290 lines
41 KiB
XML
290 lines
41 KiB
XML
<document id="64F4852EB476612C2FC74C4672D2EFBB" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862729D06EECCFCD2FE55EC2D" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus brandiae Borkent 2012, new species" docType="treatment" docUuid="0B421129-5E09-42CA-94F5-9E98BF18FCF0" docUuidSource="ZooBank" docVersion="4" lastPageNumber="21" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="20" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="F52E20F4E396579255DF6F5C6EB2E49B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4BFEC2CB591D14B6AD9968C22C164B76">
|
||
<mods:title id="5AA9CB2B5E957E333CC075D8E9E57BB0">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="EDBEF9A678672184FE5C4F8895187962" type="personal">
|
||
<mods:role id="FC317E7EEC7AA128C46B3A98615416B4">
|
||
<mods:roleTerm id="9C07FF399AE494335AFCF6144331EEE3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="950E5CF47D3F39D1A3DB40A5107AE39B">Borkent, Christopher J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="517D61735A8742975D44280C97CB699C" type="personal">
|
||
<mods:role id="5B7456315E9199A9692BD807F65CDDE0">
|
||
<mods:roleTerm id="ADB9EEF24D7B8815B6202862CFDB50A4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DD25DF9BFCAEC4E15DECBBF6D1ABB01D">Wheeler, Terry A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="87A4EB4F944B04BA29481CD417E600B6">text</mods:typeOfResource>
|
||
<mods:relatedItem id="84C0391ADD1C64386B70E3EE9FF25D18" type="host">
|
||
<mods:titleInfo id="0B07409C8865981E1363A4DCF19BCC68">
|
||
<mods:title id="5CD3AF8AF030271FE40808B28CDF9964">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="964A3EFB47EA1100FFA13CF67806B96A">
|
||
<mods:date id="E58D8DC3966F2F1D56329E227E9AFE7A">2012</mods:date>
|
||
<mods:detail id="CD6EBB631BD84AB311C44F544F5543C3" type="pubDate">
|
||
<mods:number id="1234AD4DF6B85D76CEBE4305AC34805B">2012-11-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="97339E80F580B14549A112F64D2AB016" type="volume">
|
||
<mods:number id="C349D7B8824F9C1A2EEFE7ADEB2ACEB9">3549</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="F97D34BE805A6571E5129A1F268CB5E8" unit="page">
|
||
<mods:start id="525F6992C3B37AD47B7D2C80A0FA8A9B">1</mods:start>
|
||
<mods:end id="09DDD0738522794A655EB133B0668AEC">117</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="04A7E809D54AB9BF499FB96C85CA7990">journal article</mods:classification>
|
||
<mods:identifier id="67608F309B56FEBCFD136EF97F989854" type="ISSN">11755334</mods:identifier>
|
||
<mods:identifier id="79337A2A1CE699FEAEFC8F38A52C1F5E" type="ZooBank">2412CB4F-4D29-4988-80C1-205D16767678</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CA3487C862729D06EECCFCD2FE55EC2D" ID-DOI="http://doi.org/10.5281/zenodo.5258327" ID-Zenodo-Dep="5258327" LSID="urn:lsid:zoobank.org:act:0B421129-5E09-42CA-94F5-9E98BF18FCF0" httpUri="http://treatment.plazi.org/id/CA3487C862729D06EECCFCD2FE55EC2D" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
<subSubSection id="0A87655562729D01EECCFCD2FD3AE95C" box="[151,739,835,862]" pageId="19" pageNumber="20" type="nomenclature">
|
||
<paragraph id="422236DE62729D01EECCFCD2FD3AE95C" blockId="19.[151,739,835,894]" box="[151,739,835,862]" pageId="19" pageNumber="20">
|
||
<heading id="196A81B262729D01EECCFCD2FD3AE95C" bold="true" box="[151,739,835,862]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
|
||
<emphasis id="70E9EACC62729D01EECCFCD2FD3AE95C" bold="true" box="[151,739,835,862]" pageId="19" pageNumber="20">
|
||
5.
|
||
<taxonomicName id="859D4D5D62729D01EEEFFCD5FD9FE95C" ID-CoL="8CJ44" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[180,582,835,862]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brandiae" status="sp. nov.">
|
||
<emphasis id="70E9EACC62729D01EEEFFCD5FE0FE95F" bold="true" box="[180,470,836,862]" italics="true" pageId="19" pageNumber="20">Leptomorphus brandiae</emphasis>
|
||
Borkent
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6BDA57B762729D01EC08FCD2FD3AE95C" box="[595,739,835,861]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562729D01EECCFCF7FDA9E97F" box="[151,624,870,894]" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="422236DE62729D01EECCFCF7FDA9E97F" blockId="19.[151,739,835,894]" box="[151,624,870,894]" pageId="19" pageNumber="20">
|
||
(
|
||
<figureCitation id="DAA62A5B62729D01EEC5FCF7FED2E97F" box="[158,267,870,894]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="19" pageNumber="20">Figures 6</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EF4CFCF7FEEFE97F" box="[279,310,870,894]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="19" pageNumber="20">46</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EF1AFCF7FE86E97F" box="[321,351,870,894]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="19" pageNumber="20">82</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EF30FCF7FE50E97F" box="[363,393,870,894]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="19" pageNumber="20">95</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EFCEFCF7FE18E97F" box="[405,449,870,894]" captionStart="FIGURES 125–133" captionStartId="106.[151,263,1891,1914]" captionTargetBox="[180,1416,183,1873]" captionTargetId="figure-98@106.[180,1416,183,1873]" captionTargetPageId="106" captionText="FIGURES 125–133. Leptomorphus immatures and host fungi. 125. Fomitopsis pinicola, host fungus to larvae of L. nebulosus. 126. Cerrena unicolor, host fungus to L. hyalinus. 127–131. Leptomorphus larvae. 127. L. brandiae. 128. L. nebulosus in situ on F. pinicola. 129. L. nebulosus (photograph: A. Borkent). 130. L. subcaeruleus. 131. Fully mature larva of L. nebulosus beginning to pupate. 132–133. Leptomorphus pupae. 132. L. hyalinus. 133. L. nebulosus. Arrow indicates final larval skin left behind on anchor line (photograph: A. Borkent)." pageId="19" pageNumber="20">127</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EF96FCF7FE20E97F" box="[461,505,870,894]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="19" pageNumber="20">143</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EC5EFCF7FDE8E97F" box="[517,561,870,894]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="19" pageNumber="20">150</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62729D01EC66FCF7FDB1E97F" box="[573,616,870,894]" captionStart="FIGURE 155" captionStartId="116.[151,250,858,881]" captionTargetBox="[151,1425,187,838]" captionTargetId="figure-58@116.[151,1436,159,885]" captionTargetPageId="116" captionText="FIGURE 155. One of three equally parsimonious trees showing the relationships between species in the L. furcatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="19" pageNumber="20">155</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562729D01EECCFC3CFA5AEE7A" pageId="19" pageNumber="20" type="diagnosis">
|
||
<paragraph id="422236DE62729D01EECCFC3CFAFAEE30" blockId="19.[151,1437,941,2005]" pageId="19" pageNumber="20">
|
||
<emphasis id="70E9EACC62729D01EECCFC3CFEEDE9C7" bold="true" box="[151,308,941,966]" pageId="19" pageNumber="20">DIAGNOSIS</emphasis>
|
||
: The only extant species of
|
||
<taxonomicName id="859D4D5D62729D01EC35FC3FFCC9E9C7" authorityName=": Edwards" authorityYear="1925" box="[622,784,942,966]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E9EACC62729D01EC35FC3FFCC9E9C7" box="[622,784,942,966]" italics="true" pageId="19" pageNumber="20">Leptomorphus</emphasis>
|
||
</taxonomicName>
|
||
with the following combination of characters: scutum and scutellum evenly covered with fine trichia; male foretibia with a dense row (comb) of short anteroventral bristles; palp segment 5 with thick covering of fine, white, setulae; male genitalia with tergite 9 rounded basally, tapering to a single, long, point posteriorly that extends beyond cerci, gonocoxite without gonocoxal lobe (
|
||
<figureCitation id="DAA62A5B62729D01EA99FB88FACFEE30" box="[1218,1302,1049,1073]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="19" pageNumber="20">Fig. 95</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="422236DE62729D01EE9CFBACFA5AEE7A" blockId="19.[151,1437,941,2005]" pageId="19" pageNumber="20">
|
||
This species can be distinguished from other Neotropical species without R
|
||
<subScript id="DE19349B62729D01EDA7FBDDFBDDEE5B" attach="left" box="[1020,1028,1100,1114]" fontSize="6" pageId="19" pageNumber="20">4</subScript>
|
||
by the distinctive pointed tergite 9 of the large male genitalia (
|
||
<figureCitation id="DAA62A5B62729D01EFFFFBF2FE2FEE7A" box="[420,502,1123,1147]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="19" pageNumber="20">Fig. 95</figureCitation>
|
||
). It also has the darkest legs of any of the species (e.g. hind femur 2/3 dark,
|
||
<figureCitation id="DAA62A5B62729D01EB6FFBF2FAAFEE7A" box="[1332,1398,1123,1147]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="19" pageNumber="20">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562729D06EE9CFB17FC84EE5D" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="422236DE62729D06EE9CFB17FB70E9CC" blockId="19.[151,1437,941,2005]" lastBlockId="20.[151,1437,151,1580]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
<emphasis id="70E9EACC62729D01EE9CFB17FE50EE9E" bold="true" box="[199,393,1158,1183]" pageId="19" pageNumber="20">DESCRIPTION</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62729D01EFC1FB16FE0AEE9E" box="[410,467,1159,1183]" italics="true" pageId="19" pageNumber="20">Male</emphasis>
|
||
. (
|
||
<figureCitation id="DAA62A5B62729D01EFB0FB16FDE9EE9E" box="[491,560,1159,1183]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="19" pageNumber="20">Fig. 6</figureCitation>
|
||
)
|
||
<emphasis id="70E9EACC62729D01EC1AFB16FD58EE9E" bold="true" box="[577,641,1159,1183]" pageId="19" pageNumber="20">Head</emphasis>
|
||
: light brown-brown, somewhat dorsoventrally compressed in anterior view. Antenna dark brown; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 1 large bristle, several setae on apicodorsal margin, patch of fine setae apicoventrally; flagellomere 1 with tapered base yellow, remainder dark brown; flagellomere 6 1.1X as long as broad. Clypeus light brown, slightly laterally compressed oval; bristles on clypeus brown, 6–8 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face light brown; shape a slightly wider than tall triangle, with few bristles ventrolaterally. Frons brown; bare, frontal furrow running 1/10–1/4 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus with segments 1–4 yellow, segment 5 white (
|
||
<figureCitation id="DAA62A5B62729D01ECA8FA59FC9EEFE1" box="[755,839,1480,1504]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="19" pageNumber="20">Fig. 82</figureCitation>
|
||
); segment 1 small but visible below eye, segments increasing in length, segment 5 2X length and 1.5X width of segment 4 with even width from base to apex and covered in fine white setulae, segment 3 with very small lateral patch of fine setae encircled by dark setae. Labellum light brown. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput brown with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black.
|
||
<emphasis id="70E9EACC62729D01EA21F9EDFB0BEC95" bold="true" box="[1146,1234,1660,1684]" pageId="19" pageNumber="20">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62729D01EB63F9EAFA42EC92" box="[1336,1435,1659,1683]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="19" pageNumber="20" unit="mm" value="1.31">1.31 mm</quantity>
|
||
(n = 1). Dark brown with yellow lateral spots. Scutum dark brown/black with blue-green specks, pair of small yellow anterolateral spots; surface of scutum covered with trichia; acrostichal setae absent; single dorsocentral seta present anteriorly; two thirds row of lateral setae present; patch of setae on scutum at wing base small. Scutellum dark brown with covering of fine trichia; bristles absent. Prescutum yellow. Mediotergite dark brown with 6 bristles on posterolateral corners, and covering of appressed trichia. Laterotergite brown, with covering of appressed trichia; anterior margin of laterotergite abutting katepisternum. Anepimeron brown with posterodorsal corner yellow. Anepisternum dark brown. Katepisternum brown with dorsal third yellow. Antepronotum and proepisternum brown. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum brown. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown.
|
||
<emphasis id="70E9EACC62759D06EFC1FF09FE0BEAB1" bold="true" box="[410,466,152,176]" pageId="20" pageNumber="21">Legs</emphasis>
|
||
: principally dark brown; coxae white except apical margin; fore- and midfemur with apical 1/3–1/2 yellow, hind femur with yellow on third quarter; extreme anteroapical corner brown on forefemur, on other femora dark brown; fore- and midtibia with basal 1/2 yellow. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia with comb of short setae along length of anteroventral surface, hind tibial spur dark brown, remainder brown, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 3/5 of its length, placed basally, shortest midtibial spur 0.84X length of longest, longest midtibial spur 4X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 3.75X apical thickness of hind tibia. Foreleg first tarsomere 1.8X length of foretibia.
|
||
<emphasis id="70E9EACC62759D06EF5EFE24FE9FEBCF" bold="true" box="[261,326,437,462]" pageId="20" pageNumber="21">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62759D06EF0DFE27FE75EBCF" box="[342,428,438,462]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="20" pageNumber="21">Fig. 46</figureCitation>
|
||
): Length
|
||
<quantity id="85659B3B62759D06EC42FE27FD58EBCC" box="[537,641,438,462]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" pageId="20" pageNumber="21" unit="mm" value="5.25">5.25 mm</quantity>
|
||
. Hyaline; apical macula dark brown but fading apically, running from anterior to posterior wing margin, beginning halfway along R
|
||
<subScript id="DE19349B62759D06ED1DFE79FC97EBF7" attach="left" box="[838,846,488,502]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
|
||
; medial macula extending from Sc to posterior wing margin (fainter on posterior third). Macrotrichia in all cells, though absent from posterobasal margin of cell a and sparse in basal cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present, apical end joining R within its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62759D06EB19FDC7FA93E865" attach="left" box="[1346,1354,598,612]" fontSize="6" pageId="20" pageNumber="21">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62759D06EEF2FDECFF68E88A" attach="left" box="[169,177,637,651]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
|
||
slightly concave for entire length. M
|
||
<subScript id="DE19349B62759D06EC0BFDECFD81E88A" attach="left" box="[592,600,637,651]" fontSize="6" pageId="20" pageNumber="21">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B62759D06ED39FDECFCB3E88A" attach="left" box="[866,874,637,651]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
|
||
, apices of M veins thinning towards wing margin. M
|
||
<subScript id="DE19349B62759D06EEEBFD35FF61E8B3" attach="left" box="[176,184,676,690]" fontSize="6" pageId="20" pageNumber="21">4</subScript>
|
||
-CuA fork arising at the same level as r-m. A
|
||
<subScript id="DE19349B62759D06ECEFFD35FD65E8B3" attach="left" box="[692,700,676,690]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62759D06ED41FD05FC57E8AC" bold="true" box="[794,910,660,685]" pageId="20" pageNumber="21">Abdomen</emphasis>
|
||
: Tergites and sternites principally dark brown, T3–5 with anterolateral yellow spots. Tergite 8 smaller than all other abdominal sclerites, without bristles.
|
||
<emphasis id="70E9EACC62759D06EECCFD4FFEDFE8F6" bold="true" box="[151,262,734,759]" pageId="20" pageNumber="21">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62759D06EF4DFD4EFEB5E8F6" box="[278,364,735,759]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="20" pageNumber="21">Fig. 95</figureCitation>
|
||
): basal half brown remainder light yellow/white. Sternite 9 very small, sclerotized, posteriorly directed Y-shape, not reaching to gonocoxal margin. Tergite 9 longer than wide, rounded basally, tapering to a single, long point posteriorly that extends beyond cerci. Gonocoxite placed 1/4 way toward apex of T9, medial margin not reaching medial line, bearing gonostylus on apical 1/3. Gonostylus a single broad-based lobe tapering to a point apically with small setae basally, gonocoxite III fused to dorsolateral margin. Aedeagus equal in length to gonocoxite, tapering to middle and then slightly bulbous apically, apodemes 1/3 total length. Parameres a slightly curved taper, 1.2X length of apodemes, apodemes strongly united with gonocoxal apodemes.
|
||
</paragraph>
|
||
<paragraph id="422236DE62759D06EE9CFC48FB17EE15" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<emphasis id="70E9EACC62759D06EE9CFC48FEC2E9F0" box="[199,283,985,1009]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
|
||
. As for male, except as follows.
|
||
<emphasis id="70E9EACC62759D06ECC1FC48FD2BE9F0" bold="true" box="[666,754,985,1009]" pageId="20" pageNumber="21">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62759D06ED07FC48FC18E9F0" box="[860,961,985,1009]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="20" pageNumber="21" unit="mm" value="1.56">1.56 mm</quantity>
|
||
(n = 1).
|
||
<emphasis id="70E9EACC62759D06EA70FC49FBB2E9F0" bold="true" box="[1067,1131,984,1009]" pageId="20" pageNumber="21">Wing</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62759D06EA8EFC48FAE2E9F0" box="[1237,1339,985,1009]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" pageId="20" pageNumber="21" unit="mm" value="6.15">6.15 mm</quantity>
|
||
(n = 1).
|
||
<emphasis id="70E9EACC62759D06EECCFC6CFF16EE14" bold="true" box="[151,207,1021,1045]" pageId="20" pageNumber="21">Legs</emphasis>
|
||
: Foretibia without comb of short setae on anteroventral surface.
|
||
<emphasis id="70E9EACC62759D06EDFFFC6DFBC0EE14" bold="true" box="[932,1049,1020,1045]" pageId="20" pageNumber="21">Abdomen</emphasis>
|
||
: Cercus yellow.
|
||
</paragraph>
|
||
<paragraph id="422236DE62759D06EE9CFBB0FC84EE5D" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<emphasis id="70E9EACC62759D06EE9CFBB0FEE7EE39" box="[199,318,1057,1080]" italics="true" pageId="20" pageNumber="21">Immatures</emphasis>
|
||
. White/ivory background colour with small, dark, spots covering the dorsal and lateral surfaces. Follows the general habitus for other known species (
|
||
<figureCitation id="DAA62A5B62759D06ECB5FBD5FC89EE5D" box="[750,848,1092,1116]" captionStart="FIGURES 125–133" captionStartId="106.[151,263,1891,1914]" captionTargetBox="[180,1416,183,1873]" captionTargetId="figure-98@106.[180,1416,183,1873]" captionTargetPageId="106" captionText="FIGURES 125–133. Leptomorphus immatures and host fungi. 125. Fomitopsis pinicola, host fungus to larvae of L. nebulosus. 126. Cerrena unicolor, host fungus to L. hyalinus. 127–131. Leptomorphus larvae. 127. L. brandiae. 128. L. nebulosus in situ on F. pinicola. 129. L. nebulosus (photograph: A. Borkent). 130. L. subcaeruleus. 131. Fully mature larva of L. nebulosus beginning to pupate. 132–133. Leptomorphus pupae. 132. L. hyalinus. 133. L. nebulosus. Arrow indicates final larval skin left behind on anchor line (photograph: A. Borkent)." pageId="20" pageNumber="21">Fig. 127</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562759D06EE9CFBF6FB53EEC6" pageId="20" pageNumber="21" type="biology_ecology">
|
||
<paragraph id="422236DE62759D06EE9CFBF6FB53EEC6" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<emphasis id="70E9EACC62759D06EE9CFBF6FE93EE81" bold="true" box="[199,330,1127,1152]" pageId="20" pageNumber="21">BIOLOGY</emphasis>
|
||
: Larvae and pupae were collected from an encrusting fungus on the underside of wet discarded lumber near La Fortuna,
|
||
<collectingCountry id="3A8A764E62759D06EFE9FB1AFDE8EEA2" box="[434,561,1163,1187]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
|
||
. Larvae had spun sheets and were feeding on the spores that accumulated, as described for other species (
|
||
<figureCitation id="DAA62A5B62759D06EF8AFB3EFDEDEEC6" box="[465,564,1199,1223]" captionStart="FIGURES 125–133" captionStartId="106.[151,263,1891,1914]" captionTargetBox="[180,1416,183,1873]" captionTargetId="figure-98@106.[180,1416,183,1873]" captionTargetPageId="106" captionText="FIGURES 125–133. Leptomorphus immatures and host fungi. 125. Fomitopsis pinicola, host fungus to larvae of L. nebulosus. 126. Cerrena unicolor, host fungus to L. hyalinus. 127–131. Leptomorphus larvae. 127. L. brandiae. 128. L. nebulosus in situ on F. pinicola. 129. L. nebulosus (photograph: A. Borkent). 130. L. subcaeruleus. 131. Fully mature larva of L. nebulosus beginning to pupate. 132–133. Leptomorphus pupae. 132. L. hyalinus. 133. L. nebulosus. Arrow indicates final larval skin left behind on anchor line (photograph: A. Borkent)." pageId="20" pageNumber="21">Fig. 127</figureCitation>
|
||
). The pupae were suspended from their head and tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562759D06EE9CFB43FC90EEEA" box="[199,841,1234,1259]" pageId="20" pageNumber="21" type="distribution">
|
||
<paragraph id="422236DE62759D06EE9CFB43FC90EEEA" blockId="20.[151,1437,151,1580]" box="[199,841,1234,1259]" pageId="20" pageNumber="21">
|
||
<emphasis id="70E9EACC62759D06EE9CFB43FE4FEEEA" bold="true" box="[199,406,1234,1259]" pageId="20" pageNumber="21">DISTRIBUTION</emphasis>
|
||
:
|
||
<collectingCountry id="3A8A764E62759D06EFFFFB42FDC7EEEA" box="[420,542,1235,1259]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B62759D06EC75FB42FD56EEEA" box="[558,655,1235,1259]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="20" pageNumber="21">Fig. 143</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62759D06ECFFFB42FC9DEEEA" box="[676,836,1235,1259]" metricMagnitude="2" metricUnit="m" metricValue="4.3" metricValueMax="5.6" metricValueMin="3.0" pageId="20" pageNumber="21" unit="m" value="430.0" valueMax="560.0" valueMin="300.0">300–560 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562759D06EE9CFB67FC90EF57" pageId="20" pageNumber="21" type="etymology">
|
||
<paragraph id="422236DE62759D06EE9CFB67FC90EF57" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<emphasis id="70E9EACC62759D06EE9CFB67FE58EF0E" bold="true" box="[199,385,1270,1295]" pageId="20" pageNumber="21">ETYMOLOGY</emphasis>
|
||
: This species is named for my wife Brandi Borkent, as it is a particularly beautiful species, and in thanks for her support and encouragement in all matters relating to life during my doctoral program as well as for her help while I was collecting the
|
||
<typeStatus id="9D26887C62759D06EC3EFAAEFD1CEF56" box="[613,709,1343,1367]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
|
||
specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562759D06EE9CFAF0FE55EC2D" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="422236DE62759D06EE9CFAF0FCA3EFE4" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="F2F53C8362759D06EE9CFAF0FCA3EFE4" collectingDate="2003-03-14" collectingDateMax="2003-03-25" collectingDateMin="2003-03-14" collectionCode="MATERIAL, R, LEM" collectorName="E. Rojas & B. Gamboa & W. Arana & Tp. Interseccion & Det. C. J. Borkent" country="Costa Rica" determinerName="Det. C. J. Borkent" elevation="560" latitude="9.66" location="Hitoy Cerere" longLatPrecision="779" longitude="-83.03" pageId="20" pageNumber="21" specimenCount="561" specimenCount-male="1" stateProvince="Limon" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62759D06EE9CFAF0FDDDEF7B" bold="true" box="[199,516,1377,1402]" pageId="20" pageNumber="21">
|
||
<collectionCode id="248CAE1B62759D06EE9CFAF0FE86EF7B" box="[199,351,1377,1402]" pageId="20" pageNumber="21">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62759D06EC4DFAF3FD5FEF7B" box="[534,646,1378,1402]" italics="true" pageId="20" pageNumber="21">
|
||
<typeStatus id="9D26887C62759D06EC4DFAF3FD59EF7B" box="[534,640,1378,1402]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
here designated, adult male, pinned with genitalia in plastic vial on pin, labelled “
|
||
<collectingCountry id="3A8A764E62759D06EF1AFA17FE2AEF9F" box="[321,499,1414,1438]" name="Costa Rica" pageId="20" pageNumber="21">COSTA RICA</collectingCountry>
|
||
: Prov.
|
||
<collectingRegion id="8059F83C62759D06EC15FA17FD45EF9F" box="[590,668,1414,1438]" country="Costa Rica" name="Limon" pageId="20" pageNumber="21">Limon</collectingRegion>
|
||
.
|
||
<collectionCode id="248CAE1B62759D06ECF5FA17FD1CEF9C" box="[686,709,1414,1437]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
|
||
.B./
|
||
<location id="4742600562759D06ECACFA17FC4AEF9F" LSID="urn:lsid:plazi:treatment:CA3487C862729D06EECCFCD2FE55EC2D:4742600562759D06ECACFA17FC4AEF9F" box="[759,915,1414,1438]" country="Costa Rica" latitude="9.66" longLatPrecision="779" longitude="-83.03" name="Hitoy Cerere" pageId="20" pageNumber="21" stateProvince="Limon">Hitoy Cerere</location>
|
||
, Send. Espavel.
|
||
<specimenCount id="549BFD5762759D06EA3AFA17FB71EF9F" box="[1121,1192,1414,1438]" pageId="20" pageNumber="21" type="generic">
|
||
<quantity id="85659B3B62759D06EA3AFA17FB7DEF9F" box="[1121,1188,1414,1438]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="21" unit="m" value="560.0">
|
||
<elevation id="C9B0D1ED62759D06EA3AFA17FB7DEF9F" box="[1121,1188,1414,1438]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="21" unit="m" value="560.0">560m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
/ (
|
||
<geoCoordinate id="27A9501962759D06EA9CFA17FAC2EF9F" box="[1223,1307,1414,1438]" degrees="9.66" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="9.66">9.66°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="27A9501962759D06EB75FA17FA4DEF9F" box="[1326,1428,1414,1438]" degrees="83.03" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="-83.03">83.03°W</geoCoordinate>
|
||
)
|
||
<date id="3623101E62759D06EECCFA38FEE2EFC0" box="[151,315,1449,1473]" pageId="20" pageNumber="21" value="2003-03-14" valueMax="2003-03-25" valueMin="2003-03-14">
|
||
<collectingDate id="2667E9F662759D06EECCFA38FEE2EFC0" box="[151,315,1449,1473]" pageId="20" pageNumber="21" value="2003-03-14" valueMax="2003-03-25" valueMin="2003-03-14">14–25.iii.2003</collectingDate>
|
||
</date>
|
||
./
|
||
<collectorName id="EF68530862759D06EF14FA3BFE68EFC0" box="[335,433,1449,1473]" pageId="20" pageNumber="21">E. Rojas</collectorName>
|
||
,
|
||
<collectorName id="EF68530862759D06EFE7FA3BFDE7EFC0" box="[444,574,1449,1473]" pageId="20" pageNumber="21">B. Gamboa</collectorName>
|
||
,
|
||
<collectorName id="EF68530862759D06EC12FA3BFD6BEFC0" box="[585,690,1449,1473]" pageId="20" pageNumber="21">W. Arana</collectorName>
|
||
/
|
||
<collectorName id="EF68530862759D06EC9AFA3BFCA0EFC0" box="[705,889,1449,1473]" pageId="20" pageNumber="21">Tp. Interseccion</collectorName>
|
||
, sample #73474;
|
||
<typeStatus id="9D26887C62759D06EA66FA38FB09EFC0" box="[1085,1232,1449,1473]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD5762759D06EA8DFA3BFB32EFC0" box="[1238,1259,1450,1473]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="859D4D5D62759D06EAA1FA38FF25EFE4" authorityName="Borkent" authorityYear="2012" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brandiae">
|
||
<emphasis id="70E9EACC62759D06EAA1FA38FF25EFE4" italics="true" pageId="20" pageNumber="21">Leptomorphus brandiae</emphasis>
|
||
</taxonomicName>
|
||
/Borkent,
|
||
<taxonomicNameLabel id="6BDA57B762759D06EF28FA5FFE23EFE4" box="[371,506,1485,1509]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="B7FEDA1A62759D06EC54FA5FFD0BEFE4" box="[527,722,1485,1509]" pageId="20" pageNumber="21">
|
||
<collectorName id="EF68530862759D06EC54FA5FFD0BEFE4" box="[527,722,1485,1509]" pageId="20" pageNumber="21">Det. C.J. Borkent</collectorName>
|
||
</determinerName>
|
||
2012” [
|
||
<collectionCode id="248CAE1B62759D06ED77FA5FFCB6EFE4" box="[812,879,1486,1509]" pageId="20" pageNumber="21">LEM</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="422236DE62759D06EE9CFA63FE55EC2D" blockId="20.[151,1437,151,1580]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="F2F53C8362759D06EE9CFA63FE5FEC2D" collectingDate="2010-08-15" collectionCode="LEM" collectorName="C. J. Borkent" country="Costa Rica" location="Alajeula" pageId="20" pageNumber="21" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
|
||
<emphasis id="70E9EACC62759D06EE9CFA63FE98EC08" box="[199,321,1522,1545]" italics="true" pageId="20" pageNumber="21">
|
||
<typeStatus id="9D26887C62759D06EE9CFA63FEE5EC08" box="[199,316,1522,1545]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="3A8A764E62759D06EF10FA60FE2CEC08" box="[331,501,1521,1545]" name="Costa Rica" pageId="20" pageNumber="21">COSTA RICA</collectingCountry>
|
||
:
|
||
<location id="4742600562759D06EC59FA60FDBAEC08" LSID="urn:lsid:plazi:treatment:CA3487C862729D06EECCFCD2FE55EC2D:4742600562759D06EC59FA60FDBAEC08" box="[514,611,1521,1545]" country="Costa Rica" name="Alajeula" pageId="20" pageNumber="21">Alajeula</location>
|
||
,
|
||
<locationDeviation id="53902FFC62759D06EC2AFA60FCA5EC08" box="[625,892,1521,1545]" location="La Fortuna" pageId="20" pageNumber="21">
|
||
<quantity id="85659B3B62759D06EC2AFA60FD61EC08" box="[625,696,1521,1545]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="20" pageNumber="21" unit="km" value="4.3">4.3km</quantity>
|
||
SSE La Fortuna
|
||
</locationDeviation>
|
||
,
|
||
<date id="3623101E62759D06EDD1FA60FBCDEC08" box="[906,1044,1521,1545]" pageId="20" pageNumber="21" value="2010-08-15">
|
||
<collectingDate id="2667E9F662759D06EDD1FA60FBCDEC08" box="[906,1044,1521,1545]" pageId="20" pageNumber="21" value="2010-08-15">15.viii.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EF68530862759D06EA79FA60FB68EC08" box="[1058,1201,1521,1545]" pageId="20" pageNumber="21">C.J. Borkent</collectorName>
|
||
[with cast off pupal skin] (
|
||
<specimenCount id="549BFD5762759D06EE85F985FEDBEC2D" box="[222,258,1556,1580]" pageId="20" pageNumber="21" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount id="549BFD5762759D06EF4BF985FEEAEC2D" box="[272,307,1556,1580]" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62759D06EF1AF985FEA7EC2D" box="[321,382,1556,1580]" pageId="20" pageNumber="21">LEM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |