; Sharaf
<emphasis id="B92D9A662C69974FFBE1574F8BDCF98D" box="[1097,1157,1578,1604]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2013), North Africa (Egypt) (
<bibRefCitation id="EFC83B852C69974FFE8D57298EEEF9A1" author="Sharaf M. R." box="[293,439,1613,1640]" journalOrPublisher="Ain Shams University, Faculty of Science, Entomology Department, Cairo [unpublished thesis]" pageId="23" pageNumber="24" refString="Sharaf M. R. 2006. Taxonomic and Ecological Studies on Family Formicidae (Order: Hymenoptera) in Egypt including some Protectorates with a Study of some Insect Fauna associated with Ant Species. Ain Shams University, Faculty of Science, Entomology Department, Cairo [unpublished thesis]." title="Taxonomic and Ecological Studies on Family Formicidae (Order: Hymenoptera) in Egypt including some Protectorates with a Study of some Insect Fauna associated with Ant Species" type="book" year="2006">Sharaf 2006</bibRefCitation>
) and along the coastal zones of Sub-Saharan Africa (
<bibRefCitation id="EFC83B852C69974FFB8C57298BE0F9AE" author="Bolton B." box="[1060,1209,1613,1639]" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology" pageId="23" pageNumber="24" pagination="263 - 452" part="54" publicationUrl="http://dx.doi.org/10.5281/zenodo.26850" refString="Bolton B. 1987. A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History), Entomology 54: 263 - 452. http: // dx. doi. org / 10.5281 / zenodo. 26850" title="A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae)" type="journal article" year="1987">Bolton 1987</bibRefCitation>
).
<taxonomicName id="4C593DF72C69974FFB6357298E5AF943" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B92D9A662C69974FFB6357298E5AF943" italics="true" pageId="23" pageNumber="24">Trichomyrmex mayri</emphasis>
</taxonomicName>
is the most successful and widely distributed species of the genus in the Arabian Peninsula and one of the most abundant ants of the Socotra Archipelago (
<bibRefCitation id="EFC83B852C69974FFCD957F78A08F967" author="Collingwood C. A. &amp; Pohl H. &amp; Gusten R. &amp; Wranik W. &amp; Harten A." box="[881,1361,1683,1710]" journalOrPublisher="Fauna of Arabia" pageId="23" pageNumber="24" pagination="473 - 495" part="20" refString="Collingwood C. A., Pohl H., Gusten R., Wranik W. &amp; Harten A. 2004. The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago. Fauna of Arabia 20: 473 - 495." title="The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago" type="journal article" year="2004">
Collingwood 2004; Sharaf
<emphasis id="B92D9A662C69974FFB1957F08BB5F964" box="[1201,1260,1683,1709]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
in press
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>