190 lines
14 KiB
XML
190 lines
14 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3789798" ID-GBIF-Dataset="a3fbe31d-fb13-452a-b213-ce4827a3b08e" ID-GBIF-Taxon="163815651" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789798" ID-ZooBank="7AB88B91-B696-4CA1-923A-AEB569AF2FC7" approvalRequired="3" approvalRequired_for_treatments="3" checkinTime="1587991013482" checkinUser="plazi" docAuthor="Ziani, Stefano & Gharakhloo, Mohammad Moradi" docDate="2010" docId="03DFAC174D61C44BFF4FFB5CFD2DFD35" docLanguage="en" docName="ZK_article_2136.pdf" docOrigin="ZooKeys 34" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@55b80e5e" docStyleName="zookeys.2008.journal_article" docTitle="Onthophagus furciceps Marseul 1869" docType="key" docVersion="4" lastPageId="6" lastPageNumber="39" masterDocId="FFE6D46F4D64C44DFFC3FFDCFFE8FF98" masterDocTitle="Studies on palearctic Onthophagus associated with burrows of small mammals. IV. A new Iranian species belonging to the furciceps group (Coleoptera, Scarabaeidae, Onthophagini)" masterLastPageNumber="40" masterPageNumber="33" pageId="5" pageNumber="38" updateTime="1643559032930" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Studies on palearctic Onthophagus associated with burrows of small mammals. IV. A new Iranian species belonging to the furciceps group (Coleoptera, Scarabaeidae, Onthophagini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ziani, Stefano</mods:namePart>
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<mods:nameIdentifier type="LSID">92C5BE3E-FA6A-4336-AA54-D8647B3374C0</mods:nameIdentifier>
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<mods:affiliation>Via S. Giovanni, 41 / a, 47014 Meldola (FC), Italy</mods:affiliation>
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<mods:nameIdentifier type="email">stefanoziani@alice.it</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gharakhloo, Mohammad Moradi</mods:namePart>
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<mods:nameIdentifier type="LSID">F8F3D316-7AFA-40F4-9D61-68BFD427BB05</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Faculty of Science, Zanjan University, Zanjan, Iran</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-01-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.34.272</mods:identifier>
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<mods:identifier type="GBIF-Dataset">a3fbe31d-fb13-452a-b213-ce4827a3b08e</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576602</mods:identifier>
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<mods:identifier type="ZooBank">7AB88B91-B696-4CA1-923A-AEB569AF2FC7</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789798" ID-GBIF-Taxon="163815651" ID-Zenodo-Dep="3789798" LSID="urn:lsid:plazi:treatment:03DFAC174D61C44BFF4FFB5CFD2DFD35" httpUri="http://treatment.plazi.org/id/03DFAC174D61C44BFF4FFB5CFD2DFD35" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
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<subSubSection box="[140,579,1152,1178]" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph blockId="5.[140,579,1152,1178]" box="[140,579,1152,1178]" pageId="5" pageNumber="38">
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<heading bold="true" box="[140,579,1152,1178]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<emphasis bold="true" box="[140,579,1152,1178]" pageId="5" pageNumber="38">
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Key to the
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<taxonomicName authorityName="Marseul" authorityYear="1869" box="[270,410,1152,1178]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="furciceps">
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<emphasis bold="true" box="[270,410,1152,1178]" italics="true" pageId="5" pageNumber="38">
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<collectionCode box="[270,294,1152,1178]" country="Norway" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="5" pageNumber="38" type="Herbarium">O</collectionCode>
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. furciceps
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</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38" type="key">
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<keyStep pageId="5" pageNumber="38">
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<paragraph blockId="5.[140,1108,1223,1672]" pageId="5" pageNumber="38">
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<keyLead pageId="5" pageNumber="38">
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1. Bristles on elytral disc simple; dorsal surface shining, disc of pronotum barely or not at all microreticulate.........................................................................
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<emphasis bold="true" box="[1092,1107,1257,1283]" pageId="5" pageNumber="38">4</emphasis>
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</keyLead>
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</paragraph>
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<paragraph blockId="5.[140,1108,1223,1672]" pageId="5" pageNumber="38">
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<keyLead pageId="5" pageNumber="38">
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– Bristles on elytral disc bifid or trifid, or lancelolate; dorsal surface dull, with distinct microreticulation............................................................................
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<emphasis bold="true" box="[1092,1107,1328,1354]" pageId="5" pageNumber="38">2</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="5" pageNumber="38">
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<paragraph blockId="5.[140,1108,1223,1672]" pageId="5" pageNumber="38">
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<keyLead pageId="5" pageNumber="38">
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2. Lamellae of antennal club thin and elongate, much longer than in most
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<taxonomicName authorityName="Latreille" authorityYear="1802" box="[246,389,1399,1425]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis box="[246,389,1399,1425]" italics="true" pageId="5" pageNumber="38">Onthophagus</emphasis>
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</taxonomicName>
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species, somewhat resembling those of some Melolonthini....... ..........................................................................
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<taxonomicName authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[794,1107,1433,1460]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="parmatus">
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<emphasis bold="true" box="[794,1107,1433,1460]" pageId="5" pageNumber="38">
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<emphasis bold="true" box="[794,945,1434,1460]" italics="true" pageId="5" pageNumber="38">
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<collectionCode box="[794,818,1434,1460]" country="Norway" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="5" pageNumber="38" type="Herbarium">O</collectionCode>
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. parmatus
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</emphasis>
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Reitter, 1892
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="5.[140,1108,1223,1672]" box="[140,1107,1469,1496]" pageId="5" pageNumber="38">
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<keyLead box="[140,1107,1469,1496]" pageId="5" pageNumber="38">
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– Lamellae of antennal club normally shaped.................................................
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<emphasis bold="true" box="[1092,1107,1469,1495]" pageId="5" pageNumber="38">3</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
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<paragraph blockId="5.[140,1108,1223,1672]" pageId="5" pageNumber="38">
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<keyLead pageId="5" pageNumber="38">
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3. Relatively small species (length
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<quantity box="[625,777,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.2" metricValueMin="4.8" pageId="5" pageNumber="38" unit="mm" value="6.5" valueMax="8.2" valueMin="4.8">4.8–8.2 mm</quantity>
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). Pronotum equally densely punctate on disc and basally. Male: pronotum with four tubercles, the two anteromedian very closely placed and smaller than the two anterolateral; female: pronotum with an anteromedian narrow regularly convex gibbosity.... ...........................................................................
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<taxonomicName authority="Harold, 1883" authorityName="Harold" authorityYear="1883" box="[803,1107,1645,1671]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="ponticus">
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<emphasis bold="true" box="[803,1107,1645,1671]" pageId="5" pageNumber="38">
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<emphasis bold="true" box="[803,939,1645,1671]" italics="true" pageId="5" pageNumber="38">
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<collectionCode box="[803,827,1645,1671]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="5" pageNumber="38" type="Herbarium">O</collectionCode>
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. ponticus
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</emphasis>
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Harold,
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<date box="[1046,1107,1645,1671]" pageId="5" pageNumber="38" value="1883">1883</date>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="6.[140,1108,166,475]" pageId="6" pageNumber="39">
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<keyLead pageId="6" pageNumber="39">
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– Relatively large species (length
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<quantity box="[594,740,166,193]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.9" metricValueMin="5.8" pageId="6" pageNumber="39" unit="mm" value="7.85" valueMax="9.9" valueMin="5.8">5.8–9.9 mm</quantity>
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). Pronotum densely punctate on disc, punctures gradually more spaced toward base. Male: pronotum with four equidistant tubercles, the two anteromedian more prominent than the two anterolateral; female: pronotum with an anteromedian transverse sunk gibbosity.................................................................
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<emphasis bold="true" box="[831,1108,307,334]" pageId="6" pageNumber="39">
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<taxonomicName authority="Ziani & Moradi, 2010" authorityName="Ziani & Moradi" authorityYear="2010" box="[831,1036,307,334]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="psychopompus" status="sp. n.">
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<emphasis bold="true" box="[831,1036,307,334]" italics="true" pageId="6" pageNumber="39">
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<collectionCode box="[831,855,307,333]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="6" pageNumber="39" type="Herbarium">O</collectionCode>
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. psychopompus
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1043,1108,307,333]" pageId="6" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="6" pageNumber="39">
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<paragraph blockId="6.[140,1108,166,475]" pageId="6" pageNumber="39">
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<keyLead pageId="6" pageNumber="39">
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4. Pronotal sides obviously sinuate behind anterior angles; pronotum approximately as long as elytra......................................
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<taxonomicName authority="Marseul, 1869" authorityName="Marseul" authorityYear="1869" box="[788,1107,377,404]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="furciceps">
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<emphasis bold="true" box="[788,1107,377,404]" pageId="6" pageNumber="39">
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<emphasis bold="true" box="[788,928,377,404]" italics="true" pageId="6" pageNumber="39">O. furciceps</emphasis>
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Marseul,
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<date box="[1046,1107,377,403]" pageId="6" pageNumber="39" value="1869">1869</date>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="6.[140,1108,166,475]" pageId="6" pageNumber="39">
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<keyLead pageId="6" pageNumber="39">
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– Pronotal sides not sinuate behind anterior angles; pronotum longer than elytra...............................................................
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<taxonomicName authority="Balthasar, 1960" authorityName="Balthasar" authorityYear="1960" box="[774,1107,448,475]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="bytinskii">
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<emphasis bold="true" box="[774,1107,448,475]" pageId="6" pageNumber="39">
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<emphasis bold="true" box="[774,915,448,475]" italics="true" pageId="6" pageNumber="39">O. bytinskii</emphasis>
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Balthasar,
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<date box="[1046,1107,448,474]" pageId="6" pageNumber="39" value="1960">1960</date>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="39" type="discussion">
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<paragraph blockId="6.[140,1108,519,686]" pageId="6" pageNumber="39">
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In
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<bibRefCitation author="Balthasar V" box="[172,364,519,546]" pageId="6" pageNumber="39" refId="ref3144" refString="Balthasar V (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 627 pp., 16 pls." type="book" year="1963">
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Balthasar (
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<date box="[292,354,519,546]" pageId="6" pageNumber="39" value="1963">1963</date>
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)
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</bibRefCitation>
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, the last taxonomic monograph covering Iranian territory,
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<taxonomicName authority="Ziani & Moradi, 2010" authorityName="Ziani & Moradi" authorityYear="2010" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="psychopompus" status="sp. n.">
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<emphasis italics="true" pageId="6" pageNumber="39">O. psychopompus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[265,325,554,580]" pageId="6" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
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can be inserted in the key at thesis 24 (21), p. 199, to form a couplet with
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<taxonomicName authorityName="Harold" authorityYear="1883" box="[197,321,590,616]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="ponticus">
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<emphasis box="[197,321,590,616]" italics="true" pageId="6" pageNumber="39">O. ponticus</emphasis>
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</taxonomicName>
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from which is easily distinguished by the characters mentioned above. Moreover, the shape of the lamella copulatrix, with the right branch, in ventral view, bifurcate, is very characteristic for the new species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |