treatments-xml/data/03/96/99/03969976961CFFB3FE9AF9EE794CAB12.xml
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<mods:title id="59751617FA61C3408E10377163FB9EB4">A new species of Neostethus (Teleostei; Atherinomorpha; Phallostethidae) from Brunei Darussalam, with comments on northwestern Borneo as an area of endemism</mods:title>
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<mods:namePart id="2251A216318AAAB978B07F056712DB77">Parenti, Lynne R.</mods:namePart>
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<paragraph id="8B802860961CFFB3FE9AF9EE7F64AFF2" blockId="2.[323,589,1662,1686]" box="[323,589,1662,1686]" pageId="2" pageNumber="177">
<heading id="D0C89F0C961CFFB3FE9AF9EE7F64AFF2" box="[323,589,1662,1686]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="177" reason="3">
<taxonomicName id="4C3F53E3961CFFB3FE9AF9EE7F64AFF2" ID-CoL="62RS" ID-ENA="300305" authority="Regan, 1916" authorityName="Regan" authorityYear="1916" box="[323,589,1662,1686]" class="Actinopterygii" family="Phallostethidae" genus="Neostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FE9AF9EE7C95AFF2" bold="true" box="[323,444,1662,1686]" italics="true" pageId="2" pageNumber="177">Neostethus</emphasis>
<bibRefCitation id="EFAE5591961CFFB3FE1CF9EF7F64AFF2" author="Regan CT" box="[453,589,1663,1686]" pageId="2" pageNumber="177" pagination="1 - 26" refId="ref12704" refString="Regan CT (1916) The morphology of the cyprinodont fishes of the subfamily Phallostethinae, with descriptions of a new genus and two new species. Proceedings of the Zoological Society of London, 1916: 1 - 26." type="journal article" year="1916">Regan, 1916</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3257BEB961CFFB3FF50F9517C89AFB2" box="[137,416,1729,1750]" pageId="2" pageNumber="177" type="reference_group">
<paragraph id="8B802860961CFFB3FF50F9517C89AFB2" blockId="2.[137,416,1729,1750]" box="[137,416,1729,1750]" pageId="2" pageNumber="177">
<taxonomicName id="4C3F53E3961CFFB3FF50F9517CA0AFB2" ID-CoL="62RS" authority="Regan, 1916: 1" authorityName="Regan" authorityPageNumber="1" authorityYear="1916" box="[137,393,1729,1750]" class="Actinopterygii" family="Phallostethidae" genus="Neostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FF50F9517DDBAFB2" box="[137,242,1729,1750]" italics="true" pageId="2" pageNumber="177">Neostethus</emphasis>
<bibRefCitation id="EFAE5591961CFFB3FF20F9517CA0AFB2" author="Regan CT" box="[249,393,1729,1750]" pageId="2" pageNumber="177" pagination="1 - 26" refId="ref12704" refString="Regan CT (1916) The morphology of the cyprinodont fishes of the subfamily Phallostethinae, with descriptions of a new genus and two new species. Proceedings of the Zoological Society of London, 1916: 1 - 26." type="journal article" year="1916">Regan, 1916:1</bibRefCitation>
</taxonomicName>
, 2
</paragraph>
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<subSubSection id="C3257BEB961CFFB3FF53F96F7C33AE32" pageId="2" pageNumber="177" type="type_taxon">
<paragraph id="8B802860961CFFB3FF53F96F7C33AE32" blockId="2.[137,776,1791,1878]" pageId="2" pageNumber="177">
<emphasis id="B94BF472961CFFB3FF53F96F7C34AE72" bold="true" box="[138,285,1791,1814]" pageId="2" pageNumber="177">
<typeStatus id="548496C2961CFFB3FF53F96F7DEBAE72" box="[138,194,1791,1814]" pageId="2" pageNumber="177">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3F53E3961CFFB3FEF8F96F7FA6AE72" ID-CoL="6RXLS" authority="Regan, 1916" authorityName="Regan" authorityYear="1916" box="[289,655,1791,1814]" class="Actinopterygii" family="Phallostethidae" genus="Neostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="species" species="lankesteri">
<emphasis id="B94BF472961CFFB3FEF8F96F7F2DAE72" box="[289,516,1791,1814]" italics="true" pageId="2" pageNumber="177">Neostethus lankesteri</emphasis>
<bibRefCitation id="EFAE5591961CFFB3FDD3F96F7FA6AE72" author="Regan CT" box="[522,655,1791,1814]" pageId="2" pageNumber="177" pagination="1 - 26" refId="ref12704" refString="Regan CT (1916) The morphology of the cyprinodont fishes of the subfamily Phallostethinae, with descriptions of a new genus and two new species. Proceedings of the Zoological Society of London, 1916: 1 - 26." type="journal article" year="1916">Regan, 1916</bibRefCitation>
</taxonomicName>
by original designation (use of “gen. et sp. n.” for one of two included new species).
</paragraph>
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<subSubSection id="C3257BEB961CFFB3FF50F8EF794CAB12" pageId="2" pageNumber="177" type="diagnosis">
<paragraph id="8B802860961CFFB3FF50F8EF79FBAB72" blockId="2.[137,776,1919,2070]" lastBlockId="2.[836,1475,191,534]" pageId="2" pageNumber="177">
<emphasis id="B94BF472961CFFB3FF50F8EF7CA3AEF2" bold="true" box="[137,394,1919,1942]" pageId="2" pageNumber="177">Differential diagnosis.</emphasis>
(modified from
<bibRefCitation id="EFAE5591961CFFB3FDE2F8EF7FD6AEF2" author="Parenti LR" box="[571,767,1919,1942]" pageId="2" pageNumber="177" pagination="243 - 277" refId="ref12409" refString="Parenti LR (1989) A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proceedings of the California Academy of Sciences, 46: 243 - 277." type="journal article" year="1989">Parenti, 1989:269</bibRefCitation>
)
<taxonomicName id="4C3F53E3961CFFB3FF50F80F7DD7AED2" authorityName="Regan" authorityYear="1916" box="[137,254,1951,1974]" class="Actinopterygii" family="Phallostethidae" genus="Neostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FF50F80F7DD7AED2" box="[137,254,1951,1974]" italics="true" pageId="2" pageNumber="177">Neostethus</emphasis>
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differs from all other phallostethines in having a priapium with an inner pulvinular bone (vs. lacking the bone), thin bony projections on the papillary bone that may number 80 or more (vs. lacking such bony projections) (
<bibRefCitation id="EFAE5591961CFFB3FC92FF2F7EC9A9B2" author="Parenti LR" box="[843,992,191,214]" pageId="2" pageNumber="177" pagination="243 - 277" refId="ref12409" refString="Parenti LR (1989) A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proceedings of the California Academy of Sciences, 46: 243 - 277." type="journal article" year="1989">Parenti, 1989</bibRefCitation>
: fig. 3) and two ctenactinia (vs. just a single short or rudimentary second (and no first) ctenactinium in
<taxonomicName id="4C3F53E3961CFFB3FC9DFF6F7EF0A872" authorityName="Regan" authorityYear="1913" box="[836,985,255,278]" class="Actinopterygii" family="Phallostethidae" genus="Phallostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FC9DFF6F7EF0A872" box="[836,985,255,278]" italics="true" pageId="2" pageNumber="177">Phallostethus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F53E3961CFFB3FBCFFF6F79EBA872" authorityName="Myers" authorityYear="1928" box="[1046,1218,255,278]" class="Actinopterygii" family="Phallostethidae" genus="Phenacostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FBCFFF6F79EBA872" box="[1046,1218,255,278]" italics="true" pageId="2" pageNumber="177">Phenacostethus</emphasis>
</taxonomicName>
and one elongate first (and no second) ctenactinium in
<taxonomicName id="4C3F53E3961CFFB3FB40FE8F7835A852" baseAuthorityName="Randall &amp; Lim" baseAuthorityYear="2000" box="[1177,1308,287,310]" class="Actinopterygii" family="Phallostethidae" genus="Gulaphallus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FB40FE8F7835A852" box="[1177,1308,287,310]" italics="true" pageId="2" pageNumber="177">Gulaphallus</emphasis>
</taxonomicName>
). Males lack an enlarged, fleshy seminal papilla and there is no translucent, membranous dome on the dorsal surface of the head as in
<taxonomicName id="4C3F53E3961CFFB3FC9DFEEF7EFCA8F2" authorityName="Regan" authorityYear="1913" box="[836,981,383,406]" class="Actinopterygii" family="Phallostethidae" genus="Phallostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FC9DFEEF7EFCA8F2" box="[836,981,383,406]" italics="true" pageId="2" pageNumber="177">Phallostethus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F53E3961CFFB3FBD1FEEF7986A8F2" authorityName="Myers" authorityYear="1928" box="[1032,1199,383,406]" class="Actinopterygii" family="Phallostethidae" genus="Phenacostethus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FBD1FEEF7986A8F2" box="[1032,1199,383,406]" italics="true" pageId="2" pageNumber="177">Phenacostethus</emphasis>
</taxonomicName>
. Males also do not have a perforated gular flap of skin through which the anterior end of the first ctenactinium projects or an aproctal axial bone that projects beyond the ventral body profile (
<bibRefCitation id="EFAE5591961CFFB3FAF2FE4F7897A892" author="Parenti LR" box="[1323,1470,479,502]" pageId="2" pageNumber="177" pagination="243 - 277" refId="ref12409" refString="Parenti LR (1989) A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proceedings of the California Academy of Sciences, 46: 243 - 277." type="journal article" year="1989">Parenti, 1989</bibRefCitation>
: fig. 4) which diagnoses
<taxonomicName id="4C3F53E3961CFFB3FB9EFE6F79E2AB72" baseAuthorityName="Randall &amp; Lim" baseAuthorityYear="2000" box="[1095,1227,511,534]" class="Actinopterygii" family="Phallostethidae" genus="Gulaphallus" kingdom="Animalia" order="Atheriniformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B94BF472961CFFB3FB9EFE6F79E2AB72" box="[1095,1227,511,534]" italics="true" pageId="2" pageNumber="177">Gulaphallus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B802860961CFFB3FC9DFDAF794CAB12" blockId="2.[836,1474,575,630]" pageId="2" pageNumber="177">
<emphasis id="B94BF472961CFFB3FC9DFDAF7EF7AB32" bold="true" box="[836,990,575,598]" pageId="2" pageNumber="177">Composition.</emphasis>
Twelve species, including the new species described herein (
<tableCitation id="C6BD1DDB961CFFB3FBDDFDCF797FAB12" box="[1028,1110,607,630]" captionStart="Table 1" captionStartId="3.[113,166,972,993]" captionText="Table 1. Classification of family Phallostethidae, following Parenti (1989) and Dyer &amp; Chernoff (1996). Species of subfamily Phallostethinae recorded from northwestern Borneo are marked with an asterisk (*)." pageId="2" pageNumber="177">Table 1</tableCitation>
).
</paragraph>
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