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<document id="E106BA6141F11D4258F43587DC38AE4D" ID-DOI="10.5281/zenodo.274748" ID-GBIF-Dataset="e687202a-a625-4aab-94de-312277c7e3fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="274748" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047074832" checkinUser="plazi" docAuthor="Aarvik, Leif &amp; Karisch, Timm" docDate="2009" docId="039687B6FFA7FFF1FF6E5921FB144B9F" docLanguage="en" docName="zt02026p032.pdf" docOrigin="Zootaxa 2026" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Multiquaestia agassizi Aarvik &amp; Karisch, new species" docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFAFFFCEFFA2FFF4FFF95853FFB14F5D" masterDocTitle="Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)" masterLastPageNumber="32" masterPageNumber="18" pageNumber="23" updateTime="1698577463540" updateUser="plazi">
<mods:mods id="34C2D3EBFA44D5AE4E81E7F4DE0EFFEE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D8A76D80AD880FDA805BFB2349E5BDAA">
<mods:title id="D2774C958171AECB75851E3D96CCFFC4">Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)</mods:title>
</mods:titleInfo>
<mods:name id="B3BA63E23D3CA696E64DFFABAFC30EB9" type="personal">
<mods:role id="03632CDCECF46C27B7F0614D75695967">
<mods:roleTerm id="AAC1ABD9136281154CD9A33F28F1F6B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0310A171427E087708F0054476B94E5">Aarvik, Leif</mods:namePart>
</mods:name>
<mods:name id="AD3832F374B1FE563D8F1DD765CD28B6" type="personal">
<mods:role id="B8E46C05F01D3DD61AA87E538F4AE728">
<mods:roleTerm id="F2AF71EF1F1A420A549BDD5B80D5147A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75443E14A26E3D1CE80C78A1DDEAFB90">Karisch, Timm</mods:namePart>
</mods:name>
<mods:typeOfResource id="3BCF3B8E53BD7169C710D0D623F1F114">text</mods:typeOfResource>
<mods:relatedItem id="6FC85C07FD926A7544DF7B12B2E39DD6" type="host">
<mods:titleInfo id="A85C2F5680EF108E597131D4BAFCEC49">
<mods:title id="DFE8DB081BCE9E0A1E1972C8409CDD2F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4722912CB955A94F3B609CE6B0B137B6">
<mods:date id="A8E9F014D1BBEB15FFAFB0A3B13D81B5">2009</mods:date>
<mods:detail id="C4DB86D8650D24A37D8C8A3EF2E1ECBA" type="volume">
<mods:number id="E21F94D4095B8DE15F7DAC5BC973ED16">2026</mods:number>
</mods:detail>
<mods:extent id="DF0CFC6CD9F57F8916D077803B090027" unit="page">
<mods:start id="2D9D4BD252C8A1D4A9DCF933060BBF7F">18</mods:start>
<mods:end id="7FCEB091C7FEC46BAD8253A8D82CBEB1">32</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="625EA51C3771A878E4F01D60915BD9D7">journal article</mods:classification>
<mods:identifier id="A79129B2A696721499FEFE1CF7108B29" type="DOI">10.5281/zenodo.274748</mods:identifier>
<mods:identifier id="4EE2643E1CEF2A8BFFA53D03AD879631" type="GBIF-Dataset">e687202a-a625-4aab-94de-312277c7e3fb</mods:identifier>
<mods:identifier id="89D395149029B919D75DE9DFA5C7D773" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="993A68AEC46FFC172168C9D370924F90" type="Zenodo-Dep">274748</mods:identifier>
</mods:mods>
<treatment id="039687B6FFA7FFF1FF6E5921FB144B9F" ID-DOI="http://doi.org/10.5281/zenodo.6216211" ID-GBIF-Taxon="119611110" ID-Zenodo-Dep="6216211" LSID="urn:lsid:plazi:treatment:039687B6FFA7FFF1FF6E5921FB144B9F" httpUri="http://treatment.plazi.org/id/039687B6FFA7FFF1FF6E5921FB144B9F" lastPageNumber="23" pageId="5" pageNumber="23">
<subSubSection id="C325652BFFA7FFF1FF6E5921FEE34EEF" pageId="5" pageNumber="23" type="nomenclature">
<paragraph id="8B8036A0FFA7FFF1FF6E5921FC9D4ED1" blockId="5.[151,812,370,434]" box="[151,812,370,396]" pageId="5" pageNumber="23">
<heading id="D0C881CCFFA7FFF1FF6E5921FC9D4ED1" bold="true" box="[151,812,370,396]" fontSize="11" level="1" pageId="5" pageNumber="23" reason="1">
<emphasis id="B94BEAB2FFA7FFF1FF6E5921FC9D4ED1" bold="true" box="[151,812,370,396]" pageId="5" pageNumber="23">
<taxonomicName id="4C3F4D23FFA7FFF1FF6E5921FD214ED1" ID-CoL="B3XJF" authority="Aarvik &amp; Karisch" authorityName="Aarvik &amp; Karisch" box="[151,656,370,396]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="agassizi" status="sp. nov.">
<emphasis id="B94BEAB2FFA7FFF1FF6E5921FE144ED1" bold="true" box="[151,421,370,396]" italics="true" pageId="5" pageNumber="23">Multiquaestia agassizi</emphasis>
Aarvik &amp; Karisch
</taxonomicName>
,
<taxonomicNameLabel id="A27857C9FFA7FFF1FD645921FC9D4ED1" box="[669,812,370,396]" pageId="5" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036A0FFA7FFF1FF6E59CBFEE34EEF" blockId="5.[151,812,370,434]" box="[151,338,408,434]" pageId="5" pageNumber="23">
(
<figureCitation id="13042A25FFA7FFF1FF6659CBFF434EEF" box="[159,242,408,434]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="5" pageNumber="23">Figs. 8</figureCitation>
,
<figureCitation id="13042A25FFA7FFF1FF0659CBFEFB4EEF" box="[255,330,408,434]" captionStart="FIGURES 18 – 22" captionStartId="8.[151,271,1510,1534]" captionTargetBox="[182,1404,197,1485]" captionTargetId="figure@8.[182,1404,197,1485]" captionTargetPageId="8" captionText="FIGURES 18 – 22. Male genitalia of Multiquaestia. 18. M. skulei n. sp. 19. M. skulei n. sp. uncus. 20. M. skulei n. sp. aedeagus. 21. M. agassizi n. sp. 22. M. agassizi n. sp. aedeagus." httpUri="https://zenodo.org/record/274752/files/figure.png" pageId="5" pageNumber="23">21, 22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA7FFF1FF6E59B1FDC64D7F" pageId="5" pageNumber="23" type="materials_examined">
<paragraph id="8B8036A0FFA7FFF1FF6E59B1FDC64D7F" blockId="5.[151,1437,482,1218]" pageId="5" pageNumber="23">
<emphasis id="B94BEAB2FFA7FFF1FF6E59B1FEFA4EA1" bold="true" box="[151,331,482,508]" pageId="5" pageNumber="23">
<typeStatus id="54848802FFA7FFF1FF6E59B1FF634EA1" box="[151,210,482,508]" pageId="5" pageNumber="23">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54848802FFA7FFF1FEAD59B1FE754EA1" box="[340,452,482,508]" pageId="5" pageNumber="23" type="holotype">Holotype</typeStatus>
, 3,
<collectingCountry id="F3287630FFA7FFF1FE0F59B1FDEC4EA1" box="[502,605,482,508]" name="Kenya" pageId="5" pageNumber="23">KENYA</collectingCountry>
, Western [Province], Kericho,
<quantity id="4CC79B45FFA7FFF1FC3159B1FB934EA1" box="[968,1058,482,508]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="5" pageNumber="23" unit="ft" value="7000.0">7000 ft</quantity>
.,
<date id="FF811060FFA7FFF1FBCB59B1FB194EA1" box="[1074,1192,482,508]" pageId="5" pageNumber="23" value="2000-05-24">24.v.2000</date>
, leg. D.J.L. Agassiz, genitalia slide 1981 T. Karisch (
<collectionCode id="ED2EAE65FFA7FFF1FDF45A5BFDDD4D7F" box="[525,620,520,546]" pageId="5" pageNumber="23">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA7FFF1FF3F5A7EFCF64B5A" pageId="5" pageNumber="23" type="description">
<paragraph id="8B8036A0FFA7FFF1FF3F5A7EFCF64B5A" blockId="5.[151,1437,482,1218]" pageId="5" pageNumber="23">
<emphasis id="B94BEAB2FFA7FFF1FF3F5A7EFEEE4D1A" bold="true" box="[198,351,557,583]" pageId="5" pageNumber="23">Description.</emphasis>
Adult (
<figureCitation id="13042A25FFA7FFF1FE445A7EFDB74D1A" box="[445,518,557,583]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="5" pageNumber="23">Fig. 8</figureCitation>
). Wingspan 13.0 mm.
<emphasis id="B94BEAB2FFA7FFF1FCE35A7EFCEB4D1B" box="[794,858,557,582]" italics="true" pageId="5" pageNumber="23">Head</emphasis>
: White, posteriorly and laterally greyish brown. Antenna light brown, scape white. Labial palpus 2 times diameter of eye, externally dark grey, internally and on dorsal edge white; due to scaling of second segment, broadly triangular in lateral view.
<emphasis id="B94BEAB2FFA7FFF1FB1A5A2BFA884DCC" box="[1251,1337,632,657]" italics="true" pageId="5" pageNumber="23">Thorax</emphasis>
: White; tegulae brownish ochreous. Legs brownish grey, tarsi with paler rings. Forewing upperside with basal 2/3 blackish brown; terminal 1/3 ochreous; on dorsum two white marks; the inner mark reaching to about 2/5 from dorsum, the distal one triangular, pointing towards apex, reaching to nearly middle of wing; a white dot present just beyond tip of distal white mark; a few dark scales present in terminal area.
<collectingCountry id="F3287630FFA7FFF1FB5D5B5EFB6E4C7A" box="[1188,1247,781,807]" name="India" pageId="5" pageNumber="23">Hind</collectingCountry>
wing upperside grey; pale ochreous hair-pencil going from base of hind margin to scale cover on dorsum of abdomen.
<emphasis id="B94BEAB2FFA7FFF1FF6E5B0BFEB64C2C" box="[151,263,856,881]" italics="true" pageId="5" pageNumber="23">Abdomen</emphasis>
: Male genitalia (
<figureCitation id="13042A25FFA7FFF1FE2C5B0BFD9D4C2F" box="[469,556,856,882]" captionStart="FIGURES 18 – 22" captionStartId="8.[151,271,1510,1534]" captionTargetBox="[182,1404,197,1485]" captionTargetId="figure@8.[182,1404,197,1485]" captionTargetPageId="8" captionText="FIGURES 18 – 22. Male genitalia of Multiquaestia. 18. M. skulei n. sp. 19. M. skulei n. sp. uncus. 20. M. skulei n. sp. aedeagus. 21. M. agassizi n. sp. 22. M. agassizi n. sp. aedeagus." httpUri="https://zenodo.org/record/274752/files/figure.png" pageId="5" pageNumber="23">Fig. 21</figureCitation>
) with tegumen subtriangular, uncus present in form of a distally rounded process, weakly sclerotised, socii atrophied; lower edge of sacculus gently curved, neck of valva broad, cucullus subovate, ventral point with short thorn-like process; aedeagus (
<figureCitation id="13042A25FFA7FFF1FBFC5BF1FBEC4CE1" box="[1029,1117,930,956]" captionStart="FIGURES 18 – 22" captionStartId="8.[151,271,1510,1534]" captionTargetBox="[182,1404,197,1485]" captionTargetId="figure@8.[182,1404,197,1485]" captionTargetPageId="8" captionText="FIGURES 18 – 22. Male genitalia of Multiquaestia. 18. M. skulei n. sp. 19. M. skulei n. sp. uncus. 20. M. skulei n. sp. aedeagus. 21. M. agassizi n. sp. 22. M. agassizi n. sp. aedeagus." httpUri="https://zenodo.org/record/274752/files/figure.png" pageId="5" pageNumber="23">Fig. 22</figureCitation>
) with broad, subtriangular base, constricted and angled 2/5 from base, widening distally; with 9 cornuti of which only two large spindleshaped ones had not been shed in the
<typeStatus id="54848802FFA7FFF1FDB65BBEFD084B5A" box="[591,697,1005,1031]" pageId="5" pageNumber="23" type="holotype">holotype</typeStatus>
; no carinae.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA7FFF1FF3F5C41FAD74B2A" pageId="5" pageNumber="23" type="diagnosis">
<paragraph id="8B8036A0FFA7FFF1FF3F5C41FAD74B2A" blockId="5.[151,1437,482,1218]" pageId="5" pageNumber="23">
<emphasis id="B94BEAB2FFA7FFF1FF3F5C41FEF74B71" bold="true" box="[198,326,1042,1068]" pageId="5" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="4C3F4D23FFA7FFF1FEB65C41FE644B76" box="[335,469,1042,1067]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B94BEAB2FFA7FFF1FEB65C41FE644B76" box="[335,469,1042,1067]" italics="true" pageId="5" pageNumber="23">M. agassizi</emphasis>
</taxonomicName>
differs from other
<taxonomicName id="4C3F4D23FFA7FFF1FD405C41FCED4B76" box="[697,860,1042,1067]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAB2FFA7FFF1FD405C41FCED4B76" box="[697,860,1042,1067]" italics="true" pageId="5" pageNumber="23">Multiquaestia</emphasis>
</taxonomicName>
species except
<taxonomicName id="4C3F4D23FFA7FFF1FBEF5C41FB174B76" box="[1046,1190,1042,1067]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="dallastai" status="sp. nov.">
<emphasis id="B94BEAB2FFA7FFF1FBEF5C41FB174B76" box="[1046,1190,1042,1067]" italics="true" pageId="5" pageNumber="23">M. dallastai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857C9FFA7FFF1FB575C41FB434B71" box="[1198,1266,1042,1068]" pageId="5" pageNumber="23" rank="species">
<emphasis id="B94BEAB2FFA7FFF1FB575C41FB434B71" bold="true" box="[1198,1266,1042,1068]" pageId="5" pageNumber="23">n. sp.</emphasis>
</taxonomicNameLabel>
by having the entire thorax, except for the tegulae, white.
<taxonomicName id="4C3F4D23FFA7FFF1FD6C5C6BFC924B0C" box="[661,803,1080,1105]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="dallastai">
<emphasis id="B94BEAB2FFA7FFF1FD6C5C6BFC924B0C" box="[661,803,1080,1105]" italics="true" pageId="5" pageNumber="23">M. dallastai</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C3F4D23FFA7FFF1FC445C6BFBF34B0C" box="[957,1090,1080,1105]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B94BEAB2FFA7FFF1FC445C6BFBF34B0C" box="[957,1090,1080,1105]" italics="true" pageId="5" pageNumber="23">M. agassizi</emphasis>
</taxonomicName>
by its larger size. In the male genitalia,
<taxonomicName id="4C3F4D23FFA7FFF1FEF35C0EFE214B2B" box="[266,400,1117,1142]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B94BEAB2FFA7FFF1FEF35C0EFE214B2B" box="[266,400,1117,1142]" italics="true" pageId="5" pageNumber="23">M. agassizi</emphasis>
</taxonomicName>
is distinct by the broad neck of the valva and the ventrodistal thorn on the cucullus.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA7FFF1FF3F5CD1FE734BC1" box="[198,450,1154,1180]" pageId="5" pageNumber="23" type="distribution">
<paragraph id="8B8036A0FFA7FFF1FF3F5CD1FE734BC1" blockId="5.[151,1437,482,1218]" box="[198,450,1154,1180]" pageId="5" pageNumber="23">
<emphasis id="B94BEAB2FFA7FFF1FF3F5CD1FED64BC1" bold="true" box="[198,359,1154,1180]" pageId="5" pageNumber="23">Distribution.</emphasis>
<collectingCountry id="F3287630FFA7FFF1FE945CD1FE0C4BC1" box="[365,445,1154,1180]" name="Kenya" pageId="5" pageNumber="23">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA7FFF1FF3F5CF4FB144B9F" box="[198,1189,1191,1218]" pageId="5" pageNumber="23" type="etymology">
<paragraph id="8B8036A0FFA7FFF1FF3F5CF4FB144B9F" blockId="5.[151,1437,482,1218]" box="[198,1189,1191,1218]" pageId="5" pageNumber="23">
<emphasis id="B94BEAB2FFA7FFF1FF3F5CF4FEE24B9C" bold="true" box="[198,339,1191,1217]" pageId="5" pageNumber="23">Etymology.</emphasis>
The species is named after the collector of the
<typeStatus id="54848802FFA7FFF1FC855CFBFC574B9F" box="[892,998,1192,1218]" pageId="5" pageNumber="23" type="holotype">holotype</typeStatus>
, David Agassiz.
</paragraph>
</subSubSection>
</treatment>
</document>
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