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<document ID-DOI="10.1080/00222933.2022.2115950" ID-ISSN="1464-5262" ID-Zenodo-Dep="7156474" approvalRequired="9" approvalRequired_for_textStreams="9" checkinTime="1665136984522" checkinUser="plazi" docAuthor="Medeiros, Rone A. F., Seidel, Matthias &amp; Grossi, Paschoal C." docDate="2022" docId="03F09661FFEC807DFEB5E4B47B5152D5" docLanguage="en" docName="JNATHIST.56.29-32.1315-1364.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 56 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2022.2115950" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Byrsopolis cribricollis Ohaus (MZUSP 1912" docType="treatment" docVersion="2" lastPageNumber="1336" masterDocId="FFC9EE19FFFF806BFFD0E116782B521F" masterDocTitle="Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay" masterLastPageNumber="1364" masterPageNumber="1315" pageNumber="1334" updateTime="1665137547684" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Medeiros, Rone A. F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>J. Nat. Hist.</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>29 - 32</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1315</mods:start>
<mods:end>1364</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.1080/00222933.2022.2115950</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2022.2115950</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">7156474</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7156502" ID-Zenodo-Dep="7156502" LSID="urn:lsid:plazi:treatment:03F09661FFEC807DFEB5E4B47B5152D5" httpUri="http://treatment.plazi.org/id/03F09661FFEC807DFEB5E4B47B5152D5" lastPageId="22" lastPageNumber="1336" pageId="19" pageNumber="1334">
<subSubSection box="[357,958,1442,1468]" pageId="19" pageNumber="1334" type="nomenclature">
<paragraph blockId="19.[357,958,1442,1468]" box="[357,958,1442,1468]" pageId="19" pageNumber="1334">
<heading bold="true" box="[357,958,1442,1468]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="1334" reason="3">
<emphasis bold="true" box="[357,958,1442,1468]" pageId="19" pageNumber="1334">
<taxonomicName authority="Ohaus, 1912" authorityName="Ohaus (MZUSP" authorityYear="1912" box="[357,781,1442,1468]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1334" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis bold="true" box="[357,616,1442,1468]" italics="true" pageId="19" pageNumber="1334">Byrsopolis cribricollis</emphasis>
<bibRefCitation author="Ohaus F" box="[624,781,1442,1468]" pageId="19" pageNumber="1334" pagination="273 - 319" refId="ref26994" refString="Ohaus F. 1912. Beitrage zur Kenntnis der Ruteliden. X. Deutsche entomologische Zeitschrift Berlin. 73: 273 - 319." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[790,893,1442,1468]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="19" pageNumber="1334">Figure 9</figureCitation>
(af))
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="1334" type="reference_group">
<paragraph blockId="19.[160,1156,1495,1659]" pageId="19" pageNumber="1334">
<treatmentCitationGroup pageId="19" pageNumber="1334">
<treatmentCitation author="Ohaus F" box="[160,589,1495,1520]" page="312" pageId="19" pageNumber="1334" year="1912">
<taxonomicName authority="Ohaus, 1912: 312" authorityName="Ohaus (MZUSP" authorityPageNumber="312" authorityYear="1912" box="[160,589,1495,1520]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1334" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis box="[160,385,1495,1520]" italics="true" pageId="19" pageNumber="1334">Byrsopolis cribricollis</emphasis>
<bibRefCitation author="Ohaus F" box="[394,589,1495,1520]" pageId="19" pageNumber="1334" pagination="273 - 319" refId="ref26994" refString="Ohaus F. 1912. Beitrage zur Kenntnis der Ruteliden. X. Deutsche entomologische Zeitschrift Berlin. 73: 273 - 319." type="journal article" year="1912">Ohaus, 1912: 312</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation author="Machatschke JW" box="[847,1084,1495,1520]" page="3" pageId="19" pageNumber="1334" year="1972">
<bibRefCitation author="Machatschke JW" box="[847,1084,1495,1520]" pageId="19" pageNumber="1334" pagination="361" refId="ref26699" refString="Machatschke JW. 1972. Scarabaeoidea: Melolonthinae, Rutelinae. Coleopterorum Catalogus. Supplementa. 66: 361." type="journal article" year="1972">Machatschke 1972: 3</bibRefCitation>
</treatmentCitation>
(catalogue, distribution);
<treatmentCitation author="Krajcik M" box="[388,564,1530,1555]" page="58" pageId="19" pageNumber="1334" year="2008">
<bibRefCitation author="Krajcik M" box="[388,564,1530,1555]" pageId="19" pageNumber="1334" pagination="142" refId="ref26488" refString="Krajcik M. 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). 2 nd ed. Plzen, Czech Republic: Milan Krajcik; p. 142." type="book chapter" year="2008">Krajcik 2008: 58</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation author="Carvalho TG &amp; Grossi PC" box="[697,1038,1530,1555]" page="372" pageId="19" pageNumber="1334" year="2018">
<bibRefCitation author="Carvalho TG &amp; Grossi PC" box="[697,1038,1530,1555]" pageId="19" pageNumber="1334" pagination="369 - 372" refId="ref25906" refString="Carvalho TG, Grossi PC. 2018. Description of a second species of the genus Moronius Grossi &amp; Vaz-deMello, 2015 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Zootaxa. 4434 (2): 369 - 372. doi: 10. 11646 / zootaxa. 4434.2.7." type="journal article" year="2018">Carvalho and Grossi 2018: 372</bibRefCitation>
</treatmentCitation>
(checklist, distribution);
<bibRefCitation author="Vaz-de-Mello FZ &amp; Grossi PC" box="[311,645,1565,1590]" pageId="19" pageNumber="1334" refId="ref27519" refString="Vaz-de-Mello FZ, Grossi PC. 2022. Melolonthidae in Catalogo Taxonomico da Fauna do Brasil. PNUD. http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127169." type="url" year="2022">Vaz-de-Mello and Grossi 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="19.[160,1156,1495,1659]" pageId="19" pageNumber="1334">
<treatmentCitationGroup pageId="19" pageNumber="1334">
<taxonomicName authority="(Ohaus, 1912)" baseAuthorityName="Ohaus" baseAuthorityYear="1912" box="[192,685,1599,1624]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1334" phylum="Arthropoda" rank="species" species="cribricollis" subGenus="Byrsopolis">
<emphasis box="[192,277,1599,1624]" italics="true" pageId="19" pageNumber="1334">Cotalpa</emphasis>
(
<emphasis box="[292,400,1599,1624]" italics="true" pageId="19" pageNumber="1334">Byrsopolis</emphasis>
)
<emphasis box="[415,523,1599,1624]" italics="true" pageId="19" pageNumber="1334">cribricollis</emphasis>
(
<bibRefCitation author="Ohaus F" box="[536,676,1599,1624]" pageId="19" pageNumber="1334" pagination="273 - 319" refId="ref26994" refString="Ohaus F. 1912. Beitrage zur Kenntnis der Ruteliden. X. Deutsche entomologische Zeitschrift Berlin. 73: 273 - 319." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
)
</taxonomicName>
: 312;
<treatmentCitation author="Ohaus F" box="[750,923,1599,1624]" page="10" pageId="19" pageNumber="1334" year="1918">
<bibRefCitation author="Ohaus F" box="[750,923,1599,1624]" pageId="19" pageNumber="1334" pagination="241" refId="ref27050" refString="Ohaus F. 1918. Coleopterorum Catalogus, pars. 66. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelianae. Vol. XX. Berlin, Germany: W. Junk; p. 241." type="book chapter" year="1918">Ohaus 1918: 10</bibRefCitation>
</treatmentCitation>
(catalogue, distribution); 1934: 38 (citation, checklist);
<treatmentCitation author="Blackwelder RE" box="[550,800,1634,1659]" page="235" pageId="19" pageNumber="1334" year="1944">
<bibRefCitation author="Blackwelder RE" box="[550,800,1634,1659]" pageId="19" pageNumber="1334" pagination="189 - 306" refId="ref25704" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Washington: Smithsonian Institution, United States National Museum. Bulletin 185, Part 2; p. 189 - 306." type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="1335" pageId="19" pageNumber="1334" type="materials_examined">
<paragraph blockId="19.[160,612,1702,1728]" box="[160,612,1702,1728]" pageId="19" pageNumber="1334">
<emphasis bold="true" box="[160,317,1702,1728]" italics="true" pageId="19" pageNumber="1334">Type locality.</emphasis>
<collectingCountry box="[332,395,1703,1728]" name="Brazil" pageId="19" pageNumber="1334">Brazil</collectingCountry>
,
<collectingRegion box="[405,518,1703,1728]" country="Brazil" name="Sao Paulo" pageId="19" pageNumber="1334">São Paulo</collectingRegion>
, Jacareí.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7156500" ID-Zenodo-Dep="7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="20" pageNumber="1335" startId="20.[160,225,952,975]" targetBox="[164,1151,145,917]" targetPageId="20">
<paragraph blockId="20.[160,1156,952,1033]" pageId="20" pageNumber="1335">
<emphasis bold="true" box="[160,250,952,975]" pageId="20" pageNumber="1335">Figure 9.</emphasis>
<taxonomicName authority="Ohaus." authorityName="Ohaus (MZUSP" authorityYear="1912" box="[257,529,952,975]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis box="[257,454,952,975]" italics="true" pageId="20" pageNumber="1335">Byrsopolis cribricollis</emphasis>
Ohaus.
</taxonomicName>
Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm.
</paragraph>
</caption>
<paragraph blockId="20.[160,1155,1135,1230]" pageId="20" pageNumber="1335">
<emphasis bold="true" box="[160,332,1135,1161]" italics="true" pageId="20" pageNumber="1335">Type material.</emphasis>
<emphasis box="[348,723,1136,1161]" italics="true" pageId="20" pageNumber="1335">
<typeStatus box="[348,452,1136,1161]" pageId="20" pageNumber="1335">Lectotype</typeStatus>
female, here designated
</emphasis>
. (a)
<taxonomicName box="[776,893,1136,1161]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/
<taxonomicName authority="Ohs." authorityName="Ohs." box="[901,1074,1136,1161]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="species" species="cribricollis">cribricollis Ohs.</taxonomicName>
[handscript by Ohaus], (b) Typus! [red label], c) S. PAULO/Jacarehy [printed label] (
<specimenCount box="[1030,1061,1170,1195]" pageId="20" pageNumber="1335" type="female">1♀</specimenCount>
<collectionCode box="[1067,1150,1170,1195]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="20" pageNumber="1335">MZUSP</collectionCode>
) (
<figureCitation box="[168,260,1205,1230]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="20" pageNumber="1335">Figure 9</figureCitation>
).
</paragraph>
<paragraph blockId="20.[160,1155,1284,1454]" pageId="20" pageNumber="1335">
<emphasis bold="true" box="[160,417,1289,1315]" italics="true" pageId="20" pageNumber="1335">
<typeStatus box="[160,323,1289,1315]" pageId="20" pageNumber="1335">Paralectotype</typeStatus>
female.
</emphasis>
(a)
<materialsCitation collectorName="Ohaus-leg" country="Brazil" location="Ypiranga" pageId="20" pageNumber="1335" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paralectotype">
<collectingRegion box="[468,582,1290,1315]" country="Brazil" name="Sao Paulo" pageId="20" pageNumber="1335">São Paulo</collectingRegion>
/
<location LSID="urn:lsid:plazi:treatment:03F09661FFEC807DFEB5E4B47B5152D5:8E8671ACFFEB807FFD9DE41C7A9D573C" box="[589,694,1290,1315]" country="Brazil" name="Ypiranga" pageId="20" pageNumber="1335" stateProvince="Sao Paulo">Ypiranga</location>
/xii, (b) COTYPUS, (c) DIC/
<collectingCountry box="[985,1070,1290,1315]" name="Brazil" pageId="20" pageNumber="1335">BRASIL</collectingCountry>
/E o
<collectingRegion country="Brazil" name="Sao Paulo" pageId="20" pageNumber="1335">SAO PAULO</collectingRegion>
/YPIRANGA/
<collectorName box="[394,509,1325,1350]" pageId="20" pageNumber="1335">Ohaus-leg</collectorName>
</materialsCitation>
./
<materialsCitation box="[519,560,1325,1350]" country="Brazil" location="Coll" pageId="20" pageNumber="1335" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paralectotype">
<location LSID="urn:lsid:plazi:treatment:03F09661FFEC807DFEB5E4B47B5152D5:8E8671ACFFEB807FFDD7E43B7A1B5759" box="[519,560,1325,1350]" country="Brazil" name="Coll" pageId="20" pageNumber="1335" stateProvince="Sao Paulo">Coll</location>
</materialsCitation>
<materialsCitation box="[560,954,1325,1350]" collectorName="Ohaus" country="Brazil" location="Martinez'" pageId="20" pageNumber="1335" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paralectotype">
.
<location LSID="urn:lsid:plazi:treatment:03F09661FFEC807DFEB5E4B47B5152D5:8E8671ACFFEB807FFD92E43B7A845759" box="[578,687,1325,1350]" country="Brazil" name="Martinez'" pageId="20" pageNumber="1335" stateProvince="Sao Paulo">Martinez</location>
, (d)
<collectorName box="[754,830,1325,1350]" pageId="20" pageNumber="1335">Ohaus</collectorName>
determin
</materialsCitation>
. 1912/
<taxonomicName box="[1036,1155,1325,1350]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
cotype
<taxonomicName authority="Ohs." authorityName="Ohs." box="[247,421,1360,1385]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="species" species="cribricollis">cribricollis Ohs.</taxonomicName>
<specimenCount box="[430,450,1360,1383]" pageId="20" pageNumber="1335" type="female"></specimenCount>
, (e) H. &amp; A. HOWDEN COLLECTION, ex. A.
<materialsCitation collectorName="Ohaus" country="Brazil" location="Martinez Coll'" pageId="20" pageNumber="1335" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paralectotype">
<location LSID="urn:lsid:plazi:treatment:03F09661FFEC807DFEB5E4B47B5152D5:8E8671ACFFEB807FFC10E4467C745776" box="[960,1119,1360,1385]" country="Brazil" name="Martinez Coll'" pageId="20" pageNumber="1335" stateProvince="Sao Paulo">Martinez Coll</location>
, (f)
<taxonomicName authority="(Byrs.)" baseAuthorityName="Byrs." box="[162,326,1394,1419]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1335" phylum="Arthropoda" rank="genus">Cotalpa (Byrs.)</taxonomicName>
/cribricollis
<specimenCount box="[460,480,1395,1418]" pageId="20" pageNumber="1335" type="female"></specimenCount>
/
<collectorName box="[488,564,1394,1419]" pageId="20" pageNumber="1335">Ohaus</collectorName>
/A
</materialsCitation>
. Martinez-DET.
<materialsCitation collectionCode="CMN" collectorName="Francois Genier" country="Brazil" pageId="20" pageNumber="1335" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paralectotype">
1951, (g) CMNEN/2001-0613 (
<specimenCount box="[1124,1155,1394,1419]" pageId="20" pageNumber="1335" type="female">1♀</specimenCount>
<collectionCode box="[160,217,1429,1454]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="20" pageNumber="1335" type="Museum">CMN</collectionCode>
, specimen examined through images by
<collectorName box="[682,860,1429,1454]" pageId="20" pageNumber="1335">François Génier</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="1335" type="diagnosis">
<paragraph blockId="20.[160,1156,1513,1712]" pageId="20" pageNumber="1335">
<emphasis bold="true" box="[160,284,1513,1539]" italics="true" pageId="20" pageNumber="1335">Diagnosis.</emphasis>
Body with reddish-brown colouration, without metallic green reflections; clypeal margin and pronotum dark brown; clypeus subrectangular with rounded anterior margin and anterior angles; antennal club about 1.5x longer than antennomeres IIVII combined; scutellar plate with broadly rounded base and sides straight; apex of elytral marginal areas with acute angles; humeri densely punctate; elytra with strong striae; protibial spur about 2.8x longer than wide.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="1336" type="description">
<paragraph blockId="21.[160,1156,142,1277]" pageId="21" pageNumber="1336">
<emphasis bold="true" box="[160,326,142,168]" italics="true" pageId="21" pageNumber="1336">Redescription.</emphasis>
<emphasis box="[342,489,143,168]" italics="true" pageId="21" pageNumber="1336">Female. Body</emphasis>
. Shape elongate oval; reddish-brown; frontoclypeal suture dark brown; antenna orange to dark brown; pronotal margins reddish brown at sides and dark brown at apex and base; scutellar plate marginal areas reddish-brown to dark brown, brighter and smaller in width at apex, increasing gradually to darker and thicker at base; orange setae scattered near apex; elytra reddish-brown, with marginal areas dark brown, without metallic green reflections (
<figureCitation box="[549,641,316,341]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(a,b)). Length 30.2831.00 mm. Humeral width
<quantity box="[160,280,351,376]" metricMagnitude="-2" metricUnit="m" metricValue="1.5939999999999999" pageId="21" pageNumber="1336" unit="mm" value="15.94">15.94 mm</quantity>
. Elytral width
<quantity box="[455,650,351,376]" metricMagnitude="-2" metricUnit="m" metricValue="1.8094999999999999" metricValueMax="1.85" metricValueMin="1.769" pageId="21" pageNumber="1336" unit="mm" value="18.095" valueMax="18.5" valueMin="17.69">17.6918.50 mm</quantity>
.
<emphasis box="[665,722,351,376]" italics="true" pageId="21" pageNumber="1336">Head</emphasis>
. Clypeus about 2x wider than long; subrectangular, with apex and angles weakly rounded; concavity weak at middle, moderate at apex; frontoclypeal suture almost straight, with curvilinear extremities very straight laterally; strongly striate; densely puctate; punctures moderate large to large; glabrous; frons about 1.50x longer than clypeus; punctures small to large, moderately to densely punctate; glabrous; mandibles with rounded external border, forming an almost right angle between scissorial region and apex; scissorial region slightly concave; small punctures at middle, moderately to densely punctate; scissorial region and apex of external border with setae sparsely to moderately distributed (apex and disc) (
<figureCitation box="[1061,1155,628,653]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(c)); maxillary galea with three teeth; apical tooth longer than basal and median teeth (of similar size); small basal tooth with a small seta; apex of prementum moderately emarginate; last labial palpomere about 0.75x shorter than palpomere II (
<figureCitation box="[930,1024,732,757]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(e)); labrum weakly emarginated (
<figureCitation box="[402,495,767,792]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(e)); antennomere II subconical, antennomere III subcylindrical, antennomeres IV and V subquadrate, antennomere VI subconical and VII subelliptical; antennal club about 1.50x longer than antennomeres IIVII combined.
<emphasis box="[1074,1150,836,861]" italics="true" pageId="21" pageNumber="1336">Thorax</emphasis>
. Pronotum transverse, about 1.60x wider than long; punctures small to moderate, moderately to densely punctate (
<figureCitation box="[469,563,906,931]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(a)); glabrous at disc; marginal areas with small and moderate, sparsely distributed setae; strongly striate; longitudinal midline with concavity at the apex; scutellar plate subtriangular, straight sides, broadly rounded at the base; setae small and moderate, moderately to densely distributed; surface moderately striate; punctures small to moderately large, sparsely to moderately punctate on the sides and densely at disc.
<emphasis box="[340,401,1079,1104]" italics="true" pageId="21" pageNumber="1336">Elytra</emphasis>
. About 1.20x longer than wide; strongly striate; punctures small to large, sparsely to densely distributed; humeri with small to moderate punctures, densely punctate; interstriae with large coalescent punctures, (
<figureCitation box="[793,890,1148,1173]" captionStart="Figure 9" captionStartId="20.[160,225,952,975]" captionTargetBox="[164,1151,145,917]" captionTargetId="figure-149@20.[162,1154,142,920]" captionTargetPageId="20" captionText="Figure 9. Byrsopolis cribricollis Ohaus. Lectotype, São Paulo, Jacareí. Dorsal habitus (a), lateral habitus (b); dorsal view of right mandible (c); ventral view of right maxilla (d); dorsal view of mentum (e); labels attached (f). Scale bars: a, b = 5 mm; ce = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156500" httpUri="https://zenodo.org/record/7156500/files/figure.png" pageId="21" pageNumber="1336">Figure 9</figureCitation>
(a,b)); elytral apex with acute angles; glabrous.
<emphasis box="[427,475,1183,1208]" italics="true" pageId="21" pageNumber="1336">Legs</emphasis>
. Protibial spur about 2.8x longer than wide; mesotibiae and metatibiae with two carinae; apex of mesotibia with about 14 spine-like setae and metatibiae with about 29 spine-like setae.
</paragraph>
<paragraph blockId="21.[160,356,1330,1356]" box="[160,356,1330,1356]" pageId="21" pageNumber="1336">
<emphasis bold="true" box="[160,225,1330,1356]" italics="true" pageId="21" pageNumber="1336">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="1336" type="etymology">
<paragraph blockId="21.[160,1155,1408,1468]" pageId="21" pageNumber="1336">
<emphasis bold="true" box="[160,291,1408,1434]" italics="true" pageId="21" pageNumber="1336">Etymology.</emphasis>
The species name is derived from Latin,
<taxonomicName box="[759,867,1408,1433]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1336" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis box="[759,867,1408,1433]" italics="true" pageId="21" pageNumber="1336">cribricollis</emphasis>
</taxonomicName>
=
<emphasis box="[897,981,1408,1433]" italics="true" pageId="21" pageNumber="1336">crībrum</emphasis>
(= riddled) and
<emphasis box="[160,213,1443,1468]" italics="true" pageId="21" pageNumber="1336">collis</emphasis>
(= neck), referring to the strongly punctate pronotum.
</paragraph>
</subSubSection>
<subSubSection box="[160,815,1520,1546]" pageId="21" pageNumber="1336" type="distribution">
<paragraph blockId="21.[160,815,1520,1546]" box="[160,815,1520,1546]" pageId="21" pageNumber="1336">
<emphasis bold="true" box="[160,307,1520,1546]" italics="true" pageId="21" pageNumber="1336">Distribution.</emphasis>
<collectingRegion box="[323,434,1521,1546]" country="Brazil" name="Sao Paulo" pageId="21" pageNumber="1336">São Paulo</collectingRegion>
(Jacareí and Ipiranga) (
<figureCitation box="[697,802,1521,1546]" captionStart="Figure 19" captionStartId="41.[160,225,961,984]" captionTargetBox="[162,1154,142,929]" captionTargetId="figure-144@41.[162,1154,142,929]" captionTargetPageId="41" captionText="Figure 19. Distribution map of Byrsopolis species." figureDoi="http://doi.org/10.5281/zenodo.7156532" httpUri="https://zenodo.org/record/7156532/files/figure.png" pageId="21" pageNumber="1336">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="1337" pageId="21" pageNumber="1336" type="discussion">
<paragraph blockId="21.[160,1155,1598,1728]" lastBlockId="22.[160,1156,142,202]" lastPageId="22" lastPageNumber="1337" pageId="21" pageNumber="1336">
<emphasis bold="true" box="[160,268,1598,1624]" italics="true" pageId="21" pageNumber="1336">Remarks.</emphasis>
According to
<bibRefCitation author="Ohaus F" box="[435,583,1599,1624]" pageId="21" pageNumber="1336" pagination="273 - 319" refId="ref26994" refString="Ohaus F. 1912. Beitrage zur Kenntnis der Ruteliden. X. Deutsche entomologische Zeitschrift Berlin. 73: 273 - 319." type="journal article" year="1912">Ohaus (1912)</bibRefCitation>
, the type specimens were deposited at Museum of São Paulo (MZSP). But with the deposition of the Ohaus collection in ZMHB the
<typeStatus box="[1050,1155,1634,1659]" pageId="21" pageNumber="1336">lectotype</typeStatus>
from Jacareí found its way to
<collectingRegion box="[511,577,1669,1694]" country="Germany" name="Berlin" pageId="21" pageNumber="1336">Berlin</collectingRegion>
, where it has been ever since. In agreement with Johannes Frisch (ZMHB), the specimen was finally transferred to the MZSP. The second specimen (
<typeStatus box="[281,439,142,167]" pageId="22" pageNumber="1337">paralectotype</typeStatus>
) from Ipiranga was found in the collection of Antonio Martínez, and is now deposited in the Canadian Museum of Nature (CMN).
</paragraph>
</subSubSection>
</treatment>
</document>