268 lines
62 KiB
XML
268 lines
62 KiB
XML
<document id="6C91E401AD61F699C5B7716443ED7322" ID-CLB-Dataset="5795" ID-DOI="10.2478/aemnp-2019-0008" ID-GBIF-Dataset="e3fd15a2-5823-4640-bd84-46058d760fb1" ID-ISSN="1804-6487" ID-Zenodo-Dep="4505000" ID-ZooBank="92E283C0-30BB-4610-8D48-12B40CF9B425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612474806317" checkinUser="felipe" docAuthor="Yin, Zi-Wei, Kurbatov, Sergey A., Cuccodoro, Giulio & Cai, Chen-Yang" docDate="2019" docId="03C53526FFCFFFC7FEC1F9B8BA39F7F6" docLanguage="en" docName="ActaEntMusNatPra.59.1.101-106.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1)" docSource="http://dx.doi.org/10.2478/aemnp-2019-0008" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Cretobrachygluta Yin & Kurbatov & Cuccodoro & Cai 2019, gen. nov." docType="treatment" docVersion="8" lastPageNumber="103" masterDocId="FFFC4D5EFFCEFFC5FFEAFFE9BB5CFFE8" masterDocTitle="Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="106" masterPageNumber="101" pageNumber="102" updateTime="1698927449831" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="5FA81D383B20D3DE46A0E7BE8ADD05B3">Cretobrachygluta gen. nov., the first and oldest Brachyglutini in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="9CDF0B774BEB3B945F08D6D9BF1EE35B" type="personal">
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<mods:roleTerm id="67BBE38B8B7D80196AA4E794B1F2AEDF">Author</mods:roleTerm>
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<mods:namePart id="1E10A238B64F76D99C3B9EC58C3ACB21">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="88BA220A23D1F034385868977B05104F">Laboratory of Environmental Entomology, College of Life Sciences, Shanghai Normal University, Shanghai, 200234, P. R. China</mods:affiliation>
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<mods:name id="81037490F7C875F9F64FCE0D4F509E2F" type="personal">
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<mods:roleTerm id="88635D00A96213A9FACC4C55E5EA867E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B5F7490C794E16E250331B6074405AD3">Kurbatov, Sergey A.</mods:namePart>
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<mods:affiliation id="53D383BB555D35B1FEB44678C596D5A2">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="1A137FE6D7589631F569E6383877F423" type="personal">
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<mods:roleTerm id="94B337B711ADE178C52562B52F8707EE">Author</mods:roleTerm>
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<mods:namePart id="D7109DBE8BC7FFCC94CA411C28278D52">Cuccodoro, Giulio</mods:namePart>
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<mods:affiliation id="3166B569C109E3B6A586934D54596F79">Muséum d’histoire naturelle, Case postale 6434, CH- 1211 Genève</mods:affiliation>
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<mods:namePart id="5E7F8E4CDF92675FF2741D5A01D2E177">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="D39A101B7FAA632B3AC6FC1A3861C376">CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, P. R. China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS</mods:affiliation>
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<mods:title id="5B6E78251D3E36AEE62465D113BF637F">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="621A86A89817C8DC48CAD6A610C7ED34">2019</mods:date>
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<mods:title id="B4D46F7EE345A53D4636510F951672CC">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="AD938CBC0544551A0E0A235649F0C53E">2019-03-04</mods:number>
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<treatment id="03C53526FFCFFFC7FEC1F9B8BA39F7F6" ID-DOI="http://doi.org/10.5281/zenodo.5062341" ID-GBIF-Taxon="183237542" ID-Zenodo-Dep="5062341" LSID="urn:lsid:plazi:treatment:03C53526FFCFFFC7FEC1F9B8BA39F7F6" httpUri="http://treatment.plazi.org/id/03C53526FFCFFFC7FEC1F9B8BA39F7F6" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102">
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<subSubSection id="C376D7BBFFCFFFC4FEC1F9B8B938F980" box="[299,612,1616,1640]" pageId="1" pageNumber="102" type="nomenclature">
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<paragraph id="8BD38430FFCFFFC4FEC1F9B8B938F980" blockId="1.[299,612,1616,1640]" box="[299,612,1616,1640]" pageId="1" pageNumber="102">
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<heading id="D09B335CFFCFFFC4FEC1F9B8B938F980" box="[299,612,1616,1640]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="102" reason="3">
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<emphasis id="B9185822FFCFFFC4FEC1F9B8B938F980" bold="true" box="[299,612,1616,1640]" pageId="1" pageNumber="102">
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†
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<taxonomicName id="4C6CFFB3FFCFFFC4FED2F9B9BAA5F980" ID-CoL="8XKLZ" authority="Yin & Kurbatov & Cuccodoro & Cai, 2019" authorityName="Yin & Kurbatov & Cuccodoro & Cai" authorityYear="2019" box="[312,505,1616,1640]" class="Insecta" family="Staphylinidae" genus="Cretobrachygluta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9185822FFCFFFC4FED2F9B9BAA5F980" bold="true" box="[312,505,1616,1640]" italics="true" pageId="1" pageNumber="102">Cretobrachygluta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22BE559FFCFFFC4FDEAF9B8B938F980" box="[512,612,1617,1640]" pageId="1" pageNumber="102" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C376D7BBFFCFFFC4FE71F999BAA8F96B" box="[411,500,1648,1667]" pageId="1" pageNumber="102" type="description">
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<paragraph id="8BD38430FFCFFFC4FE71F999BAA8F96B" blockId="1.[151,759,1648,1739]" box="[411,500,1648,1667]" pageId="1" pageNumber="102">
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(
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<figureCitation id="135798B5FFCFFFC4FE4BF999BAB1F96B" box="[417,493,1648,1667]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[150,181,1388,1407]" captionStartId-1="3.[151,182,1498,1517]" captionTargetBox-0="[198,1389,197,1338]" captionTargetBox-1="[151,1436,190,1468]" captionTargetId-0="figure-329@2.[198,1389,189,1368]" captionTargetId-1="figure-19@3.[151,1436,189,1468]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Habitus of †Cretobrachygluta laurasiensis gen. et sp. nov. A – dorsal view; B – ventral view. Scale bar: 0.5 mm." captionText-1="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi-0="http://doi.org/10.5281/zenodo.4505002" figureDoi-1="http://doi.org/10.5281/zenodo.4505008" httpUri-0="https://zenodo.org/record/4505002/files/figure.png" httpUri-1="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Figs 1–2</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C376D7BBFFCFFFC4FF7DF97DBA19F923" pageId="1" pageNumber="102" type="type_taxon">
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<paragraph id="8BD38430FFCFFFC4FF7DF97DBA19F923" blockId="1.[151,759,1648,1739]" pageId="1" pageNumber="102">
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<emphasis id="B9185822FFCFFFC4FF7DF97DBA73F943" bold="true" box="[151,303,1684,1707]" pageId="1" pageNumber="102">
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<typeStatus id="54D73A92FFCFFFC4FF7DF97DBB93F943" box="[151,207,1684,1707]" pageId="1" pageNumber="102">Type</typeStatus>
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species.
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</emphasis>
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†
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<taxonomicName id="4C6CFFB3FFCFFFC4FEADF97DB9CDF943" ID-CoL="8XYV5" authority="Yin & Kurbatov & Cuccodoro & Cai, 2019" authorityName="Yin & Kurbatov & Cuccodoro & Cai" authorityYear="2019" box="[327,657,1684,1707]" class="Insecta" family="Staphylinidae" genus="Cretobrachygluta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="laurasiensis" status="sp. nov.">
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<emphasis id="B9185822FFCFFFC4FEADF97DB9CDF943" box="[327,657,1684,1707]" italics="true" pageId="1" pageNumber="102">Cretobrachygluta laurasiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22BE559FFCFFFC4FD71F97DB9A8F943" box="[667,756,1684,1707]" pageId="1" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
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, here designated.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376D7BBFFCFFFC4FF7DF90ABF60FEFC" pageId="1" pageNumber="102" type="diagnosis">
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<paragraph id="8BD38430FFCFFFC4FF7DF90ABF60FEFC" blockId="1.[151,760,1763,2074]" lastBlockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
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<emphasis id="B9185822FFCFFFC4FF7DF90ABA57F912" bold="true" box="[151,267,1763,1786]" pageId="1" pageNumber="102">Diagnosis.</emphasis>
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Head with small, nude vertexal foveae, lacking sulcus connecting foveae; antennae with club formed by apical three antennomeres; gular region with broad median ridge. Pronotum with small, nude median and lateral antebasal foveae. Each elytron with four basal foveae; subhumeral foveae and marginal striae present; discal striae absent. Metacoxae contiguous, moderately projecting posteriorly; tarsomere 2 distinctly shorter than tarsomere 3; tarsi each with single pretarsal claw. Abdomen with tergite 1 (IV) much longer than 2 (V); tergites 1–4 (IV–VII) with broad paratergites; tergite 1 (IV) with two basolateral foveae, lacking mediobasal fovea; sternite 1 (III) as long as 2 (IV) along midline.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376D7BBFFCFFFC7FCD6FEF4BAF0F876" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102" type="description">
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<paragraph id="8BD38430FFCFFFC4FCD6FEF4BE5CFE9C" blockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
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<emphasis id="B9185822FFCFFFC4FCD6FEF4B89BFEDC" bold="true" box="[828,967,285,308]" pageId="1" pageNumber="102">Description.</emphasis>
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Body length slightly over
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<quantity id="4C9429D5FFCFFFC4FB06FEF4BE1FFEDC" box="[1260,1347,285,308]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="102" unit="mm" value="1.8">1.8 mm</quantity>
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; habitus (
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<figureCitation id="135798B5FFCFFFC4FCAEFED4B8D8FEBC" box="[836,900,317,340]" captionStart="Fig" captionStartId="2.[150,181,1388,1407]" captionTargetBox="[198,1389,197,1338]" captionTargetId="figure-329@2.[198,1389,189,1368]" captionTargetPageId="2" captionText="Fig. 1. Habitus of †Cretobrachygluta laurasiensis gen. et sp. nov. A – dorsal view; B – ventral view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4505002" httpUri="https://zenodo.org/record/4505002/files/figure.png" pageId="1" pageNumber="102">Fig. 1</figureCitation>
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) strongly convex dorso-ventrally, in dorsal view strongly broadened posterior to pronotum.
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</paragraph>
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<paragraph id="8BD38430FFCFFFC4FCB0FE94BFC7FC1C" blockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
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Head (
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<figureCitation id="135798B5FFCFFFC4FC49FE94B8ABFE7C" box="[931,1015,381,404]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
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) roundly quadrate, slightly wider than pronotum; vertex raised, with pair of small and nude vertexal foveae (
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<figureCitation id="135798B5FFCFFFC4FC06FE54BF1DFE3C" box="[1004,1089,445,468]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2A</figureCitation>
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;
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<emphasis id="B9185822FFCFFFC4FBA1FE54BF02FE3C" box="[1099,1118,445,468]" italics="true" pageId="1" pageNumber="102">vf</emphasis>
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); frons shallowly and broadly impressed, impression flanked laterally by widely separated antennal tubercles, and demarcated anteriorly by broad frontal-clypeal ridge; ocular-mandibular carinae (
|
||
<figureCitation id="135798B5FFCFFFC4FCAEFDD4B8C7FDBC" box="[836,923,573,596]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2D</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FC40FDD4B88AFDBC" box="[938,982,573,596]" italics="true" pageId="1" pageNumber="102">omc</emphasis>
|
||
) distinct; gular region broadly impressed at middle, two close but clearly separated foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FA9BFDB4B802FD7C" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FC80FD94B823FD7C" box="[874,895,637,660]" italics="true" pageId="1" pageNumber="102">gf</emphasis>
|
||
) in impression; broad, longitudinally oval median gular ridge present anterior to foveae and extending to mouthparts, demarcated laterally by sulci. Antennae (
|
||
<figureCitation id="135798B5FFCFFFC4FA9BFD54B802FD1C" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2A</figureCitation>
|
||
) eleven-segmented; club well-developed, formed by apical three antennomeres, with sub-oval and basally truncate antennomeres 11 (
|
||
<figureCitation id="135798B5FFCFFFC4FBB2FCF4BF98FCDC" box="[1112,1220,797,820]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">
|
||
Fig. 2A;
|
||
<emphasis id="B9185822FFCFFFC4FB5DFCF4BF98FCDC" box="[1207,1220,797,820]" italics="true" pageId="1" pageNumber="102">a</emphasis>
|
||
</figureCitation>
|
||
11) much wider than preceding antennomeres. Maxillary palpi four-segmented; palpomere 1 minute, elongate, palpomere 2 pedunculate in basal half, strongly broadening toward apex, palpomere 3 subtriangular, approximately as wide and half as long as palpomere 4, palpomere 4 (
|
||
<figureCitation id="135798B5FFCFFFC4FBBFFC54BFFAFC3C" box="[1109,1190,957,980]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB45FC54BF81FC3C" box="[1199,1245,957,980]" italics="true" pageId="1" pageNumber="102">mp4</emphasis>
|
||
) largest, sub-oval, with elongate apical palpal cone.
|
||
</paragraph>
|
||
<paragraph id="8BD38430FFCFFFC4FCB0FC14BF7BFB1C" blockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
|
||
Pronotum (
|
||
<figureCitation id="135798B5FFCFFFC4FC39FC14BF79FBFC" box="[979,1061,1021,1044]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
) slightly elongate, widest at apical third; lateral margins rounded, sides convergent toward apex at apical two-fifths and narrowing toward base at basal third; disc with nude median (
|
||
<figureCitation id="135798B5FFCFFFC4FB4EFBB4BFA8FB9C" box="[1188,1268,1117,1140]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB17FBB4BE78FB9C" box="[1277,1316,1117,1140]" italics="true" pageId="1" pageNumber="102">maf</emphasis>
|
||
) and lateral antebasal (
|
||
<figureCitation id="135798B5FFCFFFC4FC45FB94BF59FB7C" box="[943,1029,1149,1172]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FBF9FB94BF73FB7C" box="[1043,1071,1149,1172]" italics="true" pageId="1" pageNumber="102">laf</emphasis>
|
||
) foveae subequal in size, lacking additional carinae, sulci or tubercles; paranotal carinae either short, or lacking; lateral procoxal foveae present, moderately separated.
|
||
</paragraph>
|
||
<paragraph id="8BD38430FFCFFFC4FCB0FB14BF97FA1C" blockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
|
||
Elytra (
|
||
<figureCitation id="135798B5FFCFFFC4FC40FB14B8A0FAFC" box="[938,1020,1277,1300]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
) slightly wider than long, widest point posterior to middle; each elytron with four small but distinct basal foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FBFBFAD4BF3AFABC" box="[1041,1126,1341,1364]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB98FAD4BFCFFABC" box="[1138,1171,1341,1364]" italics="true" pageId="1" pageNumber="102">bef</emphasis>
|
||
); sutural striae (
|
||
<figureCitation id="135798B5FFCFFFC4FAA9FAD4BECBFABC" box="[1347,1431,1341,1364]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FCD6FAB4B80DFA9C" box="[828,849,1373,1396]" italics="true" pageId="1" pageNumber="102">ss</emphasis>
|
||
) complete; discal striae absent; subhumeral foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FA87FAB4B8D7FA7C" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Figs 2B, 2D</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FC7CFA94B89DFA7C" box="[918,961,1405,1428]" italics="true" pageId="1" pageNumber="102">shef</emphasis>
|
||
) present; short marginal striae (
|
||
<figureCitation id="135798B5FFCFFFC4FAF8FA94BECBFA7C" box="[1298,1431,1405,1428]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Figs 2B, 2D</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FCD6FA74B806FA5C" box="[828,858,1437,1460]" italics="true" pageId="1" pageNumber="102">ms</emphasis>
|
||
) extending from subhumeral foveae to basal third of elytral length; posterolateral clefts (
|
||
<figureCitation id="135798B5FFCFFFC4FB46FA54BFA2FA3C" box="[1196,1278,1469,1492]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FAEDFA54BE7BFA3C" box="[1287,1319,1469,1492]" italics="true" pageId="1" pageNumber="102">plc</emphasis>
|
||
) broad and shallow. Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph id="8BD38430FFCFFFC4FCB0FA14BF72F93C" blockId="1.[828,1437,189,2068]" pageId="1" pageNumber="102">
|
||
Thorax with narrowly separated median mesoventral foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FC93F9F4B890F9DC" box="[889,972,1565,1588]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FC3FF9F4BF51F9DC" box="[981,1037,1565,1588]" italics="true" pageId="1" pageNumber="102">mmsf</emphasis>
|
||
); lateral mesoventral foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FAAEF9F4BECBF9DC" box="[1348,1431,1565,1588]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2D</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FCD6F9D4B834F9BC" box="[828,872,1597,1620]" italics="true" pageId="1" pageNumber="102">lmsf</emphasis>
|
||
) simple; lateral mesocoxal foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FB37F9D4BE6CF9BC" box="[1245,1328,1597,1620]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FAD0F9D4BE34F9BC" box="[1338,1384,1597,1620]" italics="true" pageId="1" pageNumber="102">lmcf</emphasis>
|
||
) and lateral metaventral foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FB88F9B4BFE4F99C" box="[1122,1208,1629,1652]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB29F9B4BFB0F99C" box="[1219,1260,1629,1652]" italics="true" pageId="1" pageNumber="102">lmtf</emphasis>
|
||
) present; lateral sutures separating meso- and metaventrite well-marked, fully developed; posterior margin of metaventrite with narrow split at middle.
|
||
</paragraph>
|
||
<paragraph id="8BD38430FFCFFFC7FCB0F934B91FF9B6" blockId="1.[828,1437,189,2068]" lastBlockId="2.[151,760,1479,2078]" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102">
|
||
Abdomen (
|
||
<figureCitation id="135798B5FFCFFFC4FC3EF934BF45F91C" box="[980,1049,1757,1780]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Figs 2</figureCitation>
|
||
A–C) dorsoventrally flattened, with generally rounded lateral margins; segments telescoping. Five tergites (IV–VIII) (
|
||
<figureCitation id="135798B5FFCFFFC4FBDBF8F4BFEDF8DC" box="[1073,1201,1821,1844]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Figs 2A, 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB50F8F4BE4CF8DC" box="[1210,1296,1821,1844]" italics="true" pageId="1" pageNumber="102">tIV–VIII</emphasis>
|
||
) visible, with broad paratergites on segments 1–4 (IV–VII); tergite 1 (IV) much longer than tergite 2 (V), sides narrowing toward base, lacking mediobasal foveae, with pair of basolateral foveae (
|
||
<figureCitation id="135798B5FFCFFFC4FC78F874B8BAF85C" box="[914,998,1949,1972]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FC18F874BF52F85C" box="[1010,1038,1949,1972]" italics="true" pageId="1" pageNumber="102">blf</emphasis>
|
||
), short triangular discal carinae (
|
||
<figureCitation id="135798B5FFCFFFC4FA9BF874B801F83C" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2B</figureCitation>
|
||
; dc), and broad, shallow transverse impression at base. Six sternites (III–VIII) (
|
||
<figureCitation id="135798B5FFCFFFC4FBDBF834BFDDF81C" box="[1073,1153,2013,2036]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="1" pageNumber="102">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCFFFC4FB61F834BFBAF81C" box="[1163,1254,2013,2036]" italics="true" pageId="1" pageNumber="102">sIII–VIII</emphasis>
|
||
) visible; sternites 1 (III) and 2 (IV) subequal in length along midline; sternite 2 (IV) lacking mediobasal foveae and basal impression, with small basolateral foveae. Foveal pattern of remaining abdominal segments barely visible based on available specimen, but one pair of basolateral foveae putatively present on each of tergites 2–4 (V–VII).
|
||
</paragraph>
|
||
<caption id="DF13D4B8FFCCFFC7FF7CFA85BFC8FA97" ID-DOI="http://doi.org/10.5281/zenodo.4505002" ID-Zenodo-Dep="4505002" box="[150,1172,1388,1407]" httpUri="https://zenodo.org/record/4505002/files/figure.png" pageId="2" pageNumber="103" startId="2.[150,181,1388,1407]" targetBox="[198,1389,197,1338]" targetPageId="2">
|
||
<paragraph id="8BD38430FFCCFFC7FF7CFA85BFC8FA97" blockId="2.[150,1172,1388,1407]" box="[150,1172,1388,1407]" pageId="2" pageNumber="103">
|
||
Fig. 1. Habitus of †
|
||
<taxonomicName id="4C6CFFB3FFCCFFC7FED7FA85B91CFA97" authority="Yin & Kurbatov & Cuccodoro & Cai, 2019" authorityName="Yin & Kurbatov & Cuccodoro & Cai" authorityYear="2019" box="[317,576,1388,1407]" class="Insecta" family="Staphylinidae" genus="Cretobrachygluta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="laurasiensis" status="gen. et sp. nov.">
|
||
<emphasis id="B9185822FFCCFFC7FED7FA85B91CFA97" box="[317,576,1388,1407]" italics="true" pageId="2" pageNumber="103">Cretobrachygluta laurasiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22BE559FFCCFFC7FDACFA85B99BFA97" box="[582,711,1388,1407]" pageId="2" pageNumber="103" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
A – dorsal view; B – ventral view. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD38430FFCCFFC7FF5FF98EBAF0F876" blockId="2.[151,760,1479,2078]" pageId="2" pageNumber="103">
|
||
Legs moderately elongate; all three pairs of coxae contiguous; metacoxae (
|
||
<figureCitation id="135798B5FFCCFFC7FE86F96EBA9FF976" box="[364,451,1671,1694]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="2" pageNumber="103">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B9185822FFCCFFC7FE3BF96EBAA4F976" box="[465,504,1671,1694]" italics="true" pageId="2" pageNumber="103">mtc</emphasis>
|
||
) moderately projecting posteriorly; dorsal margin of all trochanters short, base of femora thus close to coxal-trochanteral articulation. Tarsi three-segmented (
|
||
<figureCitation id="135798B5FFCCFFC7FEBDF90EBACAF916" box="[343,406,1767,1790]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
|
||
box;
|
||
<emphasis id="B9185822FFCCFFC7FE3FF90EB959F916" box="[469,517,1767,1790]" italics="true" pageId="2" pageNumber="103">t1–3</emphasis>
|
||
), with short tarsomere 1, and longer tarsomeres 2 and 3; tarsomere 2 distinctly shorter than tarsomere 3; tarsi each with one elongate claw (
|
||
<figureCitation id="135798B5FFCCFFC7FF75F8AEBBBEF8B6" box="[159,226,1863,1886]" captionStart="Fig" captionStartId="3.[151,182,1498,1517]" captionTargetBox="[151,1436,190,1468]" captionTargetId="figure-19@3.[151,1436,189,1468]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of †Cretobrachygluta laurasiensis gen. et sp. nov. A – left antenna; B–D – main body parts in dorsal (B), ventral (C), and lateral (D) view, with foveal pattern and other important characters annotated. Scale bar: 0.2 mm in A; 0.5 mm in B–D. Abbreviations: a1–2, 9–11 = antennomeres 1–2, 9–11; bef = basal elytral foveae; blf = basolateral fovea; dc = discal carina; gf = gular fovea; gr = gular ridge; laf = lateral antebasal fovea; lmcf = lateral mesocoxal fovea; lmsf = lateral mesoventral fovea; lmtf = lateral metaventral fovea; maf = median antebasal fovea; mmsf = median mesoventral foveae; mp4 = maxillary palpomere 4; ms = marginal stria; mtc = metacoxa; omc = ocular-mandibular carina; p = projection; plc = posterolateral cleft; s = spur; shef = subhumeral elytral fovea; sIII–VIII = sternites III–VIII; ss = sutural stria; tIV–VIII = tergites IV–VIII; tc = pretarsal claw; ts1–3 = tarsomeres 1–3; vf = vertexal fovea." figureDoi="http://doi.org/10.5281/zenodo.4505008" httpUri="https://zenodo.org/record/4505008/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
|
||
box;
|
||
<emphasis id="B9185822FFCCFFC7FECDF8AEBA66F8B6" box="[295,314,1863,1886]" italics="true" pageId="2" pageNumber="103">tc</emphasis>
|
||
). Male mesotrochanters and mesotibiae modified on ventral margin and at near apex, respectively; fore and hind legs simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376D7BBFFCCFFC7FF7DF84EBA39F7F6" pageId="2" pageNumber="103" type="etymology">
|
||
<paragraph id="8BD38430FFCCFFC7FF7DF84EBA39F7F6" blockId="2.[151,760,1479,2078]" pageId="2" pageNumber="103">
|
||
<emphasis id="B9185822FFCCFFC7FF7DF84EBA45F856" bold="true" box="[151,281,1959,1982]" pageId="2" pageNumber="103">Etymology.</emphasis>
|
||
The generic name is a combination of the prefix ‘
|
||
<emphasis id="B9185822FFCCFFC7FF0FF82EBA7DF836" box="[229,289,1991,2014]" italics="true" pageId="2" pageNumber="103">Creto</emphasis>
|
||
-’ derived from the Cretaceous, and
|
||
<taxonomicName id="4C6CFFB3FFCCFFC7FD48F82EBADFF816" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Brachygluta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9185822FFCCFFC7FD48F82EBB92F816" italics="true" pageId="2" pageNumber="103">Brachygluta</emphasis>
|
||
Thomson, 1859
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54D73A92FFCCFFC7FE65F80EBAE1F816" box="[399,445,2023,2046]" pageId="2" pageNumber="103">type</typeStatus>
|
||
genus of
|
||
<taxonomicName id="4C6CFFB3FFCCFFC7FDC6F80EB99EF816" authorityName="Raffray" authorityYear="1904" box="[556,706,2023,2046]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="tribe" tribe="Brachyglutini">Brachyglutini</taxonomicName>
|
||
. The gender is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |