294 lines
38 KiB
XML
294 lines
38 KiB
XML
<document id="CC4A07E8F43B764248076CA2A121F011" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C8626E9D02EECCFA1EFC6FEDA3" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus babai Sasakawa" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="16" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="70CFF6F14CA1C16F9D7D6573FA3DFDB3">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<mods:namePart id="B9B76B98E303C48C7D6A2146BB3068D7">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="895F6C7E583CDA47FAC39A4EC0E6CB4A">Wheeler, Terry A.</mods:namePart>
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<mods:date id="7C443756BF541FAA2B30458E2FFFFFC8">2012</mods:date>
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<mods:number id="82EC94C2DFB726705320C9744D735AB9">2012-11-15</mods:number>
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<mods:number id="A57A45E8BDFEB695FF0959AA7B50ACB4">3549</mods:number>
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<treatment id="CA3487C8626E9D02EECCFA1EFC6FEDA3" ID-DOI="http://doi.org/10.5281/zenodo.5258325" ID-Zenodo-Dep="5258325" LSID="urn:lsid:plazi:treatment:CA3487C8626E9D02EECCFA1EFC6FEDA3" httpUri="http://treatment.plazi.org/id/CA3487C8626E9D02EECCFA1EFC6FEDA3" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection id="0A876555626E9D1DEECCFA1EFDE9EFA8" box="[151,560,1423,1449]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="422236DE626E9D1DEECCFA1EFDE9EFA8" blockId="15.[151,560,1423,1481]" box="[151,560,1423,1449]" pageId="15" pageNumber="16">
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<heading id="196A81B2626E9D1DEECCFA1EFDE9EFA8" bold="true" box="[151,560,1423,1449]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="70E9EACC626E9D1DEECCFA1EFDE9EFA8" bold="true" box="[151,560,1423,1449]" pageId="15" pageNumber="16">
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3.
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<taxonomicName id="859D4D5D626E9D1DEEEFFA1EFDE9EFA8" ID-CoL="8CJ3Q" authority="Sasakawa" authorityName="Sasakawa" box="[180,560,1423,1449]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="babai">
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<emphasis id="70E9EACC626E9D1DEEEFFA1EFE77EFA8" bold="true" box="[180,430,1423,1449]" italics="true" pageId="15" pageNumber="16">Leptomorphus babai</emphasis>
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Sasakawa
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||
</taxonomicName>
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||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A876555626E9D1DEECCFA20FDD7EFC8" box="[151,526,1457,1481]" pageId="15" pageNumber="16" type="description">
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<paragraph id="422236DE626E9D1DEECCFA20FDD7EFC8" blockId="15.[151,560,1423,1481]" box="[151,526,1457,1481]" pageId="15" pageNumber="16">
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(
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||
<figureCitation id="DAA62A5B626E9D1DEEC5FA20FED2EFC8" box="[158,267,1457,1481]" captionStart="FIGURES 1–4" captionStartId="83.[151,263,1566,1589]" captionTargetBox="[153,1436,289,1543]" captionTargetId="figure-45@83.[151,1436,287,1544]" captionTargetPageId="83" captionText="FIGURES 1–4. Leptomorphus habitus. 1. L. aliciae, male. 2. L. babai, paratype male. 3. L. amorimi, paratype female. 4. L. bifasciatus, female." pageId="15" pageNumber="16">Figures 2</figureCitation>
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,
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<figureCitation id="DAA62A5B626E9D1DEF4CFA20FEEFEFC8" box="[279,310,1457,1481]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="15" pageNumber="16">44</figureCitation>
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,
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<figureCitation id="DAA62A5B626E9D1DEF1AFA20FE86EFC8" box="[321,351,1457,1481]" captionStart="FIGURES 91–94" captionStartId="97.[151,263,1788,1811]" captionTargetBox="[192,1391,186,1708]" captionTargetId="figure-96@97.[181,1403,176,1722]" captionTargetPageId="97" captionText="FIGURES 91–94. Leptomorphus male genitalia, ventral. 91. L. aliciae. 92. L. amorimi. 93. L. babai. 94. L. bifasciatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="15" pageNumber="16">93</figureCitation>
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,
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<figureCitation id="DAA62A5B626E9D1DEF30FA20FE4EEFC8" box="[363,407,1457,1481]" captionStart="FIGURE 140–141" captionStartId="109.[151,249,1594,1617]" captionTargetBox="[151,1436,193,1573]" captionTargetId="figure-53@109.[151,1436,193,1573]" captionTargetPageId="109" captionText="FIGURE 140–141. Distribution of Palaearctic Leptomorphus. 140. L. forcipatus (black square), L. subforcipatus (blue triangle), L. talyshensis (black circle), L. walkeri (red star). 141. L. babai (blue triangle), L. panorpiformis (red star), L. quadrimaculatus (black square)." pageId="15" pageNumber="16">141</figureCitation>
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,
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<figureCitation id="DAA62A5B626E9D1DEFF8FA20FE16EFC8" box="[419,463,1457,1481]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="15" pageNumber="16">150</figureCitation>
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,
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<figureCitation id="DAA62A5B626E9D1DEF80FA20FDDFEFC8" box="[475,518,1457,1481]" captionStart="FIGURE 152" captionStartId="114.[151,250,1784,1807]" captionTargetBox="[162,1422,1059,1758]" captionTargetId="figure-112@114.[156,1428,1046,1779]" captionTargetPageId="114" captionText="FIGURE 152. One of three equally parsimonious trees showing the relationships between species in the L. ornatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="15" pageNumber="16">152</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A876555626E9D1DEECCFA6BFE1EEC96" pageId="15" pageNumber="16" type="reference_group">
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||
<paragraph id="422236DE626E9D1DEECCFA6BFD94EC11" blockId="15.[151,731,1530,1582]" box="[151,589,1530,1552]" pageId="15" pageNumber="16">
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<treatmentCitationGroup id="628D11F0626E9D1DEECCFA6BFD94EC11" box="[151,589,1530,1552]" pageId="15" pageNumber="16">
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<treatmentCitation id="C33C10CF626E9D1DEECCFA6BFD90EC11" author="Sasakawa, M." box="[151,585,1530,1552]" page="187" pageId="15" pageNumber="16" year="1961">
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||
<taxonomicName id="859D4D5D626E9D1DEECCFA6BFD90EC11" ID-CoL="8CJ3Q" authority="Sasakawa, 1961: 187" authorityName="Sasakawa" authorityPageNumber="187" authorityYear="1961" box="[151,585,1530,1552]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="babai">
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||
<emphasis id="70E9EACC626E9D1DEECCFA6BFEB0EC11" box="[151,361,1530,1552]" italics="true" pageId="15" pageNumber="16">Leptomorphus babai</emphasis>
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||
<bibRefCitation id="260C4B2F626E9D1DEF34FA6BFD90EC11" author="Sasakawa, M." box="[367,585,1530,1552]" pageId="15" pageNumber="16" refId="ref95678" refString="Sasakawa, M. (1961) Japanese Fungivoridae. II. New or little known fungus gnats, with descriptions of five new species. Kontyu, 29,186 - 194." type="book" year="1961">Sasakawa, 1961: 187</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="422236DE626E9D1DEECCF989FD02EC2F" blockId="15.[151,731,1530,1582]" box="[151,731,1560,1582]" pageId="15" pageNumber="16">
|
||
<treatmentCitationGroup id="628D11F0626E9D1DEECCF989FD02EC2F" box="[151,731,1560,1582]" pageId="15" pageNumber="16">
|
||
<treatmentCitation id="C33C10CF626E9D1DEECCF989FD0EEC2F" author="Matile, L." box="[151,727,1560,1582]" page="144" pageId="15" pageNumber="16" year="1977">
|
||
<taxonomicName id="859D4D5D626E9D1DEECCF989FD0EEC2F" ID-CoL="8CJ3Q" authority=": Matile, 1977: 144" authorityName=": Matile" authorityPageNumber="144" authorityYear="1977" box="[151,727,1560,1582]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="babai" subGenus="Leptomorphus">
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||
<emphasis id="70E9EACC626E9D1DEECCF989FEF0EC2F" box="[151,297,1560,1582]" italics="true" pageId="15" pageNumber="16">Leptomorphus</emphasis>
|
||
(
|
||
<emphasis id="70E9EACC626E9D1DEF63F989FE13EC2F" box="[312,458,1560,1582]" italics="true" pageId="15" pageNumber="16">Leptomorphus</emphasis>
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||
)
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||
<emphasis id="70E9EACC626E9D1DEF82F989FDCAEC2F" box="[473,531,1560,1582]" italics="true" pageId="15" pageNumber="16">babai</emphasis>
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||
:
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||
<bibRefCitation id="260C4B2F626E9D1DEC44F989FD0EEC2F" author="Matile, L." box="[543,727,1560,1582]" pageId="15" pageNumber="16" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 144</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="422236DE626E9D1DEECCF9CBFE1EEC96" blockId="15.[151,1437,1626,2011]" pageId="15" pageNumber="16">
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||
<emphasis id="70E9EACC626E9D1DEECCF9CBFEC0EC72" bold="true" box="[151,281,1626,1651]" pageId="15" pageNumber="16">References</emphasis>
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||
:
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<bibRefCitation id="260C4B2F626E9D1DEF72F9CAFE35EC72" author="Matile, L." box="[297,492,1627,1651]" pageId="15" pageNumber="16" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977: 144</bibRefCitation>
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(subgeneric placement);
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<bibRefCitation id="260C4B2F626E9D1DED57F9CAFC0EEC72" author="Matile, L." box="[780,983,1627,1651]" pageId="15" pageNumber="16" pagination="231 - 241" refId="ref94679" refString="Matile, L. (1988) Tribe Sciophilini. In: Soos, A. and Papp, L. Catalogue of Palaearctic Diptera: Volume 3: Ceratopogonidae - Mycetophilidae. Elsevier, Amsterdam, pp. 231 - 241." type="book chapter" year="1988">Matile, 1988: 234</bibRefCitation>
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(catalogue);
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<bibRefCitation id="260C4B2F626E9D1DEA35F9CAFF20EC96" author="Matsumoto, R. & Sasakawa, M." pageId="15" pageNumber="16" pagination="13 - 30" refId="ref94759" refString="Matsumoto, R. and Sasakawa, M. (2006) A list of the holotype-specimens of Diptera described by Dr. M. Sasakawa and coworkers, and deposited in the Osaka Museum of Natural History. Bulletin of the Osaka Museum of Natural History, 60, 13 - 30." type="journal article" year="2006">Matsumoto and Sasakawa, 2006: 16</bibRefCitation>
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(
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<typeStatus id="9D26887C626E9D1DEF53F9EEFEE3EC96" box="[264,314,1663,1687]" pageId="15" pageNumber="16">type</typeStatus>
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specimens).
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</paragraph>
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</subSubSection>
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<subSubSection id="0A876555626E9D1DEE9CF933FE29ED4A" pageId="15" pageNumber="16" type="diagnosis">
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||
<paragraph id="422236DE626E9D1DEE9CF933FBA1ED02" blockId="15.[151,1437,1626,2011]" pageId="15" pageNumber="16">
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||
<emphasis id="70E9EACC626E9D1DEE9CF933FEBDECBA" bold="true" box="[199,356,1698,1723]" pageId="15" pageNumber="16">DIAGNOSIS</emphasis>
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: The only extant species of
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<taxonomicName id="859D4D5D626E9D1DEC92F932FCB2ECBA" authorityName=": Edwards" authorityYear="1925" box="[713,875,1699,1723]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
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||
<emphasis id="70E9EACC626E9D1DEC92F932FCB2ECBA" box="[713,875,1699,1723]" italics="true" pageId="15" pageNumber="16">Leptomorphus</emphasis>
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||
</taxonomicName>
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with the following combination of characters: laterotergite and anepisternum brown; wing with pale brown apical wing spot reaching to wing tip, pale medial wing spot present (
|
||
<figureCitation id="DAA62A5B626E9D1DEF37F97AFE19ED02" box="[364,448,1771,1795]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="15" pageNumber="16">Fig. 44</figureCitation>
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||
); male genitalia with gonostylus a single blunt taper (
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||
<figureCitation id="DAA62A5B626E9D1DEA4CF97AFBB2ED02" box="[1047,1131,1771,1795]" captionStart="FIGURES 91–94" captionStartId="97.[151,263,1788,1811]" captionTargetBox="[192,1391,186,1708]" captionTargetId="figure-96@97.[181,1403,176,1722]" captionTargetPageId="97" captionText="FIGURES 91–94. Leptomorphus male genitalia, ventral. 91. L. aliciae. 92. L. amorimi. 93. L. babai. 94. L. bifasciatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="15" pageNumber="16">Fig. 93</figureCitation>
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).
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</paragraph>
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<paragraph id="422236DE626E9D1DEE9CF89EFE29ED4A" blockId="15.[151,1437,1626,2011]" pageId="15" pageNumber="16">
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||
This species can be distinguished from all other Palaearctic and
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||
<collectingRegion id="8059F83C626E9D1DEDC0F89EFC2EED26" box="[923,1015,1807,1831]" country="Republic of the Congo" name="Orientale" pageId="15" pageNumber="16">Oriental</collectingRegion>
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species by the striped abdomen (
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||
<figureCitation id="DAA62A5B626E9D1DEB35F89EFF7FED4A" captionStart="FIGURES 1–4" captionStartId="83.[151,263,1566,1589]" captionTargetBox="[153,1436,289,1543]" captionTargetId="figure-45@83.[151,1436,287,1544]" captionTargetPageId="83" captionText="FIGURES 1–4. Leptomorphus habitus. 1. L. aliciae, male. 2. L. babai, paratype male. 3. L. amorimi, paratype female. 4. L. bifasciatus, female." pageId="15" pageNumber="16">Fig. 2</figureCitation>
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) and/or the brown scutellum.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0A876555626E9D02EE9CF8C7FE1BEC5F" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="422236DE626E9D02EE9CF8C7FC06EC14" blockId="15.[151,1437,1626,2011]" lastBlockId="16.[151,1436,151,1954]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
<emphasis id="70E9EACC626E9D1DEE9CF8C7FE50ED6E" bold="true" box="[199,393,1878,1903]" pageId="15" pageNumber="16">DESCRIPTION</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC626E9D1DEFC1F8C6FE0AED6E" box="[410,467,1879,1903]" italics="true" pageId="15" pageNumber="16">Male</emphasis>
|
||
. (
|
||
<figureCitation id="DAA62A5B626E9D1DEFB7F8C6FDEBED6E" box="[492,562,1879,1903]" captionStart="FIGURES 1–4" captionStartId="83.[151,263,1566,1589]" captionTargetBox="[153,1436,289,1543]" captionTargetId="figure-45@83.[151,1436,287,1544]" captionTargetPageId="83" captionText="FIGURES 1–4. Leptomorphus habitus. 1. L. aliciae, male. 2. L. babai, paratype male. 3. L. amorimi, paratype female. 4. L. bifasciatus, female." pageId="15" pageNumber="16">Fig. 2</figureCitation>
|
||
)
|
||
<emphasis id="70E9EACC626E9D1DEC18F8C6FD5AED6E" bold="true" box="[579,643,1879,1903]" pageId="15" pageNumber="16">Head</emphasis>
|
||
: brown, sometimes with posterior margin yellow, circular in anterior view. Antenna with basal 2 flagellomeres yellow/light brown, brown apically; scape yellow, with light brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 1–2 large bristles, few setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base light brown remainder light brown; flagellomere 6 1.0X as long as broad. Clypeus yellow, dorsoventrally elongate oval; bristles on clypeus yellow, 4–6 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face yellow; shape a just longer than wide triangle, bare. Frons light brown to brown; bare, frontal furrow running 3/4 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus yellow; segment 1 hidden behind eye, segments increasing in length, segment 5 subequal in length to segment 4 with even width from base to apex, segment 3 with apicolateral patch of fine yellow setae weakly encircled by strong dark setae. Labellum yellow. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput brown on anterior 1/2–2/3, remainder yellow with appressed, anteriorly directed setae. Ocelli with median slightly in front of laterals, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black with electric blue green specks.
|
||
<emphasis id="70E9EACC62719D02EAE2FD91FAC8E819" bold="true" box="[1209,1297,512,536]" pageId="16" pageNumber="17">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62719D02EB22FE6EFF1BE83A" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="17" unit="mm" value="1.2">1.2 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62719D02EE8FFDB5FEA7E83A" box="[212,382,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.27" metricValueMin="1.19" pageId="16" pageNumber="17" unit="mm" value="1.23" valueMax="1.27" valueMin="1.19">1.19–1.27 mm</quantity>
|
||
, n = 2). Mostly brown, with anterolateral area yellow. Scutum dark brown with blue-green specks; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; double row of lateral setae present; patch of setae on scutum at wing base present. Scutellum dark brown; with 6 large bristles and many small bristles. Prescutum brown. Mediotergite brown with 8–10 bristles on posterolateral corners, absent. Laterotergite brown; anterior margin of laterotergite not reaching katepisternum. Anepimeron brown. Anepisternum brown. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter stem light brown, knob dark brown.
|
||
<emphasis id="70E9EACC62719D02EDEFFCB1FC35E939" bold="true" box="[948,1004,800,824]" pageId="16" pageNumber="17">Legs</emphasis>
|
||
: principally yellow; hind femur with basal 1/3 and apex light brown; extreme anteroapical corner dark brown on all femora. Midfemur without apical spine-like process. Tibia with covering of yellow macrotrichia, foretibia without comb of short setae along length of anteroventral surface, tibial spurs light brown, foretibial spur length 2X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 3/4 of its length, placed basally, shortest midtibial spur 0.75X length of longest, longest midtibial spur 4.5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 5X apical thickness of hind tibia. Foreleg first tarsomere 1.5–1.6X length of foretibia.
|
||
<emphasis id="70E9EACC62719D02EF78FB8AFEBDEE35" bold="true" box="[291,356,1051,1076]" pageId="16" pageNumber="17">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62719D02EF2FFB8DFE13EE35" box="[372,458,1052,1076]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="16" pageNumber="17">Fig. 44</figureCitation>
|
||
): Length
|
||
<quantity id="85659B3B62719D02EC6FFB8DFD53EE32" box="[564,650,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="16" pageNumber="17" unit="mm" value="5.3">5.3 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62719D02ECC2FB8DFCFDEE32" box="[665,804,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="16" pageNumber="17" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.1–5.4 mm</quantity>
|
||
, n = 2). Hyaline; apical macula light, reaching wing tip, extending faintly along posterior wing margin into apex of cell cua
|
||
<subScript id="DE19349B62719D02EDD4FBDEFC4EEE5C" attach="left" box="[911,919,1103,1117]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
|
||
, cup, not joining with medial macula; medial macula light, extending from R
|
||
<subScript id="DE19349B62719D02EFA8FBE7FE22EE85" attach="left" box="[499,507,1142,1156]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
|
||
to just beyond fork of M
|
||
<subScript id="DE19349B62719D02ED4AFBE7FCC0EE85" attach="left" box="[785,793,1142,1156]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
|
||
and M
|
||
<subScript id="DE19349B62719D02ED3CFBE7FCB6EE85" attach="left" box="[871,879,1142,1156]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
|
||
. Macrotrichia in all cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present, apical end joining R at 2X its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62719D02ED28FB51FCA2EECF" attach="left" box="[883,891,1216,1230]" fontSize="6" pageId="16" pageNumber="17">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62719D02EDB3FB51FC29EECF" attach="left" box="[1000,1008,1216,1230]" fontSize="6" pageId="16" pageNumber="17">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B62719D02EEEBFB76FF61EEF4" attach="left" box="[176,184,1255,1269]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B62719D02EF9AFB76FE10EEF4" attach="left" box="[449,457,1255,1269]" fontSize="6" pageId="16" pageNumber="17">5</subScript>
|
||
, apices of M veins thinning towards wing margin. M
|
||
<subScript id="DE19349B62719D02EA43FB76FBF9EEF4" attach="left" box="[1048,1056,1255,1269]" fontSize="6" pageId="16" pageNumber="17">4</subScript>
|
||
-CuA fork arising at same level as r-m. A
|
||
<subScript id="DE19349B62719D02EEBAFA9FFF30EF1D" attach="left" box="[225,233,1294,1308]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62719D02EF1EFB6FFE63EF16" bold="true" box="[325,442,1278,1303]" pageId="16" pageNumber="17">Abdomen</emphasis>
|
||
: Tergites principally dark brown, T2–6 with anterolateral yellow spots. Tergite 8 smaller than all other abdominal sclerites and ~15 bristles laterally, centre bare.
|
||
<emphasis id="70E9EACC62719D02ED98FAB4FBE8EF3F" bold="true" box="[963,1073,1317,1342]" pageId="16" pageNumber="17">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62719D02EA19FAB7FB4FEF3F" box="[1090,1174,1318,1342]" captionStart="FIGURES 91–94" captionStartId="97.[151,263,1788,1811]" captionTargetBox="[192,1391,186,1708]" captionTargetId="figure-96@97.[181,1403,176,1722]" captionTargetPageId="97" captionText="FIGURES 91–94. Leptomorphus male genitalia, ventral. 91. L. aliciae. 92. L. amorimi. 93. L. babai. 94. L. bifasciatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="16" pageNumber="17">Fig. 93</figureCitation>
|
||
): yellow. Sternite 9 not clearly visible. Tergite 9 longer than wide, rounded margins tapering into two short pointed lobes and shallow medial indentation. Gonocoxite placed basally on T9, medial margins almost touching near base due to triangular projection, a ventromedial lobe covering base of gonostylus, bearing gonostylus halfway to apex. Gonostylus a single broad-based lobe tapering towards rounded apex, gonocoxite III fused to dorsolateral margin. Aedeagus 2/3 length of gonocoxite, slightly tapering towards apex, apodemes 1/5 total length with small dorsally facing hook basally. Parameres a short, wide taper, apodemes ~4X length of parameres.
|
||
</paragraph>
|
||
<paragraph id="422236DE62719D02EE9CF9B3FE47EC3B" blockId="16.[151,1436,151,1954]" box="[199,414,1570,1594]" pageId="16" pageNumber="17">
|
||
<emphasis id="70E9EACC62719D02EE9CF9B3FEC2EC3B" box="[199,283,1570,1594]" italics="true" pageId="16" pageNumber="17">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="422236DE62719D02EE9CF9D6FE1BEC5F" blockId="16.[151,1436,151,1954]" box="[199,450,1606,1630]" pageId="16" pageNumber="17">
|
||
<emphasis id="70E9EACC62719D02EE9CF9D6FEE7EC5F" box="[199,318,1607,1630]" italics="true" pageId="16" pageNumber="17">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562719D02EE9CF9F8FE16EC83" box="[199,463,1641,1666]" pageId="16" pageNumber="17" type="biology_ecology">
|
||
<paragraph id="422236DE62719D02EE9CF9F8FE16EC83" blockId="16.[151,1436,151,1954]" box="[199,463,1641,1666]" pageId="16" pageNumber="17">
|
||
<emphasis id="70E9EACC62719D02EE9CF9F8FE93EC83" bold="true" box="[199,330,1641,1666]" pageId="16" pageNumber="17">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562719D02EE9CF91CFD11ECA7" box="[199,712,1677,1702]" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="422236DE62719D02EE9CF91CFD11ECA7" blockId="16.[151,1436,151,1954]" box="[199,712,1677,1702]" pageId="16" pageNumber="17">
|
||
<emphasis id="70E9EACC62719D02EE9CF91CFE4FECA7" bold="true" box="[199,406,1677,1702]" pageId="16" pageNumber="17">DISTRIBUTION</emphasis>
|
||
:
|
||
<collectingCountry id="3A8A764E62719D02EFFFF91FFE3DECA4" box="[420,484,1678,1701]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B62719D02EFA8F91FFD8DECA7" box="[499,596,1678,1702]" captionStart="FIGURE 140–141" captionStartId="109.[151,249,1594,1617]" captionTargetBox="[151,1436,193,1573]" captionTargetId="figure-53@109.[151,1436,193,1573]" captionTargetPageId="109" captionText="FIGURE 140–141. Distribution of Palaearctic Leptomorphus. 140. L. forcipatus (black square), L. subforcipatus (blue triangle), L. talyshensis (black circle), L. walkeri (red star). 141. L. babai (blue triangle), L. panorpiformis (red star), L. quadrimaculatus (black square)." pageId="16" pageNumber="17">Fig. 141</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62719D02EC32F91FFD1DECA7" box="[617,708,1678,1702]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="17" unit="m" value="40.0">40 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562719D02EE9CF920FC69ED13" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="422236DE62719D02EE9CF920FC69ED13" blockId="16.[151,1436,151,1954]" pageId="16" pageNumber="17">
|
||
<emphasis id="70E9EACC62719D02EE9CF920FEABECCB" bold="true" box="[199,370,1713,1738]" pageId="16" pageNumber="17">DISCUSSION</emphasis>
|
||
: As discussed below in the phylogeny section, the placement of
|
||
<taxonomicName id="859D4D5D62719D02EA27F923FAA3ECCB" box="[1148,1402,1714,1738]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="70E9EACC62719D02EA27F923FAC7ECCB" box="[1148,1310,1714,1738]" italics="true" pageId="16" pageNumber="17">Leptomorphus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in subgenera (
|
||
<bibRefCitation id="260C4B2F62719D02EF43F947FE7DECEF" author="Matile, L." box="[280,420,1750,1774]" pageId="16" pageNumber="17" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
|
||
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
|
||
<taxonomicName id="859D4D5D62719D02EF5DF96BFE71ED13" authorityName=": Edwards" authorityYear="1925" box="[262,424,1786,1810]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E9EACC62719D02EF5DF96BFE71ED13" box="[262,424,1786,1810]" italics="true" pageId="16" pageNumber="17">Leptomorphus</emphasis>
|
||
</taxonomicName>
|
||
and placed solely in the genus
|
||
<emphasis id="70E9EACC62719D02ED5DF96BFC69ED13" box="[774,944,1786,1810]" italics="true" pageId="16" pageNumber="17">
|
||
<taxonomicName id="859D4D5D62719D02ED5DF96BFC75ED13" authorityName=": Edwards" authorityYear="1925" box="[774,940,1786,1810]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562719D02EE9CF88CFC6FEDA3" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="422236DE62719D02EE9CF88CFBA7ED7F" blockId="16.[151,1436,151,1954]" pageId="16" pageNumber="17">
|
||
<materialsCitation id="F2F53C8362719D02EE9CF88CFBA7ED7F" collectingDate="1955-07-02" collectionCode="MATERIAL, OMNH" collectorName="Col. Kintaro" country="Japan" location="Kurokawa" municipality="Echigo" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62719D02EE9CF88CFDDAED37" bold="true" box="[199,515,1821,1846]" pageId="16" pageNumber="17">
|
||
<collectionCode id="248CAE1B62719D02EE9CF88CFE86ED37" box="[199,351,1821,1846]" pageId="16" pageNumber="17">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62719D02EC48F88FFD5BED37" box="[531,642,1822,1846]" italics="true" pageId="16" pageNumber="17">
|
||
<typeStatus id="9D26887C62719D02EC48F88FFDA5ED37" box="[531,636,1822,1846]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
adult male, pointed, with wing in separate glass and paper mount, all on same pin, genitalia missing, labelled “[translucent paper]
|
||
<location id="4742600562719D02ED12F8D3FC67ED5B" LSID="urn:lsid:plazi:treatment:CA3487C8626E9D02EECCFA1EFC6FEDA3:4742600562719D02ED12F8D3FC67ED5B" box="[841,958,1858,1882]" country="Japan" municipality="Echigo" name="Kurokawa" pageId="16" pageNumber="17">Kurokawa</location>
|
||
/
|
||
<collectingMunicipality id="A246ACA462719D02ED96F8D3FBC6ED5B" box="[973,1055,1858,1882]" pageId="16" pageNumber="17">Echigo</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="3A8A764E62719D02EA77F8D3FBB2ED5B" box="[1068,1131,1858,1882]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
|
||
/
|
||
<date id="3623101E62719D02EA27F8D3FB2CED5B" box="[1148,1269,1858,1882]" pageId="16" pageNumber="17" value="1955-07-02">
|
||
<collectingDate id="2667E9F662719D02EA27F8D3FB2CED5B" box="[1148,1269,1858,1882]" pageId="16" pageNumber="17" value="1955-07-02">2.VII.1955</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="EF68530862719D02EB5CF8D3FA41ED5B" box="[1287,1432,1858,1882]" pageId="16" pageNumber="17">Col. Kintaro</collectorName>
|
||
; [pink label]
|
||
<typeStatus id="9D26887C62719D02EF7BF8F7FE6DED7F" box="[288,436,1894,1918]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD5762719D02EFE1F8F7FE16ED7C" box="[442,463,1894,1917]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
|
||
/ KPU 0163/
|
||
<taxonomicName id="859D4D5D62719D02EC3FF8F7FCDEED7F" authorityName=": Edwards" authorityYear="1925" box="[612,775,1894,1918]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
/ babai/ SASAKAWA” [
|
||
<collectionCode id="248CAE1B62719D02EA4EF8F7FBADED7F" box="[1045,1140,1894,1918]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="16" pageNumber="17" type="Museum">OMNH</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="422236DE62719D02EE9CF81AFC6FEDA3" blockId="16.[151,1436,151,1954]" box="[199,950,1930,1954]" pageId="16" pageNumber="17">
|
||
<materialsCitation id="F2F53C8362719D02EE9CF81AFD16EDA3" box="[199,719,1930,1954]" collectionCode="VI" pageId="16" pageNumber="17" specimenCount="2" typeStatus="paratype">
|
||
<emphasis id="70E9EACC62719D02EE9CF81AFEEFEDA3" box="[199,310,1931,1954]" italics="true" pageId="16" pageNumber="17">
|
||
<typeStatus id="9D26887C62719D02EE9CF81AFEE8EDA3" box="[199,305,1931,1954]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
labelled as for
|
||
<typeStatus id="9D26887C62719D02EFB8F81BFD9CEDA3" box="[483,581,1930,1954]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
except
|
||
<date id="3623101E62719D02ECC7F81BFD16EDA3" box="[668,719,1930,1954]" pageId="16" pageNumber="17" value="1955-06-07">
|
||
7.
|
||
<collectionCode id="248CAE1B62719D02ECF5F81BFD16EDA3" box="[686,719,1930,1954]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="17">VI</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="F2F53C8362719D02EC94F81BFC6BEDA3" box="[719,946,1930,1954]" collectionCode="OMNH" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
.1955 (
|
||
<specimenCount id="549BFD5762719D02ED7BF81BFC98EDA0" box="[800,833,1930,1954]" pageId="16" pageNumber="17" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62719D02ED14F81BFC73EDA3" box="[847,938,1930,1954]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="16" pageNumber="17" type="Museum">OMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |