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<document id="2F0F5F9801DF7A8BA220F65D63E990C0" ID-CLB-Dataset="94528" ID-DOI="10.11646/zootaxa.5102.1.1" ID-GBIF-Dataset="ad30faa6-eab1-4dae-b686-9e5fee56f55c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6245602" ID-ZooBank="BEA963B7-F7B9-458A-92EE-A4740AC390AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645601333553" checkinUser="plazi" docAuthor="Ward, Philip S." docDate="2022" docId="03A33567FFA354231CD081E1FCDB38A4" docLanguage="en" docName="zootaxa.5102.1.1.pdf" docOrigin="Zootaxa 5102 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tetraponera exactor Ward 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FF9A4D1FFFBD543D1C478336FFC13E45" masterDocTitle="The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="31" updateTime="1699316131195" updateUser="ExternalLinkService">
<mods:mods id="DE45EA96E136B1F9C9A008FA58411C86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CE324CB9B736B8C1530C0CD9489B7416">
<mods:title id="78FE2C5DEF333CDC67ABFC3EB9A6E407">The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species</mods:title>
</mods:titleInfo>
<mods:name id="2CD41972F03B5C8D6D518717CBE9D733" type="personal">
<mods:role id="8501831D8E21F57C775FDDFDBA002C10">
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<mods:namePart id="52C782B4AEC81F6C6A31E222549C87E6">Ward, Philip S.</mods:namePart>
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<mods:title id="E48E36DD86F8AF7E4B4E7949418E2DF7">Zootaxa</mods:title>
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<mods:part id="C385B9F8419D4DC751F5DCB4E54A18B5">
<mods:date id="6D005D4B3AA67DE33CC121467662140E">2022</mods:date>
<mods:detail id="DD5A63B24614050F8DA915CB90BA7E01" type="pubDate">
<mods:number id="CAE8A7A678F44BC50B546134F3C2331D">2022-02-23</mods:number>
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<mods:number id="34AE3403711480C865CF16EC6ECB060B">5102</mods:number>
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<mods:classification id="64F287274116B667FEB4267FB4275DD6">journal article</mods:classification>
<mods:identifier id="35AD88FDB8C1C2BFD01DEA936FC8C0CF" type="CLB-Dataset">94528</mods:identifier>
<mods:identifier id="F9330BDCA053DA173D9743511C7B3EE4" type="DOI">10.11646/zootaxa.5102.1.1</mods:identifier>
<mods:identifier id="8F05D259C57CFAF04ADA80B27AF035BD" type="GBIF-Dataset">ad30faa6-eab1-4dae-b686-9e5fee56f55c</mods:identifier>
<mods:identifier id="0A1A25E23D1CC8154D6AF8745ADEF5F0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DBAB8EEF28E1FD71FFE82400BD79F075" type="Zenodo-Dep">6245602</mods:identifier>
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<treatment id="03A33567FFA354231CD081E1FCDB38A4" ID-DOI="http://doi.org/10.5281/zenodo.6315493" ID-GBIF-Taxon="197227887" ID-Zenodo-Dep="6315493" LSID="urn:lsid:plazi:treatment:03A33567FFA354231CD081E1FCDB38A4" httpUri="http://treatment.plazi.org/id/03A33567FFA354231CD081E1FCDB38A4" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="C310D7FAFFA354231CD081E1FE313CB4" box="[151,496,727,753]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BB58471FFA354231CD081E1FE313CB4" blockId="30.[151,496,727,789]" box="[151,496,727,753]" pageId="30" pageNumber="31">
<heading id="D0FD331DFFA354231CD081E1FE313CB4" bold="true" box="[151,496,727,753]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="B97E5863FFA354231CD081E1FE313CB4" bold="true" box="[151,496,727,753]" pageId="30" pageNumber="31">
<taxonomicName id="4C0AFFF2FFA354231CD081E1FE4D3CB4" authority="Ward, 2022" authorityName="Ward" authorityYear="2022" box="[151,396,727,753]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="exactor" status="sp. nov.">
<emphasis id="B97E5863FFA354231CD081E1FE4D3CB4" bold="true" box="[151,396,727,753]" italics="true" pageId="30" pageNumber="31">Tetraponera exactor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DE518FFA354231DD481E1FE313CB4" box="[403,496,727,753]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA354231CD081CDFED83D50" box="[151,281,763,789]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BB58471FFA354231CD081CDFED83D50" blockId="30.[151,496,727,789]" box="[151,281,763,789]" pageId="30" pageNumber="31">
(
<figureCitation id="133198F4FFA354231CD881CDFF293D50" box="[159,232,763,789]" captionStart="FIGURES 7–9" captionStartId="14.[151,264,1692,1716]" captionTargetBox="[152,1437,181,1666]" captionTargetId="figure-29@14.[151,1437,181,1667]" captionTargetPageId="14" captionText="FIGURES 7–9. Tetraponera allaborans group, full-face (dorsal) view of head (a) and lateral profile of body (b). 7, T. dispar, worker, holotype, Republic of Congo (CASENT0888518); 8, T. emeryi, worker, South Africa (CASENT0220743); 9, T. exactor, alate queen, holotype, South Africa (CASENT0794350). Images from AntWeb (www.antweb.org); photographers Peter Hawkes (7), Matthew Prebus (8, 9)." figureDoi="http://doi.org/10.5281/zenodo.6245620" httpUri="https://zenodo.org/record/6245620/files/figure.png" pageId="30" pageNumber="31">Figs 9</figureCitation>
,
<figureCitation id="133198F4FFA354231CB381CDFED03D50" box="[244,273,763,789]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[156,1433,182,1919]" captionTargetId="figure-17@23.[155,1433,181,1919]" captionTargetPageId="23" captionText="FIGURES 35–40. Distribution maps for Afrotropical Tetraponera. 35, T. clypeata (circles), T. tessmanni (triangles); 36, T. emeryi (circles), T. dispar (triangle); 37, T. exactor (circle), T. furtiva (triangle), T. liengmei (squares); 38, T. continua (circles), T. cortina (triangles), T. gerdae (squares); 39, T. pedana (circles), T. penzigi (triangles); 40, T. pumila (circles)." figureDoi="http://doi.org/10.5281/zenodo.6245653" httpUri="https://zenodo.org/record/6245653/files/figure.png" pageId="30" pageNumber="31">37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA354231CD08075FD553B48" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BB58471FFA354231CD08075FC6B3DC4" blockId="30.[151,1437,834,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231CD08075FE863D19" bold="true" box="[151,327,834,861]" pageId="30" pageNumber="31">Type material.</emphasis>
<materialsCitation id="3B628E2CFFA354231D178075FC6B3DC4" ID-GBIF-Occurrence="3861810368" collectingDate="1994-10-15" collectingDateMax="1994-10-17" collectingDateMin="1994-10-15" collectionCode="SAMC" collectorName="R. Danielsson" country="South Africa" latitude="-33.966667" location="Groot Rivier" longLatPrecision="1202" longitude="23.55" municipality="Nature's Valley" pageId="30" pageNumber="31" specimenCode="CASENT0794350" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B97E5863FFA354231D178075FDC23D19" box="[336,515,835,861]" italics="true" pageId="30" pageNumber="31">
<typeStatus id="54B13AD3FFA354231D178075FE763D18" box="[336,439,835,861]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
queen
</emphasis>
:
<collectingCountry id="F31DC4E1FFA354231E538075FD663D18" box="[532,679,835,861]" name="South Africa" pageId="30" pageNumber="31">South Africa</collectingCountry>
,
<collectingRegion id="49CE4A93FFA354231EF28075FC943D18" box="[693,853,835,861]" country="South Africa" name="Western Cape" pageId="30" pageNumber="31">Western Cape</collectingRegion>
,
<collectingMunicipality id="6BD11E0BFFA354231F248075FBD23D18" box="[867,1043,835,861]" pageId="30" pageNumber="31">Nature’s Valley</collectingMunicipality>
at
<location id="8ED5D2AAFFA35423187E8075FB0A3D18" LSID="urn:lsid:plazi:treatment:03A33567FFA354231CD081E1FCDB38A4:8ED5D2AAFFA35423187E8075FB0A3D18" box="[1081,1227,835,861]" country="South Africa" latitude="-33.966667" longLatPrecision="1202" longitude="23.55" municipality="Nature's Valley" name="Groot Rivier" pageId="30" pageNumber="31" stateProvince="Western Cape">Groot Rivier</location>
,
<geoCoordinate id="EE3EE2B6FFA35423189E8072FAF33D1A" box="[1241,1330,836,863]" degrees="33" direction="south" minutes="58" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-33.966667">33°58′S</geoCoordinate>
<geoCoordinate id="EE3EE2B6FFA35423197C8072FA593D1A" box="[1339,1432,836,863]" degrees="23" direction="east" minutes="33" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="23.55">23°33′E</geoCoordinate>
,
<date id="FFB4A2B1FFA354231CD08050FE8E3DC4" box="[151,335,870,897]" pageId="30" pageNumber="31" value="1994-10-15" valueMax="1994-10-17" valueMin="1994-10-15">
<collectingDate id="EFF05B59FFA354231CD08050FE8E3DC4" box="[151,335,870,897]" pageId="30" pageNumber="31" value="1994-10-15" valueMax="1994-10-17" valueMin="1994-10-15">15–17 Oct 1994</collectingDate>
</date>
,
<collectorName id="26FFE1A7FFA354231D1D8051FE383DC4" box="[346,505,871,897]" pageId="30" pageNumber="31">R. Danielsson</collectorName>
, loc. 21 (
<specimenCode id="DBAC2C0AFFA354231E188050FCF33DC4" box="[607,818,870,897]" collectionCode="CASENT" pageId="30" pageNumber="31">CASENT0794350</specimenCode>
) (
<collectionCode id="ED1B1CB4FFA354231F0F8051FC5F3DC4" box="[840,926,871,897]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="30" pageNumber="31" type="Museum">SAMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB58471FFA354231C8080BCFC743D8C" blockId="30.[151,1437,834,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231C8080BCFE063DE0" bold="true" box="[199,455,906,933]" pageId="30" pageNumber="31">Queen measurements</emphasis>
(n = 1). HW 0.55, HL 0.70, LHT 0.43, CI 0.79, FCI 0.19, REL 0.29, REL2 0.37, SI 0.56, SI3 1.50, FI 0.37, PLI 0.53, PWI 0.46, LHT/HW 0.79, CSC 4, MSC16.
</paragraph>
<paragraph id="8BB58471FFA354231C8080E4FD553B48" blockId="30.[151,1437,834,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231C8080E4FE4B3DA9" bold="true" box="[199,394,978,1004]" pageId="30" pageNumber="31">Queen diagnosis</emphasis>
. Relatively small species (
<collectionCode id="ED1B1CB4FFA354231EFF80E5FD273DA8" box="[696,742,979,1005]" pageId="30" pageNumber="31">HW</collectionCode>
0.55); head moderately elongate, with broadly rounded sides; median clypeal lobe bluntly bidentate; frontal carinae well separated, the distance between them much greater than basal scape width (
<collectionCode id="ED1B1CB4FFA354231D2F872DFE543A70" box="[360,405,1051,1077]" pageId="30" pageNumber="31">FCI</collectionCode>
0.19); scape short (SI 0.56), somewhat expanded apically; profemur slender (FI 0.37); dorsal face of propodeum rounding gently into declivitous face, both of similar length; petiole elongate (
<collectionCode id="ED1B1CB4FFA3542318AA8709FAD83A1C" box="[1261,1305,1087,1113]" pageId="30" pageNumber="31">PLI</collectionCode>
0.53), with differentiated anterior peduncle and weak anteroventral tooth (
<figureCitation id="133198F4FFA354231F1D8755FC6E3A3B" box="[858,943,1123,1150]" captionStart="FIGURES 7–9" captionStartId="14.[151,264,1692,1716]" captionTargetBox="[152,1437,181,1666]" captionTargetId="figure-29@14.[151,1437,181,1667]" captionTargetPageId="14" captionText="FIGURES 7–9. Tetraponera allaborans group, full-face (dorsal) view of head (a) and lateral profile of body (b). 7, T. dispar, worker, holotype, Republic of Congo (CASENT0888518); 8, T. emeryi, worker, South Africa (CASENT0220743); 9, T. exactor, alate queen, holotype, South Africa (CASENT0794350). Images from AntWeb (www.antweb.org); photographers Peter Hawkes (7), Matthew Prebus (8, 9)." figureDoi="http://doi.org/10.5281/zenodo.6245620" httpUri="https://zenodo.org/record/6245620/files/figure.png" pageId="30" pageNumber="31">Fig. 9b</figureCitation>
); postpetiole longer than broad, lacking anteroventral process. Integument smooth and shiny, with scattered fine punctures. Standing pilosity present on frontal carinae and as two long posterolateral pairs (
<quantity id="4CF22994FFA354231ECC879DFCF13A80" box="[651,816,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="1.5500000000000003" metricValueMax="1.7000000000000002" metricValueMin="1.4000000000000001" pageId="30" pageNumber="31" unit="mm" value="0.155" valueMax="0.17" valueMin="0.14">0.14–0.17 mm</quantity>
long) on upper half of head; scattered standing pilosity also present on pronotum (10 setae), mesoscutum (5), propodeum (1), petiole (4), postpetiole (4), and succeeding abdominal segments. Light yellowish-brown.
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA354231C808620FD6C384F" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BB58471FFA354231C808620FD6C384F" blockId="30.[151,1437,834,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231C808620FE893B75" bold="true" box="[199,328,1302,1328]" pageId="30" pageNumber="31">Comments</emphasis>
. This species is evidently closely related to
<taxonomicName id="4C0AFFF2FFA354231F038621FBF43B74" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[836,1077,1303,1329]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA354231F038621FBF43B74" box="[836,1077,1303,1329]" italics="true" pageId="30" pageNumber="31">Tetraponera clypeata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0AFFF2FFA3542318288621FB193B75" baseAuthorityName="Forel" baseAuthorityYear="1911" box="[1135,1240,1303,1328]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B97E5863FFA3542318288621FB193B75" box="[1135,1240,1303,1328]" italics="true" pageId="30" pageNumber="31">T. emeryi</emphasis>
</taxonomicName>
. The queen of
<taxonomicName id="4C0AFFF2FFA3542319CF8621FF2A3B11" authorityName="Ward" authorityYear="2022" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="exactor">
<emphasis id="B97E5863FFA3542319CF8621FF2A3B11" italics="true" pageId="30" pageNumber="31">T. exactor</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="4C0AFFF2FFA354231DA3860DFD9E3B10" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[484,607,1339,1365]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA354231DA3860DFD9E3B10" box="[484,607,1339,1365]" italics="true" pageId="30" pageNumber="31">T. clypeata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0AFFF2FFA354231ED0860DFCC13B11" baseAuthorityName="Forel" baseAuthorityYear="1911" box="[663,768,1339,1364]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B97E5863FFA354231ED0860DFCC13B11" box="[663,768,1339,1364]" italics="true" pageId="30" pageNumber="31">T. emeryi</emphasis>
</taxonomicName>
by its smaller size (HW 0.55, versus 0.63–0.80 [n = 12] in queens of the other species), more broadly rounded head (CI 0.79, versus 0.67–0.72), more widely separated frontal carinae (FCI 0.19, versus 0.11–0.17), shorter but apically expanded scape (SI 0.56, versus 0.57–0.66), smoother integument, and more abundant standing pilosity. In
<taxonomicName id="4C0AFFF2FFA354231EB68691FCA33B85" authorityName="Ward" authorityYear="2022" box="[753,866,1447,1472]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="exactor">
<emphasis id="B97E5863FFA354231EB68691FCA33B85" box="[753,866,1447,1472]" italics="true" pageId="30" pageNumber="31">T. exactor</emphasis>
</taxonomicName>
standing hairs are conspicuous on the mesosoma, petiole and postpetiole (MSC 16;
<figureCitation id="133198F4FFA354231E5E86FDFDAA3BA3" box="[537,619,1483,1510]" captionStart="FIGURES 7–9" captionStartId="14.[151,264,1692,1716]" captionTargetBox="[152,1437,181,1666]" captionTargetId="figure-29@14.[151,1437,181,1667]" captionTargetPageId="14" captionText="FIGURES 7–9. Tetraponera allaborans group, full-face (dorsal) view of head (a) and lateral profile of body (b). 7, T. dispar, worker, holotype, Republic of Congo (CASENT0888518); 8, T. emeryi, worker, South Africa (CASENT0220743); 9, T. exactor, alate queen, holotype, South Africa (CASENT0794350). Images from AntWeb (www.antweb.org); photographers Peter Hawkes (7), Matthew Prebus (8, 9)." figureDoi="http://doi.org/10.5281/zenodo.6245620" httpUri="https://zenodo.org/record/6245620/files/figure.png" pageId="30" pageNumber="31">Fig. 9b</figureCitation>
) whereas in queens of
<taxonomicName id="4C0AFFF2FFA354231F3486FDFC2E3BA0" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[883,1007,1483,1509]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA354231F3486FDFC2E3BA0" box="[883,1007,1483,1509]" italics="true" pageId="30" pageNumber="31">T. clypeata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0AFFF2FFA35423186F86FDFB503BA1" baseAuthorityName="Forel" baseAuthorityYear="1911" box="[1064,1169,1483,1508]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B97E5863FFA35423186F86FDFB503BA1" box="[1064,1169,1483,1508]" italics="true" pageId="30" pageNumber="31">T. emeryi</emphasis>
</taxonomicName>
standing hairs are very sparse to absent on these structures (MSC 0–6).
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA354231C808524FD5E38F8" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BB58471FFA354231C808524FD5E38F8" blockId="30.[151,1437,834,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231C808524FE2F3869" bold="true" box="[199,494,1554,1581]" pageId="30" pageNumber="31">Distribution and biology</emphasis>
. This species is known only from a single alate queen collected in Nature’s Valley,
<collectingRegion id="49CE4A93FFA354231CD08501FEF43814" box="[151,309,1591,1617]" country="South Africa" name="Western Cape" pageId="30" pageNumber="31">Western Cape</collectingRegion>
,
<collectingCountry id="F31DC4E1FFA354231D068501FE123814" box="[321,467,1591,1617]" name="South Africa" pageId="30" pageNumber="31">South Africa</collectingCountry>
. No workers have been found that could be attributed to this species, suggesting that it might be a workerless social parasite. Traits indicative of social parasitism include the small size, shiny integument, and increased amounts of standing pilosity compared to related congeners.
<taxonomicName id="4C0AFFF2FFA354231F998549FB8438DD" baseAuthorityName="Forel" baseAuthorityYear="1911" box="[990,1093,1663,1688]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B97E5863FFA354231F998549FB8438DD" box="[990,1093,1663,1688]" italics="true" pageId="30" pageNumber="31">T. emeryi</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C0AFFF2FFA3542319558549FA4038DD" authorityName="Ward" authorityYear="2022" box="[1298,1409,1663,1688]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="exactor">
<emphasis id="B97E5863FFA3542319558549FA4038DD" box="[1298,1409,1663,1688]" italics="true" pageId="30" pageNumber="31">T. exactor</emphasis>
</taxonomicName>
at Nature’s Valley and is a potential host species.
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA354231C8085F0FCDB38A4" box="[199,794,1734,1761]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BB58471FFA354231C8085F0FCDB38A4" blockId="30.[151,1437,834,1761]" box="[199,794,1734,1761]" pageId="30" pageNumber="31">
<emphasis id="B97E5863FFA354231C8085F0FE6838A5" bold="true" box="[199,425,1734,1760]" pageId="30" pageNumber="31">Material examined</emphasis>
. Known only from the
<typeStatus id="54B13AD3FFA354231EF785F1FCD738A4" box="[688,790,1735,1761]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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