277 lines
39 KiB
XML
277 lines
39 KiB
XML
<document id="BD8AB65A19350DB75929B09767E6A4E8" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C8625F9D2DEECCFDFAFD18EFA5" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus tabatius Borkent 2012, new species" docType="treatment" docUuid="71ACD33E-E56E-4A28-8E24-8D74297FE85E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="64" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="63" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="61F543EBB289EB937A4223D89EF6B832" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="DAE58746A938E8C5B12FC5AE36A8AAF5">
|
||
<mods:title id="FC2BACEC2AB331633A99C8892B4441EB">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="10F94288405CCA58481BA060E47DB973" type="personal">
|
||
<mods:role id="2943FB946BCFD8CA87462E0EAB9E3524">
|
||
<mods:roleTerm id="642BDD9E63EEB0097B6E21C54EE7C879">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="61C751B32F013439EF20283A5A35FA0F">Borkent, Christopher J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="45B606A74789D86AB574886F92179F22" type="personal">
|
||
<mods:role id="9B6B8649D29CFEDC0EA9F2440BDA313A">
|
||
<mods:roleTerm id="FC1DDDC22FEBCA8046A7601C523D4C42">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8DB9BF9854C0811A8F58185383C3D76C">Wheeler, Terry A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="51541420749F7582AC0F0D523CEF77A5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BEF20364735011DABE4A156B6DD24E7E" type="host">
|
||
<mods:titleInfo id="5B9F21AF7D7C01DF6795B5E1EFA5B4C6">
|
||
<mods:title id="CE2C8E54EE33AD847F6D86BACC7F943C">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="761F826858EF2C89CF468740B0434015">
|
||
<mods:date id="AF92037128F49BCFA2271551E7513045">2012</mods:date>
|
||
<mods:detail id="BE182C4CC827457D788A9BDDA4BBCFB9" type="pubDate">
|
||
<mods:number id="CFE9C4D3A8B05D4FDC5E773D06E2A1F9">2012-11-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A9E9AEFADE0A2A972459003E76C760DF" type="volume">
|
||
<mods:number id="5443066C2B5559C9F10B29018B86575E">3549</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8EB8ADED8E03D7A45154991890AE2A81" unit="page">
|
||
<mods:start id="EADE6BF6553F0846C72BC07A2B7C7615">1</mods:start>
|
||
<mods:end id="29D4A660C7A364BFD8DFEB3D60261EBB">117</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="E2484309321D89F27637FA2465B8FF48">journal article</mods:classification>
|
||
<mods:identifier id="82C631D2FC148BE63C2806784C40A3A3" type="ISSN">11755334</mods:identifier>
|
||
<mods:identifier id="7BA6AA530633F0B78CFFE1C1763A17A6" type="ZooBank">2412CB4F-4D29-4988-80C1-205D16767678</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CA3487C8625F9D2DEECCFDFAFD18EFA5" ID-DOI="http://doi.org/10.5281/zenodo.5258367" ID-Zenodo-Dep="5258367" LSID="urn:lsid:zoobank.org:act:71ACD33E-E56E-4A28-8E24-8D74297FE85E" httpUri="http://treatment.plazi.org/id/CA3487C8625F9D2DEECCFDFAFD18EFA5" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
<subSubSection id="0A876555625F9D2CEECCFDFAFD3FE884" box="[151,742,619,646]" pageId="62" pageNumber="63" type="nomenclature">
|
||
<paragraph id="422236DE625F9D2CEECCFDFAFD3FE884" blockId="62.[151,742,619,678]" box="[151,742,619,646]" pageId="62" pageNumber="63">
|
||
<heading id="196A81B2625F9D2CEECCFDFAFD3FE884" bold="true" box="[151,742,619,646]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
|
||
<emphasis id="70E9EACC625F9D2CEECCFDFAFD3FE884" bold="true" box="[151,742,619,646]" pageId="62" pageNumber="63">
|
||
32.
|
||
<taxonomicName id="859D4D5D625F9D2CEE98FDFDFD90E884" ID-CoL="8CJ4H" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[195,585,619,646]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tabatius" status="sp. nov.">
|
||
<emphasis id="70E9EACC625F9D2CEE98FDFDFE00E887" bold="true" box="[195,473,620,646]" italics="true" pageId="62" pageNumber="63">Leptomorphus tabatius</emphasis>
|
||
Borkent
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6BDA57B7625F9D2CEC0DFDFAFD3FE884" box="[598,742,619,645]" pageId="62" pageNumber="63" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625F9D2CEECCFD1FFD8DE8A7" box="[151,596,654,678]" pageId="62" pageNumber="63" type="description">
|
||
<paragraph id="422236DE625F9D2CEECCFD1FFD8DE8A7" blockId="62.[151,742,619,678]" box="[151,596,654,678]" pageId="62" pageNumber="63">
|
||
(
|
||
<figureCitation id="DAA62A5B625F9D2CEEC5FD1FFEC0E8A7" box="[158,281,654,678]" captionStart="FIGURES 34–37" captionStartId="90.[151,263,1623,1646]" captionTargetBox="[152,1434,181,1602]" captionTargetId="figure-41@90.[151,1436,179,1602]" captionTargetPageId="90" captionText="FIGURES 34–37. Leptomorphus habitus. 34. L. stigmatus, holotype male. 35. L. tabatius, holotype male. 36. L. subcaeruleus, male. 37. L. subcaeruleus, female." pageId="62" pageNumber="63">Figures 35</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEF7EFD1FFE9AE8A7" box="[293,323,654,678]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="62" pageNumber="63">73</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEF14FD1FFEB4E8A7" box="[335,365,654,678]" captionStart="FIGURES 78–81" captionStartId="95.[151,263,1524,1547]" captionTargetBox="[151,1436,193,1503]" captionTargetId="figure-60@95.[151,1436,193,1503]" captionTargetPageId="95" captionText="FIGURES 78–81. Leptomorphus morphology. 78–79. Dorsal view of head showing ocelli placement and arrangement. 78. L. tabatius. 79. L. perplexus. 80–81. Dorsal view of scutum. 80. L. fasciculatus showing covering of fine trichia. 81. L. stigmatus showing pattern and lack of setae or trichia." pageId="62" pageNumber="63">78</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEF22FD1FFE7CE8A7" box="[377,421,654,678]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev & Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="62" pageNumber="63">120</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEFEAFD1FFE04E8A7" box="[433,477,654,678]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp & Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="62" pageNumber="63">137</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEFB2FD1FFDCCE8A7" box="[489,533,654,678]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="62" pageNumber="63">150</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B625F9D2CEC7AFD1FFD95E8A7" box="[545,588,654,678]" captionStart="FIGURE 152" captionStartId="114.[151,250,1784,1807]" captionTargetBox="[162,1422,1059,1758]" captionTargetId="figure-112@114.[156,1428,1046,1779]" captionTargetPageId="114" captionText="FIGURE 152. One of three equally parsimonious trees showing the relationships between species in the L. ornatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="62" pageNumber="63">152</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625F9D2CEECCFD47FCA9E9EB" pageId="62" pageNumber="63" type="diagnosis">
|
||
<paragraph id="422236DE625F9D2CEECCFD47FDAFE95B" blockId="62.[151,1437,726,2011]" pageId="62" pageNumber="63">
|
||
<emphasis id="70E9EACC625F9D2CEECCFD47FEEDE8EE" bold="true" box="[151,308,726,751]" pageId="62" pageNumber="63">DIAGNOSIS</emphasis>
|
||
: The only extant species of
|
||
<taxonomicName id="859D4D5D625F9D2CECEAFD47FC8AE8EF" authorityName=": Edwards" authorityYear="1925" box="[689,851,726,750]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E9EACC625F9D2CECEAFD47FC8AE8EF" box="[689,851,726,750]" italics="true" pageId="62" pageNumber="63">Leptomorphus</emphasis>
|
||
</taxonomicName>
|
||
with the following combination of characters: laterotergite and anepisternum brown; wing without apical spot; segments 3–5 of abdomen noticeably swollen relative to other segments; male genitalia with tergite 9 bearing two long lateral processes posteriorly, so that tergite 9 is ~2X as long as wide (
|
||
<figureCitation id="DAA62A5B625F9D2CEC5CFCD3FDB0E95B" box="[519,617,834,858]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev & Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="62" pageNumber="63">Fig. 120</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="422236DE625F9D2CEE9CFCF7FCA9E9EB" blockId="62.[151,1437,726,2011]" pageId="62" pageNumber="63">
|
||
This species can be distinguished from the other
|
||
<collectingRegion id="8059F83C625F9D2CECAEFCF7FC89E97F" box="[757,848,870,894]" country="Republic of the Congo" name="Orientale" pageId="62" pageNumber="63">Oriental</collectingRegion>
|
||
and eastern Palaearctic species either by the brown katepisternum and metepisternum (
|
||
<figureCitation id="DAA62A5B625F9D2CEC7EFC1BFDA1E9A3" box="[549,632,906,930]" captionStart="FIGURES 34–37" captionStartId="90.[151,263,1623,1646]" captionTargetBox="[152,1434,181,1602]" captionTargetId="figure-41@90.[151,1436,179,1602]" captionTargetPageId="90" captionText="FIGURES 34–37. Leptomorphus habitus. 34. L. stigmatus, holotype male. 35. L. tabatius, holotype male. 36. L. subcaeruleus, male. 37. L. subcaeruleus, female." pageId="62" pageNumber="63">Fig. 35</figureCitation>
|
||
, both completely yellow in
|
||
<emphasis id="70E9EACC625F9D2CEDEFFC1AFB4FE9A3" box="[948,1174,907,931]" italics="true" pageId="62" pageNumber="63">
|
||
<taxonomicName id="859D4D5D625F9D2CEDEFFC1AFBCCE9A2" authorityName=": Matile" authorityYear="1977" box="[948,1045,907,931]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="babai">L. babai</taxonomicName>
|
||
,
|
||
<taxonomicName id="859D4D5D625F9D2CEA79FC1AFB4FE9A3" authorityName=": Matile" authorityYear="1977" box="[1058,1174,907,930]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ornatus">L. ornatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="859D4D5D625F9D2CEA95FC1AFA55E9A3" authorityName="Borkent" authorityYear="2012" box="[1230,1420,907,930]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="titiwangsensis">
|
||
<emphasis id="70E9EACC625F9D2CEA95FC1AFA55E9A3" box="[1230,1420,907,930]" italics="true" pageId="62" pageNumber="63">L. titiwangsensis</emphasis>
|
||
</taxonomicName>
|
||
), or by the posterolateral yellow spots on the scutum and the long evaginations (more than half the length of the genitalia) at the posterior margin of the male genitalia (
|
||
<figureCitation id="DAA62A5B625F9D2CED5AFC43FCBDE9EB" box="[769,868,978,1002]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev & Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="62" pageNumber="63">Fig. 120</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625F9D2DEE9CFC67FE1BE989" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
|
||
<paragraph id="422236DE625F9D2DEE9CFC67FDC1E941" blockId="62.[151,1437,726,2011]" lastBlockId="63.[151,1436,151,1444]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
<emphasis id="70E9EACC625F9D2CEE9CFC67FE55EE0E" bold="true" box="[199,396,1014,1039]" pageId="62" pageNumber="63">DESCRIPTION</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC625F9D2CEFC4FC67FE00EE0F" box="[415,473,1014,1038]" italics="true" pageId="62" pageNumber="63">Male</emphasis>
|
||
. (
|
||
<figureCitation id="DAA62A5B625F9D2CEFAEFC67FD92EE0F" box="[501,587,1014,1038]" captionStart="FIGURES 34–37" captionStartId="90.[151,263,1623,1646]" captionTargetBox="[152,1434,181,1602]" captionTargetId="figure-41@90.[151,1436,179,1602]" captionTargetPageId="90" captionText="FIGURES 34–37. Leptomorphus habitus. 34. L. stigmatus, holotype male. 35. L. tabatius, holotype male. 36. L. subcaeruleus, male. 37. L. subcaeruleus, female." pageId="62" pageNumber="63">Fig. 35</figureCitation>
|
||
)
|
||
<emphasis id="70E9EACC625F9D2CEC04FC66FD46EE0E" bold="true" box="[607,671,1015,1039]" pageId="62" pageNumber="63">Head</emphasis>
|
||
: (
|
||
<figureCitation id="DAA62A5B625F9D2CECE0FC67FCCAEE0F" box="[699,787,1014,1038]" captionStart="FIGURES 78–81" captionStartId="95.[151,263,1524,1547]" captionTargetBox="[151,1436,193,1503]" captionTargetId="figure-60@95.[151,1436,193,1503]" captionTargetPageId="95" captionText="FIGURES 78–81. Leptomorphus morphology. 78–79. Dorsal view of head showing ocelli placement and arrangement. 78. L. tabatius. 79. L. perplexus. 80–81. Dorsal view of scutum. 80. L. fasciculatus showing covering of fine trichia. 81. L. stigmatus showing pattern and lack of setae or trichia." pageId="62" pageNumber="63">Fig. 78</figureCitation>
|
||
) light yellow, somewhat dorsoventrally compressed in anterior view. Antenna with basal 2–3 flagellomeres lighter brown, darkening apically; scape yellow, with black setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae absent; pedicel light brown, with 5–6 setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base light brown remainder brown; flagellomere 6 1.1X as long as broad. Clypeus light yellow, dorsoventrally elongate oval; bristles on clypeus brown, 4 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 1.6X as long as face. Face light yellow; shape a slightly wider than tall triangle, with few bristles ventrolaterally. Frons yellow; bare, frontal furrow running full distance from dorsal margin towards ventral margin, frontal cleft more than 1X diameter of median ocellus anterior of median ocellus. Palpus yellow; segment 1 hidden behind eye, segments increasing in length, segment 5 2X length of segment 4 with apex slightly swollen, segment 3 spherical with apicolateral depressed patch of fine yellow setae partially encircled by strong dark setae. Labellum yellow. Eye with inter-ommatidial setulae absent. Occiput yellow with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle black.
|
||
<emphasis id="70E9EACC625F9D2CEFADFA7EFD89EC06" bold="true" box="[502,592,1519,1543]" pageId="62" pageNumber="63">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B625F9D2CECE3FA7FFCC4EC07" box="[696,797,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="62" pageNumber="63" unit="mm" value="1.19">1.19 mm</quantity>
|
||
(n = 1). Dark brown. Scutum dark brown with a pair of small yellow posterolateral spots; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; multiple rows of lateral setae present; patch of setae on scutum at wing base present. Scutellum light yellow; without large bristles but few small bristles. Prescutum light brown. Mediotergite dark brown with 8 bristles on posterolateral corners, anteromedial patch of small setae. Laterotergite brown; anterior margin of laterotergite not reaching katepisternum. Anepimeron brown with posterodorsal corner yellow. Anepisternum brown. Katepisternum brown. Antepronotum and proepisternum brown. Margin of anterior spiracle brown and posterior spiracle yellow, both with light brown trichia. Metepisternum light brown. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/ 3 of stem ivory, apical portion and knob dark brown.
|
||
<emphasis id="70E9EACC625F9D2CECA1F8A2FCEAED4A" bold="true" box="[762,819,1843,1867]" pageId="62" pageNumber="63">Legs</emphasis>
|
||
: Light yellow except for joint of hind femur and tibia which is brown; extreme anteroapical corner dark brown on all femora. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, foretibial and anterior midtibial spurs brown, remainder yellow, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 2/3 of its length, placed basally, shortest midtibial spur subequal to length of longest, longest midtibial spur 4X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 3.5X apical thickness of hind tibia. Foreleg first tarsomere 1.7X length of foretibia.
|
||
<emphasis id="70E9EACC625E9D2DED38FF4EFC7CEAF9" bold="true" box="[867,933,223,248]" pageId="63" pageNumber="64">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B625E9D2DEDEEFF4EFBD5EAF6" box="[949,1036,223,247]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="63" pageNumber="64">Fig. 73</figureCitation>
|
||
): Length 5.0 mm (n = 1). Hyaline; apical macula absent; medial macula absent. Macrotrichia in all cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a group of macrotrichia. Vein sc-r present, apical end joining R at 2X its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B625E9D2DED7DFECAFCF7EB68" attach="left" box="[806,814,347,361]" fontSize="6" pageId="63" pageNumber="64">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B625E9D2DEDF9FECAFC73EB68" attach="left" box="[930,938,347,361]" fontSize="6" pageId="63" pageNumber="64">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B625E9D2DEBCFFECAFA45EB68" attach="left" box="[1428,1436,347,361]" fontSize="6" pageId="63" pageNumber="64">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B625E9D2DEFFBFE13FE71EB91" attach="left" box="[416,424,386,400]" fontSize="6" pageId="63" pageNumber="64">5</subScript>
|
||
, apices of M veins fading before wing margin. M
|
||
<subScript id="DE19349B625E9D2DED85FE13FC3FEB91" attach="left" box="[990,998,386,400]" fontSize="6" pageId="63" pageNumber="64">4</subScript>
|
||
-CuA fork arising apically of origin of r-m. A
|
||
<subScript id="DE19349B625E9D2DEEBEFE39FF34EBB7" attach="left" box="[229,237,424,438]" fontSize="6" pageId="63" pageNumber="64">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC625E9D2DEF0AFE08FE1EEBB3" bold="true" box="[337,455,409,434]" pageId="63" pageNumber="64">Abdomen</emphasis>
|
||
: Abdominal segments 3–5 swollen to 2X thickness of other segments; tergites 3–5 with anterior 1/3–1/2 yellow, remainder brown. Tergite 8 smaller than all other abdominal sclerites, with 8 bristles on each apicolateral corner and few bristles along posterior margin but not as complete row (missing from medial 1/4).
|
||
<emphasis id="70E9EACC625E9D2DEF20FD99FE35E820" bold="true" box="[379,492,520,545]" pageId="63" pageNumber="64">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B625E9D2DEC59FD99FDB7E821" box="[514,622,520,544]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev & Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="63" pageNumber="64">Fig. 120</figureCitation>
|
||
): light yellow. Sternite 9 reduced to long thin sclerite attached to aedeagus. Tergite 9 longer than wide, basal 1/3 rounded and then spreading apically into strongly pointed lateral lobes and deep V-shaped medial notch, mediobasal corner of lobes with two sclerotized points ventrally. Gonocoxite strongly fused with T9 but visible due to difference in sclerotization, placed centrally on T9, medial margin not reaching median line, bearing gonostylus apically. Gonostylus a single broad-based lobe tapering to apex which is bifurcated into two short sclerotized points, gonocoxite III fused to gonostylus basally. Aedeagus 1/4 length of genitalia, broad, rounded base tapering to apex, strongly united basally with S9, apodemes highly reduced. Parameres squared off so essentially only apodemes remain (posterior margin flat), apodemes strongly united with gonocoxal apodemes.
|
||
</paragraph>
|
||
<paragraph id="422236DE625E9D2DEE9CFCDDFE06E965" blockId="63.[151,1436,151,1444]" box="[199,479,844,868]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFCDDFE85E965" box="[199,348,844,868]" italics="true" pageId="63" pageNumber="64">Female adult</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="422236DE625E9D2DEE9CFCE0FE1BE989" blockId="63.[151,1436,151,1444]" box="[199,450,880,904]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFCE0FEE7E989" box="[199,318,881,904]" italics="true" pageId="63" pageNumber="64">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625E9D2DEE9CFC05FE16E9AD" box="[199,463,916,941]" pageId="63" pageNumber="64" type="biology_ecology">
|
||
<paragraph id="422236DE625E9D2DEE9CFC05FE16E9AD" blockId="63.[151,1436,151,1444]" box="[199,463,916,941]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFC05FE93E9AC" bold="true" box="[199,330,916,941]" pageId="63" pageNumber="64">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625E9D2DEE9CFC29FCAEE9D1" box="[199,887,952,977]" pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph id="422236DE625E9D2DEE9CFC29FCAEE9D1" blockId="63.[151,1436,151,1444]" box="[199,887,952,977]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFC29FE4FE9D0" bold="true" box="[199,406,952,977]" pageId="63" pageNumber="64">DISTRIBUTION</emphasis>
|
||
:
|
||
<materialsCitation id="F2F53C83625E9D2DEFFFFC29FCAAE9D1" box="[420,883,952,976]" country="Indonesia" elevation="820" location="Sulawesi" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Sulawesi Utara">
|
||
<collectingRegion id="8059F83C625E9D2DEFFFFC29FDD5E9D1" box="[420,524,952,976]" country="Indonesia" name="Sulawesi Utara" pageId="63" pageNumber="64">Sulawesi</collectingRegion>
|
||
,
|
||
<collectingCountry id="3A8A764E625E9D2DEC43FC29FD5CE9D1" box="[536,645,952,976]" name="Indonesia" pageId="63" pageNumber="64">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B625E9D2DECCFFC29FD2CE9D1" box="[660,757,952,976]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp & Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="63" pageNumber="64">Fig. 137</figureCitation>
|
||
),
|
||
<quantity id="85659B3B625E9D2DED52FC29FCAAE9D1" box="[777,883,952,976]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="63" pageNumber="64" unit="m" value="820.0">
|
||
<elevation id="C9B0D1ED625E9D2DED52FC29FCAAE9D1" box="[777,883,952,976]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="63" pageNumber="64" unit="m" value="820.0">820 masl</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625E9D2DEE9CFC4DFDE8EE3D" pageId="63" pageNumber="64" type="discussion">
|
||
<paragraph id="422236DE625E9D2DEE9CFC4DFDE8EE3D" blockId="63.[151,1436,151,1444]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFC4DFEABE9F4" bold="true" box="[199,370,988,1013]" pageId="63" pageNumber="64">DISCUSSION</emphasis>
|
||
: This species appears to have male genitalia similar to, but sufficiently distinct from, those of
|
||
<taxonomicName id="859D4D5D625E9D2DEECCFB90FE24EE19" authority="Papp & Sevcik. We" authorityName="Papp & Sevcik. We" authorityYear="2011" box="[151,509,1024,1049]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="alienus">
|
||
<emphasis id="70E9EACC625E9D2DEECCFB90FED3EE19" box="[151,266,1024,1048]" italics="true" pageId="63" pageNumber="64">L. alienus</emphasis>
|
||
Papp & Ševčík. We
|
||
</taxonomicName>
|
||
were unable to examine specimens of that species for comparison and cannot comment further on the relationship.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625E9D2DEE9CFBD9FE46EEA9" pageId="63" pageNumber="64" type="etymology">
|
||
<paragraph id="422236DE625E9D2DEE9CFBD9FE46EEA9" blockId="63.[151,1436,151,1444]" pageId="63" pageNumber="64">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFBD9FE58EE60" bold="true" box="[199,385,1096,1121]" pageId="63" pageNumber="64">ETYMOLOGY</emphasis>
|
||
:
|
||
<materialsCitation id="F2F53C83625E9D2DEFCBFBD9FE42EEA9" county="This" location="Tolaki" municipality="Tolaki" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<collectingCounty id="AB434E52625E9D2DEFCBFBD9FE1BEE61" box="[400,450,1096,1120]" pageId="63" pageNumber="64">This</collectingCounty>
|
||
species name is derived from the
|
||
<collectingMunicipality id="A246ACA4625E9D2DED1DFBD9FC54EE61" box="[838,909,1096,1120]" pageId="63" pageNumber="64">Tolaki</collectingMunicipality>
|
||
for ‘fat’ (taba) ‘belly’ (tia), in reference to the swollen abdomen.
|
||
<location id="47426005625E9D2DEF31FBFDFE68EE85" LSID="urn:lsid:plazi:treatment:CA3487C8625F9D2DEECCFDFAFD18EFA5:47426005625E9D2DEF31FBFDFE68EE85" box="[362,433,1132,1156]" county="This" municipality="Tolaki" name="Tolaki" pageId="63" pageNumber="64" stateProvince="Sulawesi Utara">Tolaki</location>
|
||
is the language spoken by the people of the
|
||
<location id="47426005625E9D2DEDFBFBFDFB4FEE85" LSID="urn:lsid:plazi:treatment:CA3487C8625F9D2DEECCFDFAFD18EFA5:47426005625E9D2DEDFBFBFDFB4FEE85" box="[928,1174,1132,1156]" county="This" municipality="Tolaki" name="Mekongga Mountains" pageId="63" pageNumber="64" stateProvince="Sulawesi Utara">Mekongga Mountains</location>
|
||
of
|
||
<collectingRegion id="8059F83C625E9D2DEAE0FBFDFAFAEE85" box="[1211,1315,1132,1156]" country="Indonesia" name="Sulawesi Utara" pageId="63" pageNumber="64">Sulawesi</collectingRegion>
|
||
, where the
|
||
<typeStatus id="9D26887C625E9D2DEECCFB01FF20EEA9" box="[151,249,1168,1192]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
was collected
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555625E9D2DEE9CFB25FD18EFA5" pageId="63" pageNumber="64" type="materials_examined">
|
||
<paragraph id="422236DE625E9D2DEE9CFB25FD18EFA5" blockId="63.[151,1436,151,1444]" pageId="63" pageNumber="64">
|
||
<materialsCitation id="F2F53C83625E9D2DEE9CFB25FD18EFA5" collectingDate="2010-06-23" collectingDateMax="2010-07-01" collectingDateMin="2010-06-23" collectionCode="MATERIAL, ICBG, MZB, NIH, LIPI" collectorName="Det. C. J. Borkent" country="Indonesia" determinerName="Det. C. J. Borkent" elevation="401" latitude="-3.639444" location="Indonesian Institute of Sciences" longLatPrecision="1" longitude="121.15111" municipality="National Institute of Health" pageId="63" pageNumber="64" specimenCode="CSCA11" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<emphasis id="70E9EACC625E9D2DEE9CFB25FDDDEECC" bold="true" box="[199,516,1204,1229]" pageId="63" pageNumber="64">
|
||
<collectionCode id="248CAE1B625E9D2DEE9CFB25FE86EECC" box="[199,351,1204,1229]" pageId="63" pageNumber="64">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC625E9D2DEC4DFB25FD5FEECD" box="[534,646,1204,1228]" italics="true" pageId="63" pageNumber="64">
|
||
<typeStatus id="9D26887C625E9D2DEC4DFB25FD59EECD" box="[534,640,1204,1228]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
here designated, adult male, pinned with genitalia in plastic vial on pin, labelled “
|
||
<collectingCountry id="3A8A764E625E9D2DEF63FB49FE73EEF1" box="[312,426,1240,1264]" name="Indonesia" pageId="63" pageNumber="64">Indonesia</collectingCountry>
|
||
, se
|
||
<collectingRegion id="8059F83C625E9D2DEF82FB49FD98EEF1" box="[473,577,1240,1264]" country="Indonesia" name="Sulawesi Utara" pageId="63" pageNumber="64">Sulawesi</collectingRegion>
|
||
, North Kolaka / Rante Angin, Tinukari, Mekongga Mt. /
|
||
<date id="3623101E625E9D2DEA87FB49FA4EEEF1" box="[1244,1431,1240,1264]" pageId="63" pageNumber="64" value="2010-06-23" valueMax="2010-07-01" valueMin="2010-06-23">
|
||
<collectingDate id="2667E9F6625E9D2DEA87FB49FA4EEEF1" box="[1244,1431,1240,1264]" pageId="63" pageNumber="64" value="2010-06-23" valueMax="2010-07-01" valueMin="2010-06-23">23.vi–1.vii.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="248CAE1B625E9D2DEECCFB6DFF03EF15" box="[151,218,1276,1300]" pageId="63" pageNumber="64">ICBG</collectionCode>
|
||
team,
|
||
<quantity id="85659B3B625E9D2DEF76FB6DFE41EF15" box="[301,408,1276,1300]" metricMagnitude="2" metricUnit="m" metricValue="4.01" pageId="63" pageNumber="64" unit="m" value="401.0">
|
||
<elevation id="C9B0D1ED625E9D2DEF76FB6DFE41EF15" box="[301,408,1276,1300]" metricMagnitude="2" metricUnit="m" metricValue="4.01" pageId="63" pageNumber="64" unit="m" value="401.0">401 masl</elevation>
|
||
</quantity>
|
||
/
|
||
<geoCoordinate id="27A95019625E9D2DEFEDFB6DFDE4EF15" box="[438,573,1276,1300]" degrees="3.639444" direction="south" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="-3.639444">3.639444°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="27A95019625E9D2DEC17FB6DFD33EF15" box="[588,746,1276,1300]" degrees="121.151111" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="121.15111">121.151111°E</geoCoordinate>
|
||
/
|
||
<specimenCode id="123B9EA5625E9D2DED53FB6DFCB7EF15" box="[776,878,1276,1300]" collectionCode="CSCA" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="63" pageNumber="64" type="Museum">CSCA11</specimenCode>
|
||
L042;
|
||
<typeStatus id="9D26887C625E9D2DEDE0FB6DFB96EF15" box="[955,1103,1276,1300]" pageId="63" pageNumber="64" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD57625E9D2DEA01FB6CFBB6EF15" box="[1114,1135,1277,1300]" pageId="63" pageNumber="64" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="859D4D5D625E9D2DEAD9FB6DFA50EF15" authorityName="Borkent" authorityYear="2012" box="[1154,1417,1276,1300]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tabatius">
|
||
<emphasis id="70E9EACC625E9D2DEAD9FB6DFA50EF15" box="[1154,1417,1276,1300]" italics="true" pageId="63" pageNumber="64">Leptomorphus tabatius</emphasis>
|
||
</taxonomicName>
|
||
/ Borkent,
|
||
<taxonomicNameLabel id="6BDA57B7625E9D2DEF59FAB0FE52EF39" box="[258,395,1312,1336]" pageId="63" pageNumber="64" rank="species">new species</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="B7FEDA1A625E9D2DEFFDFAB0FDB7EF39" box="[422,622,1312,1336]" pageId="63" pageNumber="64">
|
||
<collectorName id="EF685308625E9D2DEFFDFAB0FDB7EF39" box="[422,622,1312,1336]" pageId="63" pageNumber="64">Det. C.J. Borkent</collectorName>
|
||
</determinerName>
|
||
2012” [
|
||
<collectionCode id="248CAE1B625E9D2DEC96FAB0FCC8EF39" box="[717,785,1313,1336]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="63" pageNumber="64" type="Museum">MZB</collectionCode>
|
||
]. The
|
||
<typeStatus id="9D26887C625E9D2DED01FAB1FC65EF39" box="[858,956,1312,1336]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
was collected as part of the International Cooperative Biodiversity Group:
|
||
<collectingCountry id="3A8A764E625E9D2DEC7BFAD5FD54EF5D" box="[544,653,1348,1372]" name="Indonesia" pageId="63" pageNumber="64">Indonesia</collectingCountry>
|
||
study of this biodiversity hotspot. This study was funded by the
|
||
<collectingMunicipality id="A246ACA4625E9D2DEECCFAF9FE17EF81" box="[151,462,1384,1408]" pageId="63" pageNumber="64">National Institute of Health</collectingMunicipality>
|
||
(
|
||
<collectionCode id="248CAE1B625E9D2DEF86FAF9FDCBEF81" box="[477,530,1384,1408]" pageId="63" pageNumber="64">NIH</collectionCode>
|
||
) with the support of the
|
||
<location id="47426005625E9D2DED77FAF9FB4DEF81" LSID="urn:lsid:plazi:treatment:CA3487C8625F9D2DEECCFDFAFD18EFA5:47426005625E9D2DED77FAF9FB4DEF81" box="[812,1172,1384,1408]" country="Indonesia" latitude="-3.639444" longLatPrecision="1" longitude="121.15111" municipality="National Institute of Health" name="Indonesian Institute of Sciences" pageId="63" pageNumber="64" stateProvince="Sulawesi Utara">Indonesian Institute of Sciences</location>
|
||
(
|
||
<collectionCode id="248CAE1B625E9D2DEAFFFAF9FB04EF81" box="[1188,1245,1384,1408]" pageId="63" pageNumber="64">LIPI</collectionCode>
|
||
). We thank them for providing us with this specimen for our study.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |