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<document id="C1C5C2E1BF851C5354B73FB939ADC6F0" ID-DOI="10.5281/zenodo.14662132" ID-ISSN="1942-1354" ID-Zenodo-Dep="14662132" ID-ZooBank="5671A77B-2ECB-445F-8F61-246A9E442CDE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736984354913" checkinUser="felipe" docAuthor="Eya, Bryan K." docDate="2024" docId="038487F8027FFF95FF140EB7FA49FA1C" docLanguage="en" docName="InsectaMundi.2024.1061.1-94.pdf" docOrigin="Insecta Mundi 2024 (61)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Mannophorus LeConte 1854" docType="key" docVersion="2" lastPageNumber="36" masterDocId="FFBDFF80025AFFB0FFD40F39FFE9FFB1" masterDocTitle="Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="36" updateTime="1736985555110" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="708A2DE9FA5F8F9498477456209CF1D6">Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)</mods:title>
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<mods:namePart id="A358B491CFD39200DC0CB8183C3B9A0E">Eya, Bryan K.</mods:namePart>
<mods:affiliation id="93071CF7E6CF11F1F6840D167E89C12B">California Academy of Sciences 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118 USA</mods:affiliation>
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<mods:date id="987E176A8CFD53AA7C24CFDDF6DF3E1A">2024</mods:date>
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<treatment id="038487F8027FFF95FF140EB7FA49FA1C" LSID="urn:lsid:plazi:treatment:038487F8027FFF95FF140EB7FA49FA1C" httpUri="http://treatment.plazi.org/id/038487F8027FFF95FF140EB7FA49FA1C" lastPageNumber="36" pageId="37" pageNumber="36">
<subSubSection id="C3376565027FFF95FF140EB7FCE0FE1B" box="[192,777,397,426]" pageId="37" pageNumber="36" type="nomenclature">
<paragraph id="8B9236EE027FFF95FF140EB7FCE0FE1B" blockId="37.[192,777,397,426]" box="[192,777,397,426]" pageId="37" pageNumber="36">
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<emphasis id="B959EAFC027FFF95FF140EB7FCE0FE1B" bold="true" box="[192,777,397,426]" pageId="37" pageNumber="36">
Key to species of
<taxonomicName id="4C2D4D6D027FFF95FE4E0EB4FCE0FE1B" ID-CoL="5KXF" authority="LeConte, 1854" authorityName="LeConte" authorityYear="1854" box="[410,777,397,426]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC027FFF95FE4E0EB4FDA2FE1B" bold="true" box="[410,587,397,426]" italics="true" pageId="37" pageNumber="36">Mannophorus</emphasis>
<bibRefCitation id="EFBC4B1F027FFF95FD850EB4FCE0FE1B" author="LeConte JL" box="[593,777,397,426]" pageId="37" pageNumber="36" pagination="439 - 448" refId="ref71415" refString="LeConte JL. 1854. Descriptions of some new Coleoptera from Texas, chiefly collected by the Mexican Boundary Commission. Proceedings of the Academy of Natural Sciences of Philadelphia 6: 439 - 448." type="journal article" year="1854">LeConte, 1854</bibRefCitation>
</taxonomicName>
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<paragraph id="8B9236EE027FFF95FF140E85FA49FE64" blockId="37.[192,1440,443,504]" box="[192,1440,443,470]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140E85FA49FE64" box="[192,1440,443,470]" pageId="37" pageNumber="36">
1. Each elytron with two ivory vittae, integument black or dark metallic bluish black or greenish......
<emphasis id="B959EAFC027FFF95FA460E82FA49FE64" bold="true" box="[1426,1440,443,469]" pageId="37" pageNumber="36">2</emphasis>
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</paragraph>
<paragraph id="8B9236EE027FFF95FF140EE7FA49FE46" blockId="37.[192,1440,443,504]" box="[192,1440,477,504]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140EE7FA49FE46" box="[192,1440,477,504]" pageId="37" pageNumber="36">
— Each elytron with three costae, integument yellowish-brown or metallic bluish with greenish luster..
<emphasis id="B959EAFC027FFF95FA460EE4FA49FE46" bold="true" box="[1426,1440,477,503]" pageId="37" pageNumber="36">5</emphasis>
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</keyStep>
<keyStep id="30D92E4B027FFF95FF140D33FA49FD39" pageId="37" pageNumber="36">
<paragraph id="8B9236EE027FFF95FF140D33FA49FDF4" blockId="37.[192,1440,521,649]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140D33FA49FDF4" pageId="37" pageNumber="36">
2(1). Pronotum with punctures contiguous, cribriform or alveolate, integument opaque, disc reddish, black or dark bluish with pubescence short (
<figureCitation id="13162A6B027FFF95FD0F0D12FC85FDF7" box="[731,876,555,582]" captionStart="Figures 152163" captionStartId="39.[193,272,1190,1214]" captionTargetBox="[194,1442,229,1161]" captionTargetId="figure-166@39.[194,1442,227,1161]" captionTargetPageId="39" captionText="Figures 152163. Mannophorus laetus LeConte, head, pronotum, thorax and elytral apices: 152155, 160, 162) M. laetus LeConte.156159, 161, 163) M. laetus (i.e., Sphenothecus cribellatus (Bates, 1892), syn. nov.). Head: 152, 156) Front short, mandibles arcuate at sides (blue arrow). 153, 157) Lateral profile with dorsal half of genae broad (red arrow) with lower eye lobes well separated from mandibles, apical margin of pronotum with collar-like projection laterally (yellow arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow). 154, 158) Frontolateral profile with front declivous in middle, and antennal tubercles divergent, each side of frontoclypeal sulcus (yellow arrow) with small, deep pit (red arrow), apical margin of pronotum elevated (green arrow), narrowly constricted behind. Pronotum: 155, 159) Apical margin with collarlike projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), disc with sides evenly rounded. Thorax lateral image: 160, 161) Pronotal disc convex, and mesosternum non-protuberant (red arrow). Elytra: 162, 163) Each elytron with two ivory vittae and apices rounded." figureDoi="http://doi.org/10.5281/zenodo.14662187" httpUri="https://zenodo.org/record/14662187/files/figure.png" pageId="37" pageNumber="36">Fig. 155, 159</figureCitation>
,
<figureCitation id="13162A6B027FFF95FCA30D12FC48FDF7" box="[887,929,555,582]" captionStart="Figures 178187" captionStartId="44.[191,270,956,980]" captionTargetBox="[193,1440,246,926]" captionTargetId="figure-282@44.[192,1440,229,926]" captionTargetPageId="44" captionText="Figures 178187. Mannophorus minor (Bates) comb. nov., dorsal and lateral images: 178179) Male, 13 mm, Puebla, MEX. 180181) Female, 13 mm, Puebla, MEX. Head: 182) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), antennal tubercles flattened (green arrows) then abruptly declivous in middle on either side of midcranial sulcus. 183) Lateral profile with dorsal half of genae very narrow (red arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow).184) Frontolateral profile with mandibles strongly angulated at sides (blue arrow). Pronotum: 185) Disc closely confluently punctate with apical margin elevated (green arrow) and narrowly constricted behind with collar-like projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), sides rounded slightly behind middle, tapered apically. Thorax lateral image: 186) Pronotal disc shallowly convex, and mesosternum nonprotuberant (red arrow), and submarginal costae starting from base of elytra (yellow arrow). Elytra: 187) Apices unarmed, rounded, vaguely serrate." figureDoi="http://doi.org/10.5281/zenodo.14662193" httpUri="https://zenodo.org/record/14662193/files/figure.png" pageId="37" pageNumber="36">185</figureCitation>
)....................................
<emphasis id="B959EAFC027FFF95FA460D12FA49FDF4" bold="true" box="[1426,1440,555,581]" pageId="37" pageNumber="36">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9236EE027FFF95FF140D74FA49FD39" blockId="37.[192,1440,521,649]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140D74FA49FD39" pageId="37" pageNumber="36">
— Pronotum with punctures coarse, irregular, integument shining, disc metallic greenish or bluish with long flying setae (
<figureCitation id="13162A6B027FFF95FDC70D57FCE7FD39" box="[531,782,622,649]" captionStart="Figures 202217" captionStartId="47.[192,271,1618,1642]" captionTargetBox="[195,1440,229,1579]" captionTargetId="figure-10@47.[192,1440,226,1588]" captionTargetPageId="47" captionText="Figures 202217. Mannophorus v. virgulata (Chemsak) comb. nov., and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology. Head: 202, 206) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), mandibles arcuate from base to apices (green arrow). 203, 207) Lateral profile with gena short, prominent (red arrow), posterior-lateral margin of eyes emarginate (green arrow). 204, 208) Frontolateral profile declivous along midcranial sulcus (yellow arrow). Pronotum: 205, 209, 212, 216) Apical margin elevated (yellow arrow), narrowly constricted behind with collar-like projection on sides (red arrow). 205, 212) Disc coarsely, subconfluently punctate (M. v. virgulata), clothed with long flying setae. 209, 216) Disc with punctures smaller, sparser, and well separated in M. v. virescens compared to M. v. virgulata. Thorax lateral images:210, 214) pronotum with collar-like projection on apical margin (yellow arrow), disc shallowly convex, densely clothed with long flying setae, punctures coarser in M. v. virgulata. mesosternum non-protuberant (red arrow). Elytra: 211, 213, 215, 217) Apices rounded or truncate, disc with punctures coarse, subconfluent." figureDoi="http://doi.org/10.5281/zenodo.14662197" httpUri="https://zenodo.org/record/14662197/files/figure.png" pageId="37" pageNumber="36">Fig. 205, 209, 212, 216</figureCitation>
)................................................
<emphasis id="B959EAFC027FFF95FA460D57FA49FD39" bold="true" box="[1426,1440,622,648]" pageId="37" pageNumber="36">4</emphasis>
</keyLead>
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</keyStep>
<keyStep id="30D92E4B027FFF95FF140DA2FA49FC52" pageId="37" pageNumber="36">
<paragraph id="8B9236EE027FFF95FF140DA2FA49FC8A" blockId="37.[192,1441,666,995]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140DA2FA49FC8A" pageId="37" pageNumber="36">
3(2). Form elongate; pronotum tapering apically, metallic dark bluish to aeneous black with an elongate, impunctate area in middle of basal half (
<figureCitation id="13162A6B027FFF95FCDC0D85FC8DFD66" box="[776,868,700,727]" captionStart="Figures 178187" captionStartId="44.[191,270,956,980]" captionTargetBox="[193,1440,246,926]" captionTargetId="figure-282@44.[192,1440,229,926]" captionTargetPageId="44" captionText="Figures 178187. Mannophorus minor (Bates) comb. nov., dorsal and lateral images: 178179) Male, 13 mm, Puebla, MEX. 180181) Female, 13 mm, Puebla, MEX. Head: 182) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), antennal tubercles flattened (green arrows) then abruptly declivous in middle on either side of midcranial sulcus. 183) Lateral profile with dorsal half of genae very narrow (red arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow).184) Frontolateral profile with mandibles strongly angulated at sides (blue arrow). Pronotum: 185) Disc closely confluently punctate with apical margin elevated (green arrow) and narrowly constricted behind with collar-like projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), sides rounded slightly behind middle, tapered apically. Thorax lateral image: 186) Pronotal disc shallowly convex, and mesosternum nonprotuberant (red arrow), and submarginal costae starting from base of elytra (yellow arrow). Elytra: 187) Apices unarmed, rounded, vaguely serrate." figureDoi="http://doi.org/10.5281/zenodo.14662193" httpUri="https://zenodo.org/record/14662193/files/figure.png" pageId="37" pageNumber="36">Fig. 185</figureCitation>
); antennal tubercles flattened, abruptly declivous on either side of mid-cranial sulcus (
<figureCitation id="13162A6B027FFF95FD130DE7FCB5FD49" box="[711,860,734,760]" captionStart="Figures 178187" captionStartId="44.[191,270,956,980]" captionTargetBox="[193,1440,246,926]" captionTargetId="figure-282@44.[192,1440,229,926]" captionTargetPageId="44" captionText="Figures 178187. Mannophorus minor (Bates) comb. nov., dorsal and lateral images: 178179) Male, 13 mm, Puebla, MEX. 180181) Female, 13 mm, Puebla, MEX. Head: 182) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), antennal tubercles flattened (green arrows) then abruptly declivous in middle on either side of midcranial sulcus. 183) Lateral profile with dorsal half of genae very narrow (red arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow).184) Frontolateral profile with mandibles strongly angulated at sides (blue arrow). Pronotum: 185) Disc closely confluently punctate with apical margin elevated (green arrow) and narrowly constricted behind with collar-like projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), sides rounded slightly behind middle, tapered apically. Thorax lateral image: 186) Pronotal disc shallowly convex, and mesosternum nonprotuberant (red arrow), and submarginal costae starting from base of elytra (yellow arrow). Elytra: 187) Apices unarmed, rounded, vaguely serrate." figureDoi="http://doi.org/10.5281/zenodo.14662193" httpUri="https://zenodo.org/record/14662193/files/figure.png" pageId="37" pageNumber="36">Fig. 182, 184</figureCitation>
); dorsal half of genae narrow with anterior margins of lower eyes almost contiguous with base of mandibles (
<figureCitation id="13162A6B027FFF95FC080DC6FBD2FCA8" box="[988,1083,767,793]" captionStart="Figures 178187" captionStartId="44.[191,270,956,980]" captionTargetBox="[193,1440,246,926]" captionTargetId="figure-282@44.[192,1440,229,926]" captionTargetPageId="44" captionText="Figures 178187. Mannophorus minor (Bates) comb. nov., dorsal and lateral images: 178179) Male, 13 mm, Puebla, MEX. 180181) Female, 13 mm, Puebla, MEX. Head: 182) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), antennal tubercles flattened (green arrows) then abruptly declivous in middle on either side of midcranial sulcus. 183) Lateral profile with dorsal half of genae very narrow (red arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow).184) Frontolateral profile with mandibles strongly angulated at sides (blue arrow). Pronotum: 185) Disc closely confluently punctate with apical margin elevated (green arrow) and narrowly constricted behind with collar-like projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), sides rounded slightly behind middle, tapered apically. Thorax lateral image: 186) Pronotal disc shallowly convex, and mesosternum nonprotuberant (red arrow), and submarginal costae starting from base of elytra (yellow arrow). Elytra: 187) Apices unarmed, rounded, vaguely serrate." figureDoi="http://doi.org/10.5281/zenodo.14662193" httpUri="https://zenodo.org/record/14662193/files/figure.png" pageId="37" pageNumber="36">Fig. 183</figureCitation>
); mandibles with sides strongly angulated (
<figureCitation id="13162A6B027FFF95FE1A0C18FDC3FC8A" box="[462,554,801,827]" captionStart="Figures 178187" captionStartId="44.[191,270,956,980]" captionTargetBox="[193,1440,246,926]" captionTargetId="figure-282@44.[192,1440,229,926]" captionTargetPageId="44" captionText="Figures 178187. Mannophorus minor (Bates) comb. nov., dorsal and lateral images: 178179) Male, 13 mm, Puebla, MEX. 180181) Female, 13 mm, Puebla, MEX. Head: 182) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), antennal tubercles flattened (green arrows) then abruptly declivous in middle on either side of midcranial sulcus. 183) Lateral profile with dorsal half of genae very narrow (red arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow).184) Frontolateral profile with mandibles strongly angulated at sides (blue arrow). Pronotum: 185) Disc closely confluently punctate with apical margin elevated (green arrow) and narrowly constricted behind with collar-like projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), sides rounded slightly behind middle, tapered apically. Thorax lateral image: 186) Pronotal disc shallowly convex, and mesosternum nonprotuberant (red arrow), and submarginal costae starting from base of elytra (yellow arrow). Elytra: 187) Apices unarmed, rounded, vaguely serrate." figureDoi="http://doi.org/10.5281/zenodo.14662193" httpUri="https://zenodo.org/record/14662193/files/figure.png" pageId="37" pageNumber="36">Fig. 184</figureCitation>
).........................................
<emphasis id="B959EAFC027FFF95FB840C18FA49FC8A" bold="true" box="[1104,1440,801,827]" pageId="37" pageNumber="36">
<taxonomicName id="4C2D4D6D027FFF95FB840C18FAFEFC8A" ID-CoL="6MXT5" authority="(Bates)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Bates" box="[1104,1303,801,827]" class="Insecta" family="Cerambycidae" genus="Ischnocnemis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="minor" status="comb. nov.">
<emphasis id="B959EAFC027FFF95FB840C18FB55FC8A" bold="true" box="[1104,1212,801,827]" italics="true" pageId="37" pageNumber="36">M. minor</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787027FFF95FAF60C18FA49FC8A" box="[1314,1440,801,827]" pageId="37" pageNumber="36" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8B9236EE027FFF95FF140C7AFA49FC52" blockId="37.[192,1441,666,995]" pageId="37" pageNumber="36">
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— Form stout, pronotum evenly rounded on sides, disc reddish with apical and basal margins black, all black or all dark bluish, punctures coarse and contiguous throughout; antennal tubercles gradually declivous from apex to mid-cranial sulcus; dorsal half of genae broader with margins of lower eye lobes well separated from base of mandibles; sides of mandibles arcuate (
<figureCitation id="13162A6B027FFF95FBAF0C9EFAE7FC73" box="[1147,1294,935,962]" captionStart="Figures 152163" captionStartId="39.[193,272,1190,1214]" captionTargetBox="[194,1442,229,1161]" captionTargetId="figure-166@39.[194,1442,227,1161]" captionTargetPageId="39" captionText="Figures 152163. Mannophorus laetus LeConte, head, pronotum, thorax and elytral apices: 152155, 160, 162) M. laetus LeConte.156159, 161, 163) M. laetus (i.e., Sphenothecus cribellatus (Bates, 1892), syn. nov.). Head: 152, 156) Front short, mandibles arcuate at sides (blue arrow). 153, 157) Lateral profile with dorsal half of genae broad (red arrow) with lower eye lobes well separated from mandibles, apical margin of pronotum with collar-like projection laterally (yellow arrow), posterior-lateral margin of lower eye lobe emarginate (green arrow). 154, 158) Frontolateral profile with front declivous in middle, and antennal tubercles divergent, each side of frontoclypeal sulcus (yellow arrow) with small, deep pit (red arrow), apical margin of pronotum elevated (green arrow), narrowly constricted behind. Pronotum: 155, 159) Apical margin with collarlike projection on sides (red arrow), base margined and more broadly constricted on sides (blue arrow), disc with sides evenly rounded. Thorax lateral image: 160, 161) Pronotal disc convex, and mesosternum non-protuberant (red arrow). Elytra: 162, 163) Each elytron with two ivory vittae and apices rounded." figureDoi="http://doi.org/10.5281/zenodo.14662187" httpUri="https://zenodo.org/record/14662187/files/figure.png" pageId="37" pageNumber="36">Fig. 152159</figureCitation>
)...............................................................................
<taxonomicName id="4C2D4D6D027FFF95FB180CF0FA49FC52" ID-CoL="6R6J5" authority="LeConte" authorityName="LeConte. Lateral" authorityYear="1854" box="[1228,1440,968,995]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B959EAFC027FFF95FB180CF0FA49FC52" bold="true" box="[1228,1440,968,995]" pageId="37" pageNumber="36">
<emphasis id="B959EAFC027FFF95FB180CF0FADFFC52" bold="true" box="[1228,1334,968,995]" italics="true" pageId="37" pageNumber="36">M. laetus</emphasis>
LeConte
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D92E4B027FFF95FF140CCCFA49FB4A" pageId="37" pageNumber="36">
<paragraph id="8B9236EE027FFF95FF140CCCFA49FBC5" blockId="37.[192,1440,1013,1275]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140CCCFA49FBC5" pageId="37" pageNumber="36">
4(2). Antennae stout, male with antennomeres IV, IX and X subequal to or slightly longer than I (
<figureCitation id="13162A6B027FFF95FAE00CCCFA7DFBBE" box="[1332,1428,1013,1039]" captionStart="Figures 218221" captionStartId="48.[192,271,1115,1139]" captionTargetBox="[225,1405,229,1083]" captionTargetId="figure-290@48.[221,1410,229,1096]" captionTargetPageId="48" captionText="Figures 218221. Mannophorus v. virgulata (Chemsak) comb. nov. and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology of antennae and dorsal habitus: 218) Mannophorus v. virgulata male with antennae stout, antennomeres IV, IX and X subequal to or slightly longer than I, subsutural raised ivory vitta narrow from base to near apex. 219) Mannophorus v. virescens male with antennae slender, antennomeres IV, IX and X much longer than I, subsutural raised ivory vitta broader anteriorly. 220) Mannophorus v. virgulata female with antennae short, stout, distal antennomeres that are expanded, antennomeres IVVII shorter than I, subsutural raised ivory vitta narrow from base to near apex. 221) Mannophorus v. virescens female with antennae slender, elongate, antennomere IV subequal to I and VVII slightly longer than I, subsutural raised ivory vitta broader anteriorly." figureDoi="http://doi.org/10.5281/zenodo.14662199" httpUri="https://zenodo.org/record/14662199/files/figure.png" pageId="37" pageNumber="36">Fig. 218</figureCitation>
), female with distal antennomeres expanded, antennomeres IVVII shorter than I (
<figureCitation id="13162A6B027FFF95FB310B2EFAADFB80" box="[1253,1348,1047,1073]" captionStart="Figures 218221" captionStartId="48.[192,271,1115,1139]" captionTargetBox="[225,1405,229,1083]" captionTargetId="figure-290@48.[221,1410,229,1096]" captionTargetPageId="48" captionText="Figures 218221. Mannophorus v. virgulata (Chemsak) comb. nov. and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology of antennae and dorsal habitus: 218) Mannophorus v. virgulata male with antennae stout, antennomeres IV, IX and X subequal to or slightly longer than I, subsutural raised ivory vitta narrow from base to near apex. 219) Mannophorus v. virescens male with antennae slender, antennomeres IV, IX and X much longer than I, subsutural raised ivory vitta broader anteriorly. 220) Mannophorus v. virgulata female with antennae short, stout, distal antennomeres that are expanded, antennomeres IVVII shorter than I, subsutural raised ivory vitta narrow from base to near apex. 221) Mannophorus v. virescens female with antennae slender, elongate, antennomere IV subequal to I and VVII slightly longer than I, subsutural raised ivory vitta broader anteriorly." figureDoi="http://doi.org/10.5281/zenodo.14662199" httpUri="https://zenodo.org/record/14662199/files/figure.png" pageId="37" pageNumber="36">Fig. 220</figureCitation>
); subsutural raised ivory vittae narrow from base to near apex; pronotum coarsely, subconfluently punctate (
<figureCitation id="13162A6B027FFF95FE8C0B63FE00FBC5" box="[344,489,1114,1141]" captionStart="Figures 202217" captionStartId="47.[192,271,1618,1642]" captionTargetBox="[195,1440,229,1579]" captionTargetId="figure-10@47.[192,1440,226,1588]" captionTargetPageId="47" captionText="Figures 202217. Mannophorus v. virgulata (Chemsak) comb. nov., and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology. Head: 202, 206) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), mandibles arcuate from base to apices (green arrow). 203, 207) Lateral profile with gena short, prominent (red arrow), posterior-lateral margin of eyes emarginate (green arrow). 204, 208) Frontolateral profile declivous along midcranial sulcus (yellow arrow). Pronotum: 205, 209, 212, 216) Apical margin elevated (yellow arrow), narrowly constricted behind with collar-like projection on sides (red arrow). 205, 212) Disc coarsely, subconfluently punctate (M. v. virgulata), clothed with long flying setae. 209, 216) Disc with punctures smaller, sparser, and well separated in M. v. virescens compared to M. v. virgulata. Thorax lateral images:210, 214) pronotum with collar-like projection on apical margin (yellow arrow), disc shallowly convex, densely clothed with long flying setae, punctures coarser in M. v. virgulata. mesosternum non-protuberant (red arrow). Elytra: 211, 213, 215, 217) Apices rounded or truncate, disc with punctures coarse, subconfluent." figureDoi="http://doi.org/10.5281/zenodo.14662197" httpUri="https://zenodo.org/record/14662197/files/figure.png" pageId="37" pageNumber="36">Fig. 205, 212</figureCitation>
)...............................
<emphasis id="B959EAFC027FFF95FC5A0B63FA49FBC5" bold="true" box="[910,1440,1113,1140]" pageId="37" pageNumber="36">
<taxonomicName id="4C2D4D6D027FFF95FC5A0B63FAFEFBC5" authority="(Chemsak)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Chemsak" baseAuthorityYear="2024" box="[910,1303,1113,1140]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="virgulata" status="comb. nov." subSpecies="virgulata">
<emphasis id="B959EAFC027FFF95FC5A0B63FB67FBC5" bold="true" box="[910,1166,1113,1140]" italics="true" pageId="37" pageNumber="36">M. virgulata virgulata</emphasis>
(Chemsak)
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787027FFF95FAF60B63FA49FBC5" box="[1314,1440,1114,1140]" pageId="37" pageNumber="36" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9236EE027FFF95FF140B45FA49FB4A" blockId="37.[192,1440,1013,1275]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140B45FA49FB4A" pageId="37" pageNumber="36">
— Antennae slender, male with antennomeres IV, IX and X much longer than I (
<figureCitation id="13162A6B027FFF95FB700B45FAECFB27" box="[1188,1285,1148,1174]" captionStart="Figures 218221" captionStartId="48.[192,271,1115,1139]" captionTargetBox="[225,1405,229,1083]" captionTargetId="figure-290@48.[221,1410,229,1096]" captionTargetPageId="48" captionText="Figures 218221. Mannophorus v. virgulata (Chemsak) comb. nov. and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology of antennae and dorsal habitus: 218) Mannophorus v. virgulata male with antennae stout, antennomeres IV, IX and X subequal to or slightly longer than I, subsutural raised ivory vitta narrow from base to near apex. 219) Mannophorus v. virescens male with antennae slender, antennomeres IV, IX and X much longer than I, subsutural raised ivory vitta broader anteriorly. 220) Mannophorus v. virgulata female with antennae short, stout, distal antennomeres that are expanded, antennomeres IVVII shorter than I, subsutural raised ivory vitta narrow from base to near apex. 221) Mannophorus v. virescens female with antennae slender, elongate, antennomere IV subequal to I and VVII slightly longer than I, subsutural raised ivory vitta broader anteriorly." figureDoi="http://doi.org/10.5281/zenodo.14662199" httpUri="https://zenodo.org/record/14662199/files/figure.png" pageId="37" pageNumber="36">Fig. 219</figureCitation>
), female with antennomere IV subequal to I and VVII slightly longer than I (
<figureCitation id="13162A6B027FFF95FBFE0BA4FB63FB06" box="[1066,1162,1181,1207]" captionStart="Figures 218221" captionStartId="48.[192,271,1115,1139]" captionTargetBox="[225,1405,229,1083]" captionTargetId="figure-290@48.[221,1410,229,1096]" captionTargetPageId="48" captionText="Figures 218221. Mannophorus v. virgulata (Chemsak) comb. nov. and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology of antennae and dorsal habitus: 218) Mannophorus v. virgulata male with antennae stout, antennomeres IV, IX and X subequal to or slightly longer than I, subsutural raised ivory vitta narrow from base to near apex. 219) Mannophorus v. virescens male with antennae slender, antennomeres IV, IX and X much longer than I, subsutural raised ivory vitta broader anteriorly. 220) Mannophorus v. virgulata female with antennae short, stout, distal antennomeres that are expanded, antennomeres IVVII shorter than I, subsutural raised ivory vitta narrow from base to near apex. 221) Mannophorus v. virescens female with antennae slender, elongate, antennomere IV subequal to I and VVII slightly longer than I, subsutural raised ivory vitta broader anteriorly." figureDoi="http://doi.org/10.5281/zenodo.14662199" httpUri="https://zenodo.org/record/14662199/files/figure.png" pageId="37" pageNumber="36">Fig. 221</figureCitation>
); subsutural raised ivory vittae broader anteriorly; pronotum moderately, coarsely punctate (
<figureCitation id="13162A6B027FFF95FBEB0B86FB39FB68" box="[1087,1232,1215,1241]" captionStart="Figures 202217" captionStartId="47.[192,271,1618,1642]" captionTargetBox="[195,1440,229,1579]" captionTargetId="figure-10@47.[192,1440,226,1588]" captionTargetPageId="47" captionText="Figures 202217. Mannophorus v. virgulata (Chemsak) comb. nov., and Mannophorus v. virescens (Eya) comb. nov., syn. nov., comparative morphology. Head: 202, 206) Front with small, deep pits (red arrow) on each side of frontoclypeal sulcus (yellow arrow), mandibles arcuate from base to apices (green arrow). 203, 207) Lateral profile with gena short, prominent (red arrow), posterior-lateral margin of eyes emarginate (green arrow). 204, 208) Frontolateral profile declivous along midcranial sulcus (yellow arrow). Pronotum: 205, 209, 212, 216) Apical margin elevated (yellow arrow), narrowly constricted behind with collar-like projection on sides (red arrow). 205, 212) Disc coarsely, subconfluently punctate (M. v. virgulata), clothed with long flying setae. 209, 216) Disc with punctures smaller, sparser, and well separated in M. v. virescens compared to M. v. virgulata. Thorax lateral images:210, 214) pronotum with collar-like projection on apical margin (yellow arrow), disc shallowly convex, densely clothed with long flying setae, punctures coarser in M. v. virgulata. mesosternum non-protuberant (red arrow). Elytra: 211, 213, 215, 217) Apices rounded or truncate, disc with punctures coarse, subconfluent." figureDoi="http://doi.org/10.5281/zenodo.14662197" httpUri="https://zenodo.org/record/14662197/files/figure.png" pageId="37" pageNumber="36">Fig. 209, 216</figureCitation>
).......................................................
<emphasis id="B959EAFC027FFF95FCB50BD8FA49FB4A" bold="true" box="[865,1440,1248,1275]" pageId="37" pageNumber="36">
<taxonomicName id="4C2D4D6D027FFF95FCB50BD8FB4DFB4B" authority="(Eya)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Eya" baseAuthorityYear="2010" box="[865,1188,1248,1275]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="virgulata" status="comb. nov., syn. nov." subSpecies="virescens">
<emphasis id="B959EAFC027FFF95FCB50BD8FBB7FB4B" bold="true" box="[865,1118,1248,1275]" italics="true" pageId="37" pageNumber="36">M. virgulata virescens</emphasis>
(Eya)
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787027FFF95FB7B0BD8FA49FB4A" box="[1199,1440,1249,1275]" pageId="37" pageNumber="36">comb. nov., syn. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D92E4B027FFF95FF140A35FA49FA1C" pageId="37" pageNumber="36">
<paragraph id="8B9236EE027FFF95FF140A35FA49FAF9" blockId="37.[192,1440,1292,1454]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140A35FA49FAF9" pageId="37" pageNumber="36">
5(1). Integument yellowish-brown; pronotum with seven black maculae, and apical third to half of elytra black (
<figureCitation id="13162A6B027FFF95FE830A17FE03FAF9" box="[343,490,1326,1352]" captionStart="Figures 164177" captionStartId="42.[192,271,1216,1240]" captionTargetBox="[193,1440,229,1194]" captionTargetId="figure-163@42.[192,1440,216,1195]" captionTargetPageId="42" captionText="Figures 164177. Mannophorus forreri Bates, dorsal and lateral images: 164165) Male, 17 mm, Pima Co., AZ, USA. 166 167) Female, 20 mm, Chihuahua, MEX. 168169) Female, 22 mm, Chihuahua, MEX. 170) Male, lectotype, Durango, MEX (Bezark 2024, id: 17269). Head: 171) Front short, mandibles arcuate at sides (blue arrow). 172) Lateral profile with genae short (red arrow), lower eye lobes well separated from mandibles, posterior-lateral margin of lower eye emarginate (green arrow), collar-like projection on apical margin of pronotum (yellow arrow).173) Frontolateral profile with front declivous in middle, and antennal tubercles divergent, each side of frontoclypeal sulcus (yellow arrow) with small, deep pit (red arrow). Pronotum: 174) Disc with sides evenly rounded, apical margin elevated (green arrow) with collar-like projection laterally (red arrow), base margined and broadly constricted on sides (blue arrow). Thorax lateral image: 175) Disc convex, and mesosternum non-protuberant (red arrow). Elytra: 176, 177) Apices unarmed, rounded to truncate with coarse, intricate punctures, each elytron with three narrow raised costae, subsutural impunctate costa not attaining apex (red arrows), submarginal costa (green arrow), and third narrow, partially punctate costae (blue arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662189" httpUri="https://zenodo.org/record/14662189/files/figure.png" pageId="37" pageNumber="36">Fig. 164177</figureCitation>
)..........................................................
<taxonomicName id="4C2D4D6D027FFF95FB3E0A16FA49FAF9" ID-CoL="3XVPT" authority="Bates" authorityName="Bates" authorityYear="1885" box="[1258,1440,1326,1353]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="forreri">
<emphasis id="B959EAFC027FFF95FB3E0A16FA49FAF9" bold="true" box="[1258,1440,1326,1353]" pageId="37" pageNumber="36">
<emphasis id="B959EAFC027FFF95FB3E0A16FAB2FAF9" bold="true" box="[1258,1371,1326,1353]" italics="true" pageId="37" pageNumber="36">M. forreri</emphasis>
Bates
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9236EE027FFF95FF140A69FA49FA1C" blockId="37.[192,1440,1292,1454]" pageId="37" pageNumber="36">
<keyLead id="30DC95DB027FFF95FF140A69FA49FA1C" pageId="37" pageNumber="36">
— Integument metallic bluish with greenish luster; sides of pronotum broadly tuberculate; front, vertex, pronotum, prosternum, and basal half of elytra clothed with long flying erect setae (
<figureCitation id="13162A6B027FFF95FB270A48FA6EFA3D" box="[1267,1415,1393,1420]" captionStart="Figures 222234" captionStartId="51.[192,271,1251,1275]" captionTargetBox="[193,1439,232,1215]" captionTargetId="figure-193@51.[192,1440,226,1221]" captionTargetPageId="51" captionText="Figures 222234. Mannophorus tricostatus Eya sp. nov., dorsal and lateral images: 222223) Male, Chiapas, 48 km SE Jct. 190/199, MEX. 224225) Female, Chiapas, 48 km SE Jct. 190/199, MEX. Head: 226) Front. 227) Lateral profile. 228) Frontolateral profile. Pronotum: 229) Disc, sides broadly tuberculate. Thorax lateral image: 230) mesosternal intercoxal process non-protuberant. Elytra: 231) Male, apices. 232) Female, apices. 233) Elytron lateral image with three costae, subsutural costa (red arrow), submarginal costa (yellow arrow), costa in between (green arrow). Metatarsi: 234) Metatarsomere 1 (T1) subequal to or shorter than tarsomeres 2+3 (T2+3), T1/T2+3: 0.9." figureDoi="http://doi.org/10.5281/zenodo.14662201" httpUri="https://zenodo.org/record/14662201/files/figure.png" pageId="37" pageNumber="36">Fig. 222234</figureCitation>
)..............................................................
<emphasis id="B959EAFC027FFF95FBB90AAAFA49FA1C" bold="true" box="[1133,1440,1427,1453]" pageId="37" pageNumber="36">
<taxonomicName id="4C2D4D6D027FFF95FBB90AAAFAD7FA1C" authority="Eya" authorityName="Eya" authorityYear="2024" box="[1133,1342,1427,1453]" class="Insecta" family="Cerambycidae" genus="Mannophorus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
<emphasis id="B959EAFC027FFF95FBB90AAAFAE5FA1C" bold="true" box="[1133,1292,1427,1453]" italics="true" pageId="37" pageNumber="36">M. tricostatus</emphasis>
Eya
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787027FFF95FA9D0AAAFA49FA1C" box="[1353,1440,1427,1453]" pageId="37" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>