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<document id="4AE55179C496E8A2D307F9C374D61463" ID-CLB-Dataset="272446" ID-DOI="10.1093/zoolinnean/zlad043" ID-GBIF-Dataset="81098796-8acf-4360-a625-ad76a281e99e" ID-ISSN="0024-4082" ID-Zenodo-Dep="8426263" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1696940821456" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan C. &amp; Cerdeño, Esperanza" docDate="2023" docId="03B17549E920476B9EFDFA19FABEFD94" docLanguage="en" docName="zlad043.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad043" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Protypotherium claudum Ameghino 1889" docType="treatment" docVersion="7" lastPageNumber="432" masterDocId="FF880D31E92F47649E20FFB4FF9EFFC7" masterDocTitle="Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" masterLastPageNumber="444" masterPageNumber="417" pageNumber="432" updateTime="1701999754155" updateUser="plazi">
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<mods:title id="525699AA3DE027D85ECC6D343C5BDDD1">Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae</mods:title>
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<treatment id="03B17549E920476B9EFDFA19FABEFD94" ID-DOI="http://doi.org/10.5281/zenodo.8426265" ID-GBIF-Taxon="213996077" ID-Zenodo-Dep="8426265" LSID="urn:lsid:plazi:treatment:03B17549E920476B9EFDFA19FABEFD94" httpUri="http://treatment.plazi.org/id/03B17549E920476B9EFDFA19FABEFD94" lastPageNumber="432" pageId="15" pageNumber="432">
<subSubSection id="C30297D4E920476B9EFDFA19FD10FA00" box="[221,654,1453,1480]" pageId="15" pageNumber="432" type="nomenclature">
<paragraph id="8BA7C45FE920476B9EFDFA19FD10FA00" blockId="15.[221,654,1453,1480]" box="[221,654,1453,1480]" pageId="15" pageNumber="432">
<heading id="D0EF7333E920476B9EFDFA19FD10FA00" bold="true" box="[221,654,1453,1480]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="432" reason="2">
<taxonomicName id="4C18BFDCE920476B9EFDFA19FD10FA00" authority="Ameghino, 1889" authorityName="Ameghino" authorityYear="1889" box="[221,654,1453,1480]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE920476B9EFDFA19FD10FA00" bold="true" box="[221,654,1453,1480]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9EFDFA19FE44FA0F" bold="true" box="[221,474,1453,1480]" italics="true" pageId="15" pageNumber="432">Protypotherium claudum</emphasis>
<bibRefCitation id="EF89B9AEE920476B9FFEFA19FD10FA00" author="Ameghino F" box="[478,654,1453,1479]" pageId="15" pageNumber="432" pagination="1 - 1028" refId="ref21871" refString="Ameghino F. Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias Cordoba 1889; 6: 1 - 1028, atlas, 98 pls." type="journal article" year="1889">Ameghino, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30297D4E920476B9E51FA60FABEFD94" pageId="15" pageNumber="432" type="description">
<paragraph id="8BA7C45FE920476B9E51FA60FDCBFA2B" blockId="15.[113,762,1492,1579]" box="[113,597,1492,1516]" pageId="15" pageNumber="432">
(
<figureCitation id="1323D8DAE920476B9E5CFA60FF7AFA2B" box="[124,228,1492,1516]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8BD</figureCitation>
; Supporting Information, Table S2)
</paragraph>
<paragraph id="8BA7C45FE920476B9EADFA47FEF6F9EC" blockId="15.[113,762,1492,1579]" pageId="15" pageNumber="432">
1889
<taxonomicName id="4C18BFDCE920476B9EE7FA47FDB3F9CC" authority="Ameghino" authorityName="Ameghino" authorityYear="1889" box="[199,557,1523,1547]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE920476B9EE7FA47FE25F9CC" box="[199,443,1523,1547]" italics="true" pageId="15" pageNumber="432">Protypotherium claudum</emphasis>
Ameghino
</taxonomicName>
, p. 480 (original description), pl. 14, fig. 22.
</paragraph>
<paragraph id="8BA7C45FE920476B9E51F9F8FD2AF964" blockId="15.[113,762,1612,1699]" pageId="15" pageNumber="432">
<materialsCitation id="3B70CE02E920476B9E51F9F8FD2AF964" ID-GBIF-Occurrence="4425895306" collectionCode="MACN-A" country="Argentina" location="Early - Middle Miocene" pageId="15" pageNumber="432" specimenCode="MACN-A 551" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<emphasis id="B96C184DE920476B9E51F9F8FF4FF9A3" box="[113,209,1612,1636]" italics="true" pageId="15" pageNumber="432">
<typeStatus id="54A37AFDE920476B9E51F9F8FF52F9A3" box="[113,204,1612,1636]" pageId="15" pageNumber="432" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBE6C24E920476B9EFAF9F8FEEEF9A3" box="[218,368,1612,1636]" collectionCode="MACN-A" pageId="15" pageNumber="432">MACN-A 551</specimenCode>
(
<figureCitation id="1323D8DAE920476B9FA3F9F8FE53F9A3" box="[387,461,1612,1636]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8B</figureCitation>
), right mandibular fragment with p2 (alveolus)p3m2 (see
<bibRefCitation id="EF89B9AEE920476B9FE0F9DFFD01F944" author="Fernandez M &amp; Fernicola JC &amp; Cerdeno E" box="[448,671,1643,1667]" pageId="15" pageNumber="432" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
Fernández
<emphasis id="B96C184DE920476B9C14F9D8FDFBF944" box="[564,613,1643,1667]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
2018
</bibRefCitation>
),
<location id="8EC79284E920476B9C98F9DFFE83F964" LSID="urn:lsid:plazi:treatment:03B17549E920476B9EFDFA19FABEFD94:8EC79284E920476B9C98F9DFFE83F964" country="Argentina" name="Early - Middle Miocene" pageId="15" pageNumber="432" stateProvince="Santa Cruz">Early Middle Miocene</location>
, SCF,
<collectingRegion id="49DC0ABDE920476B9F7DF93FFDB0F964" box="[349,558,1675,1699]" country="Argentina" name="Santa Cruz" pageId="15" pageNumber="432">Santa Cruz Province</collectingRegion>
(
<collectingCountry id="F30F84CFE920476B9C1AF93FFD36F964" box="[570,680,1675,1699]" name="Argentina" pageId="15" pageNumber="432">Argentina</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA7C45FE920476B9E51F970FD12F8FD" blockId="15.[113,763,1732,1850]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9E51F970FEB5F91B" box="[113,299,1732,1756]" italics="true" pageId="15" pageNumber="432">Referred materials:</emphasis>
MLP 73-VII-6-4 d (ex
<taxonomicName id="4C18BFDCE920476B9C00F970FD29F91B" authorityName="Ameghino" authorityYear="1887" box="[544,695,1732,1756]" class="Mammalia" family="Interatheriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="subFamily" subFamily="Interatheriinae">Interatheriinae</taxonomicName>
indet;
<figureCitation id="1323D8DAE920476B9E51F957FF58F93C" box="[113,198,1763,1787]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8C</figureCitation>
), less mandibular fragment with p2 (broken)m2 (trigonid), and MPEF-PV 1394 (
<figureCitation id="1323D8DAE920476B9FE2F8B7FD92F8DC" box="[450,524,1795,1819]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8D</figureCitation>
), incomplete mandible with less i1dp1p24, and right i1 and dp1p2m3.
</paragraph>
<paragraph id="8BA7C45FE920476B9E51F8EFFBA1FE44" blockId="15.[113,765,1883,1970]" lastBlockId="15.[810,1462,144,387]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9E51F8EFFDDBF8B4" box="[113,581,1883,1907]" italics="true" pageId="15" pageNumber="432">
Extended diagnosis:
<taxonomicName id="4C18BFDCE920476B9F69F8EFFDDBF8B4" authorityName="Ameghino" authorityYear="1889" box="[329,581,1883,1907]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="claudum">Protypotherium claudum</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C18BFDCE920476B9CFFF8E8FF67F854" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE920476B9CFFF8E8FF67F854" italics="true" pageId="15" pageNumber="432">Pr. compressidens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9F2DF8CFFE0BF855" authorityName="Ameghino" authorityYear="1885" box="[269,405,1915,1938]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE920476B9F2DF8CFFE0BF855" box="[269,405,1915,1938]" italics="true" pageId="15" pageNumber="432">Pr. antiquum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9F87F8CFFD82F854" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[423,540,1915,1939]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE920476B9F87F8CFFD82F854" box="[423,540,1915,1939]" italics="true" pageId="15" pageNumber="432">Pr. australe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9C10F8CFFD52F854" authorityName="Ameghino" authorityYear="1887" box="[560,716,1915,1939]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE920476B9C10F8CFFD52F854" box="[560,716,1915,1939]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9CFFF8CFFF49F875" authorityName="Ameghino" authorityYear="1902" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9CFFF8CFFF49F875" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9EC5F82FFEFCF875" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[229,354,1947,1970]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE920476B9EC5F82FFEFCF875" box="[229,354,1947,1970]" italics="true" pageId="15" pageNumber="432">Pr. minutum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C18BFDCE920476B9FBDF82FFDDDF875" box="[413,579,1946,1970]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE920476B9FBDF82FFDDDF875" box="[413,579,1946,1970]" italics="true" pageId="15" pageNumber="432">Pr. colloncurensis</emphasis>
</taxonomicName>
by p23 without the entoflexid and with the ectoflexid wider and slightly more mesially placed, and p4m3 with shallower entoflexid; from
<taxonomicName id="4C18BFDCE920476B9D0AFF7BFC06FF20" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[810,920,207,231]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE920476B9D0AFF7BFC06FF20" box="[810,920,207,231]" italics="true" pageId="15" pageNumber="432">Pr. australe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE920476B9DEDFF7BFBCBFF20" authorityName="Cabrera and Kraglievich" authorityYear="1931" box="[973,1109,207,231]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="distinctum">
<emphasis id="B96C184DE920476B9DEDFF7BFBCBFF20" box="[973,1109,207,231]" italics="true" pageId="15" pageNumber="432">Pr. distinctum</emphasis>
</taxonomicName>
by having a smaller size; from
<taxonomicName id="4C18BFDCE920476B9BB8FF7BFC2CFEC1" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE920476B9BB8FF7BFC2CFEC1" italics="true" pageId="15" pageNumber="432">Pr. compressidens</emphasis>
</taxonomicName>
by having relatively wider lower cheek teeth; from
<taxonomicName id="4C18BFDCE920476B9D0AFEBAFC32FEE2" authorityName="Ameghino" authorityYear="1885" box="[810,940,270,293]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE920476B9D0AFEBAFC32FEE2" box="[810,940,270,293]" italics="true" pageId="15" pageNumber="432">Pr. antiquum</emphasis>
</taxonomicName>
by the talonid of m3 without labial groove; from
<taxonomicName id="4C18BFDCE920476B9D0AFE99FC4EFE82" box="[810,976,301,325]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE920476B9D0AFE99FC4EFE82" box="[810,976,301,325]" italics="true" pageId="15" pageNumber="432">Pr. colloncurensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE920476B9A26FE99FB10FE82" authorityName="Ameghino" authorityYear="1902" box="[1030,1166,301,325]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9A26FE99FB10FE82" box="[1030,1166,301,325]" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
by the talonid of p24 sub-triangular in outline; and from
<taxonomicName id="4C18BFDCE920476B9A43FEF9FB77FEA3" authorityName="Ameghino" authorityYear="1902" box="[1123,1257,332,356]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9A43FEF9FB77FEA3" box="[1123,1257,332,356]" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
by the talonid of p4 narrower than the trigonid.
</paragraph>
<paragraph id="8BA7C45FE920476B9D0AFE11FB5DFDDC" blockId="15.[810,1459,421,539]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AFE11FB56FE7B" box="[810,1224,421,445]" italics="true" pageId="15" pageNumber="432">Geographic and stratigraphic provenance:</emphasis>
<collectingRegion id="49DC0ABDE920476B9AFAFE11FA2DFE7A" box="[1242,1459,421,445]" country="Argentina" name="Santa Cruz" pageId="15" pageNumber="432">Santa Cruz Province</collectingRegion>
(
<collectingCountry id="F30F84CFE920476B9D11FE70FC01FE1B" box="[817,927,452,476]" name="Argentina" pageId="15" pageNumber="432">Argentina</collectingCountry>
), SCF, EarlyMiddle Miocene, Santacrucian SALMA;
<collectingRegion id="49DC0ABDE920476B9DB2FE57FBFBFE3C" box="[914,1125,483,508]" country="Argentina" name="Rio Negro" pageId="15" pageNumber="432">Río Negro Province</collectingRegion>
(
<collectingCountry id="F30F84CFE920476B9A56FE57FB7AFE3C" box="[1142,1252,483,507]" name="Argentina" pageId="15" pageNumber="432">Argentina</collectingCountry>
),
<collectorName id="26EDA189E920476B9AD8FE57FA2EFE3C" box="[1272,1456,483,507]" pageId="15" pageNumber="432">Collón Curá Fm.</collectorName>
, Middle Miocene, Colloncuran SALMA.
</paragraph>
<paragraph id="8BA7C45FE920476B9D0AFD89FABEFD94" blockId="15.[810,1312,572,596]" box="[810,1312,572,596]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AFD89FC22FD93" box="[810,956,573,596]" italics="true" pageId="15" pageNumber="432">Measurements:</emphasis>
Supporting Information, Table S2.
</paragraph>
</subSubSection>
</treatment>
</document>