156 lines
19 KiB
XML
156 lines
19 KiB
XML
<document id="EE179EDF65A4E8FF04605C1894C004EA" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B0DFFFBFF79F2BAFEB8FA9C" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena buloba Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="105" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="105" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CF47622A5F79A0169771295362E18741">
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<mods:title id="541174CF231D7D6AFDE2881E8792953E">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:name id="F7CD009C116BE65E3ABF6AEA5680AB9D" type="personal">
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<mods:roleTerm id="5F1C3238796C5E224C7A8022407F55C4">Author</mods:roleTerm>
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<mods:namePart id="3026CA0D4F756873FA9FCEFAB433EDDA">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="86EEF781B700D0F61F00F7682F9E2D8C">text</mods:typeOfResource>
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<mods:title id="CF61FE1A9D6BABF7430BEC15C91635F5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DBB7E4373167BEB38DF6B40C27237EF8">
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<mods:date id="4791A7C3A86FC76F4D81B7D50F76B834">2011</mods:date>
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<mods:number id="3DD2FA0D7F139F91DDCA7829AE006E9A">2011-07-08</mods:number>
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<mods:detail id="A85DFCE77386CB7365BCC75912520748" type="volume">
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<mods:number id="02220BDAE90F9471223706CB9942709A">2944</mods:number>
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<mods:url id="5F3DEED76D6F78A0F985856E994DEE26">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="4F8D5925A91D1118F49F71D9368B6705">journal article</mods:classification>
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<mods:identifier id="F46AD97782838E54640780FECA099121" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="DAF22507C103FBF3CF6B72B1A3347FBF" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55B0DFFFBFF79F2BAFEB8FA9C" ID-DOI="http://doi.org/10.5281/zenodo.5291702" ID-Zenodo-Dep="5291702" LSID="urn:lsid:plazi:treatment:03B087E55B0DFFFBFF79F2BAFEB8FA9C" httpUri="http://treatment.plazi.org/id/03B087E55B0DFFFBFF79F2BAFEB8FA9C" lastPageNumber="105" pageId="104" pageNumber="105">
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<subSubSection id="C30365785B0DFFFBFF79F2BAFDBEFF27" box="[151,520,151,177]" pageId="104" pageNumber="105" type="nomenclature">
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<paragraph id="8BA636F35B0DFFFBFF79F2BAFDBEFF27" blockId="104.[151,520,151,210]" box="[151,520,151,177]" pageId="104" pageNumber="105">
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<heading id="D0EE819F5B0DFFFBFF79F2BAFDBEFF27" bold="true" box="[151,520,151,177]" fontSize="11" level="1" pageId="104" pageNumber="105" reason="1">
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<emphasis id="B96DEAE15B0DFFFBFF79F2BAFDBEFF27" bold="true" box="[151,520,151,177]" pageId="104" pageNumber="105">
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<taxonomicName id="4C194D705B0DFFFBFF79F2BAFEDFFF27" ID-CoL="3N372" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,361,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="buloba" status="sp. nov.">
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<emphasis id="B96DEAE15B0DFFFBFF79F2BAFEDFFF27" bold="true" box="[151,361,151,177]" italics="true" pageId="104" pageNumber="105">Hydraena buloba</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B0DFFFBFE96F2BAFDBEFF27" box="[376,520,151,177]" pageId="104" pageNumber="105" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785B0DFFFBFF79F297FE34FF44" box="[151,386,186,210]" pageId="104" pageNumber="105" type="description">
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||
<paragraph id="8BA636F35B0DFFFBFF79F297FE34FF44" blockId="104.[151,520,151,210]" box="[151,386,186,210]" pageId="104" pageNumber="105">
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(
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<figureCitation id="13222A765B0DFFFBFF71F297FEBDFF44" box="[159,267,186,210]" captionStart="FIGURE 182" captionStartId="281.[151,250,1753,1776]" captionTargetBox="[245,1339,211,1724]" captionTargetId="figure-26@281.[231,1355,193,1732]" captionTargetPageId="281" captionText="FIGURE 182. Hydraena buloba, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283756" httpUri="https://zenodo.org/record/5283756/files/figure.png" pageId="104" pageNumber="105">Figs. 182</figureCitation>
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,
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<figureCitation id="13222A765B0DFFFBFEF8F297FEF4FF44" box="[278,322,186,210]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="104" pageNumber="105">184</figureCitation>
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,
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<figureCitation id="13222A765B0DFFFBFEA0F297FECFFF44" box="[334,377,186,210]" captionStart="FIGURES 515–518" captionStartId="404.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1611]" captionTargetId="figure-23@404.[165,1421,193,1629]" captionTargetPageId="404" captionText="FIGURES 515–518. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284127" httpUri="https://zenodo.org/record/5284127/files/figure.png" pageId="104" pageNumber="105">518</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785B0DFFFBFF79F32EFC6CFEA8" pageId="104" pageNumber="105" type="materials_examined">
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<paragraph id="8BA636F35B0DFFFBFF79F32EFC6CFEA8" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">
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Type Material.
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<materialsCitation id="3B713CAE5B0DFFFBFEBFF32EFC6CFEA8" collectingDate="1965-01-08" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1128" latitude="-6.8" location="Vagau" longLatPrecision="1303" longitude="146.8" municipality="Herzog Mts." pageId="104" pageNumber="105" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
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||
<typeStatus id="54A288515B0DFFFBFEBFF32EFE08FE8D" box="[337,446,259,283]" pageId="104" pageNumber="105" type="holotype">
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||
<emphasis id="B96DEAE15B0DFFFBFEBFF32EFE08FE8D" bold="true" box="[337,446,259,283]" pageId="104" pageNumber="105">Holotype</emphasis>
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||
</typeStatus>
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(male):
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<emphasis id="B96DEAE15B0DFFFBFDC6F32EFCB7FE8D" bold="true" box="[552,769,258,283]" pageId="104" pageNumber="105">
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||
<collectingRegion id="49DDF8115B0DFFFBFDC6F32EFD4DFE8D" box="[552,763,258,283]" country="Papua New Guinea" name="Morobe" pageId="104" pageNumber="105">Morobe Province</collectingRegion>
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:
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</emphasis>
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<collectingMunicipality id="6BC2AC895B0DFFFBFCFCF32EFC17FE8C" box="[786,929,259,283]" pageId="104" pageNumber="105">Herzog Mts.</collectingMunicipality>
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,
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<location id="8EC660285B0DFFFBFC40F32EFC42FE8D" LSID="urn:lsid:plazi:treatment:03B087E55B0DFFFBFF79F2BAFEB8FA9C:8EC660285B0DFFFBFC40F32EFC42FE8D" box="[942,1012,259,283]" latitude="-6.8" longLatPrecision="1303" longitude="146.8" municipality="Herzog Mts." name="Vagau" pageId="104" pageNumber="105" stateProvince="Morobe">Vagau</location>
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[Wagau], ex shallow pools in marsh,
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<quantity id="4CE19B165B0DFFFBFF79F30BFF5BFEA8" box="[151,237,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="104" pageNumber="105" unit="m" value="1128.0">
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||
<elevation id="0034D1C05B0DFFFBFF79F30BFF5BFEA8" box="[151,237,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="104" pageNumber="105" unit="m" value="1128.0">1128 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B0DFFFBFF17F30BFEE4FEA8" box="[249,338,294,318]" degrees="6" direction="south" minutes="48" orientation="latitude" pageId="104" pageNumber="105" precision="925" value="-6.8">6° 48' S</geoCoordinate>
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,
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||
<geoCoordinate id="EE2D50345B0DFFFBFEB0F30BFE63FEA8" box="[350,469,294,318]" degrees="146" direction="east" minutes="48" orientation="longitude" pageId="104" pageNumber="105" precision="925" value="146.8">146° 48' E</geoCoordinate>
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,
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<date id="FFA710335B0DFFFBFE0FF30BFD89FEA8" box="[481,575,294,318]" pageId="104" pageNumber="105" value="1965-01-08">
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||
<collectingDate id="EFE3E9DB5B0DFFFBFE0FF30BFD89FEA8" box="[481,575,294,318]" pageId="104" pageNumber="105" value="1965-01-08">8 i 1965</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B0DFFFBFDA5F30AFD46FEA8" box="[587,752,294,318]" pageId="104" pageNumber="105">M. E. Bacchus</collectorName>
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(MEB 144) (
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<collectionCode id="ED08AE365B0DFFFBFC68F30BFC78FEA8" box="[902,974,294,318]" pageId="104" pageNumber="105">NHM</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30365785B0DFFFBFF29F367FCEAFE58" pageId="104" pageNumber="105" type="diagnosis">
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<paragraph id="8BA636F35B0DFFFBFF29F367FCEAFE58" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">
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<emphasis id="B96DEAE15B0DFFFBFF29F367FE63FEF5" bold="true" box="[199,469,330,355]" pageId="104" pageNumber="105">Differential Diagnosis.</emphasis>
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Recognized from the two other members of the
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<emphasis id="B96DEAE15B0DFFFBFBE0F366FB2BFEF5" box="[1038,1181,331,355]" italics="true" pageId="104" pageNumber="105">Luridipennis</emphasis>
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group in
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<collectingCountry id="F30E76635B0DFFFBFAFCF367FAFCFEF4" box="[1298,1354,330,354]" name="Papua New Guinea" pageId="104" pageNumber="105">PNG</collectingCountry>
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by the more robust body, the more markedly angulate sides of the very transverse pronotum (PL/PW ca. 0.50), and the striate-impressed elytral series (
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<figureCitation id="13222A765B0DFFFBFDEEF3BFFDDBFE3C" box="[512,621,402,426]" captionStart="FIGURE 178" captionStartId="278.[151,250,1739,1762]" captionTargetBox="[220,1367,193,1718]" captionTargetId="figure-29@278.[220,1367,193,1718]" captionTargetPageId="278" captionText="FIGURE 178. Hydraena impercepta, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5299418" httpUri="https://zenodo.org/record/5299418/files/figure.png" pageId="104" pageNumber="105">Figs. 178</figureCitation>
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,
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<figureCitation id="13222A765B0DFFFBFD94F3BFFD10FE3C" box="[634,678,402,426]" captionStart="FIGURE 179" captionStartId="279.[151,250,1753,1776]" captionTargetBox="[209,1373,204,1724]" captionTargetId="figure-26@279.[199,1387,193,1732]" captionTargetPageId="279" captionText="FIGURE 179. Hydraena cavifrons, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5299422" httpUri="https://zenodo.org/record/5299422/files/figure.png" pageId="104" pageNumber="105">179</figureCitation>
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,
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<figureCitation id="13222A765B0DFFFBFD5AF3BFFD54FE3C" box="[692,738,402,426]" captionStart="FIGURE 182" captionStartId="281.[151,250,1753,1776]" captionTargetBox="[245,1339,211,1724]" captionTargetId="figure-26@281.[231,1355,193,1732]" captionTargetPageId="281" captionText="FIGURE 182. Hydraena buloba, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283756" httpUri="https://zenodo.org/record/5283756/files/figure.png" pageId="104" pageNumber="105">182</figureCitation>
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). The pro- and metatibiae of males differ in all three species, and the male genitalia distinctively differ (
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<figureCitation id="13222A765B0DFFFBFD9DF39BFCA0FE58" box="[627,790,438,462]" captionStart="FIGURES 180–181" captionStartId="280.[151,263,1650,1673]" captionTargetBox="[250,1331,205,1619]" captionTargetId="figure-42@280.[241,1346,193,1629]" captionTargetPageId="280" captionText="FIGURES 180–181. Aedeagi. 180. H. impercepta, non-type specimen from Moitaka, 7 mi. N. of Port Moresby. 181. H. cavifrons, holotype." figureDoi="http://doi.org/10.5281/zenodo.5283750" httpUri="https://zenodo.org/record/5283750/files/figure.png" pageId="104" pageNumber="105">Figs. 180, 181</figureCitation>
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,
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<figureCitation id="13222A765B0DFFFBFCCCF39BFCE6FE58" box="[802,848,438,462]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="104" pageNumber="105">184</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30365785B0DFFFBFF29F3F7FD38FB08" pageId="104" pageNumber="105" type="description">
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||
<paragraph id="8BA636F35B0DFFFBFF29F3F7FD53FDAC" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">
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||
<emphasis id="B96DEAE15B0DFFFBFF29F3F7FEEEFE65" bold="true" box="[199,344,474,499]" inLineHeading="true" pageId="104" pageNumber="105" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B0DFFFBFE4AF3F7FDB0FE64" box="[420,518,474,498]" pageId="104" pageNumber="105" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.20/0.98; head 0.30/0.48; pronotum 0.49/0.75, PA 0.55, PB 0.64; elytra 1.43/0.98. Dorsum of head and pronotal disc piceous, remainder of dorsum dark brown; legs and maxillary palpi brown.
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</paragraph>
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<paragraph id="8BA636F35B0DFFFBFF29F06AFABBFC84" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">Frons punctures ca. 2–3xef; interstices shining, narrow walls to 1xpd. Clypeus laterally microreticulate and as coarsely punctate as frons, medially less coarsely punctate. Mentum shining, finely sparsely punctate; postmentum medially microreticulate, otherwise finely sparsely punctate, shining. Genae raised, lacking posterior ridge, midlongitudinally impressed. Pronotum densely coarsely punctate, punctures on disc deep, ca. 3–4xpd those of frons, interstices narrow walls to 1xpd, some interstices uniting to form short longitudinal lines; PF1 shallow, in midline extended to ca. midlength of pronotum; PF2 deep, oval, oblique; PF3 and PF4 deep, shallowly joined.</paragraph>
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<paragraph id="8BA636F35B0DFFFBFF29F133FB1BFCE8" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">Elytral punctures striate-impressed, ca. 1.5xpd of largest pronotal punctures. Intervals slightly raised, width ca. 1xpd, interstices between punctures of a row less, some punctures contiguous. Explanate margin wide. Apices in dorsal aspect separately rounded, in posterior aspect margins forming angle with one another.</paragraph>
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<paragraph id="8BA636F35B0DFFFBFF29F1A7FD38FB08" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">
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Metaventral disc densely punctate beneath hydrofuge pubescence, plaques very small, oval, located basolaterally on sides of deep, inverted V-shaped median depression. P1 laminate; median carina angulate in profile. P2 very narrow, l/w ca. 5/3, sides parallel, apex blunt. No carina between mesoventral intercoxal process and median depression. AIS flat, width at slightly arcuate posterior margin 3x P2. Abdominal apex asymmetrical, with short setae; last tergite notched on ventral face. Protrochanter weakly keeled. Protibia straight, slightly widened on medial margin at about distal 1/3. Mesotibia straight. Metafemur with brush of long setae on entire length of lower surface. Metatibia straight, medial margin slightly widened in distal 1/2, dorsomedian margin with brush of setae in distal 1/2. Aedeagus as illustrated (
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<figureCitation id="13222A765B0DFFFBFDF1F6ABFD34FB08" box="[543,642,1158,1182]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="104" pageNumber="105">Fig. 184</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30365785B0DFFFBFF29F686FBA1FB54" box="[199,1047,1194,1219]" pageId="104" pageNumber="105" type="etymology">
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||
<paragraph id="8BA636F35B0DFFFBFF29F686FBA1FB54" blockId="104.[151,1437,258,1290]" box="[199,1047,1194,1219]" pageId="104" pageNumber="105">
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||
<emphasis id="B96DEAE15B0DFFFBFF29F686FEFBFB55" bold="true" box="[199,333,1195,1219]" inLineHeading="true" pageId="104" pageNumber="105" reason="1">Etymology.</emphasis>
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Named in reference to the large rounded lobes of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B0DFFFBFF29F6E3FEB8FA9C" pageId="104" pageNumber="105" type="distribution">
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<paragraph id="8BA636F35B0DFFFBFF29F6E3FEB8FA9C" blockId="104.[151,1437,258,1290]" pageId="104" pageNumber="105">
|
||
<emphasis id="B96DEAE15B0DFFFBFF29F6E3FED6FB71" bold="true" box="[199,352,1230,1255]" inLineHeading="true" pageId="104" pageNumber="105" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B0DFFFBFD35F6E2FCBDFB71" box="[731,779,1231,1255]" pageId="104" pageNumber="105">type</typeStatus>
|
||
locality in the Herzog Mts. of Area 6; elevation
|
||
<quantity id="4CE19B165B0DFFFBFAA8F6E3FA2AFB70" box="[1350,1436,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="104" pageNumber="105" unit="m" value="1128.0">1128 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B0DFFFBFF71F6DFFEB4FA9C" box="[159,258,1266,1290]" captionStart="FIGURES 515–518" captionStartId="404.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1611]" captionTargetId="figure-23@404.[165,1421,193,1629]" captionTargetPageId="404" captionText="FIGURES 515–518. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284127" httpUri="https://zenodo.org/record/5284127/files/figure.png" pageId="104" pageNumber="105">Fig. 518</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |