treatments-xml/data/CA/34/87/CA3487C8625E9D52EECCFA7CFE4DED95.xml

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<mods:title id="71D9497AA182024A27425CFAEF4A9D1C">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<subSubSection id="0A876555625E9D2DEECCFA7CFD2EEC06" box="[151,759,1517,1543]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="422236DE625E9D2DEECCFA7CFD2EEC06" blockId="63.[151,759,1517,1576]" box="[151,759,1517,1543]" pageId="63" pageNumber="64">
<heading id="196A81B2625E9D2DEECCFA7CFD2EEC06" bold="true" box="[151,759,1517,1543]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
<emphasis id="70E9EACC625E9D2DEECCFA7CFD2EEC06" bold="true" box="[151,759,1517,1543]" pageId="63" pageNumber="64">
33. 
<taxonomicName id="859D4D5D625E9D2DEE98FA7CFD82EC06" ID-CoL="8CJ4J" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[195,603,1517,1543]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tagbanua" status="sp. nov.">
<emphasis id="70E9EACC625E9D2DEE98FA7CFE32EC06" bold="true" box="[195,491,1517,1543]" italics="true" pageId="63" pageNumber="64">Leptomorphus tagbanua</emphasis>
Borkent
</taxonomicName>
, 
<taxonomicNameLabel id="6BDA57B7625E9D2DEC33FA7CFD2EEC06" box="[616,759,1517,1543]" pageId="63" pageNumber="64" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A876555625E9D2DEECCF981FD8AEC29" box="[151,595,1552,1576]" pageId="63" pageNumber="64" type="description">
<paragraph id="422236DE625E9D2DEECCF981FD8AEC29" blockId="63.[151,759,1517,1576]" box="[151,595,1552,1576]" pageId="63" pageNumber="64">
(
<figureCitation id="DAA62A5B625E9D2DEEC5F981FEC0EC29" box="[158,281,1552,1576]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="63" pageNumber="64">Figures 40</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEF7EF981FE9AEC29" box="[293,323,1552,1576]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="63" pageNumber="64">74</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEF14F981FEB4EC29" box="[335,365,1552,1576]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="63" pageNumber="64">88</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEF22F981FE7DEC29" box="[377,420,1552,1576]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev &amp; Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="63" pageNumber="64">119</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEFEBF981FE05EC29" box="[432,476,1552,1576]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp &amp; Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="63" pageNumber="64">137</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEFB3F981FDCDEC29" box="[488,532,1552,1576]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values&gt;50% are shown in bold below the branches." pageId="63" pageNumber="64">150</figureCitation>
, 
<figureCitation id="DAA62A5B625E9D2DEC7BF981FD92EC29" box="[544,587,1552,1576]" captionStart="FIGURE 152" captionStartId="114.[151,250,1784,1807]" captionTargetBox="[162,1422,1059,1758]" captionTargetId="figure-112@114.[156,1428,1046,1779]" captionTargetPageId="114" captionText="FIGURE 152. One of three equally parsimonious trees showing the relationships between species in the L. ornatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="63" pageNumber="64">152</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0A876555625E9D2DEECCF9C9FB93ED49" pageId="63" pageNumber="64" type="diagnosis">
<paragraph id="422236DE625E9D2DEECCF9C9FE9FECDD" blockId="63.[151,1437,1624,2008]" pageId="63" pageNumber="64">
<emphasis id="70E9EACC625E9D2DEECCF9C9FEEDEC70" bold="true" box="[151,308,1624,1649]" pageId="63" pageNumber="64">DIAGNOSIS</emphasis>
: The only extant species of 
<taxonomicName id="859D4D5D625E9D2DECEAF9C9FC8AEC71" authorityName=": Edwards" authorityYear="1925" box="[689,851,1624,1648]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC625E9D2DECEAF9C9FC8AEC71" box="[689,851,1624,1648]" italics="true" pageId="63" pageNumber="64">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: laterotergite and anepisternum brown; wing without apical spot; vein sc-r joining R at ~2X its own length from origin of Rs; segments 3–5 of abdomen noticeably swollen relative to other segments; male genitalia wider than long (
<figureCitation id="DAA62A5B625E9D2DEE83F955FEE0ECDD" box="[216,313,1732,1756]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev &amp; Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="63" pageNumber="64">Fig. 119</figureCitation>
).
</paragraph>
<paragraph id="422236DE625E9D2DEE9CF979FB93ED49" blockId="63.[151,1437,1624,2008]" pageId="63" pageNumber="64">
This species can be distinguished from the other 
<collectingRegion id="8059F83C625E9D2DECAEF979FC89ED01" box="[757,848,1768,1792]" country="Republic of the Congo" name="Orientale" pageId="63" pageNumber="64">Oriental</collectingRegion>
and eastern Palaearctic species either by the brown katepisternum (
<figureCitation id="DAA62A5B625E9D2DEF12F89DFE46ED25" box="[329,415,1804,1828]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="63" pageNumber="64">Fig. 40</figureCitation>
, yellow in 
<emphasis id="70E9EACC625E9D2DEC7CF89CFCCBED25" box="[551,786,1804,1828]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="859D4D5D625E9D2DEC7CF89CFD52ED25" authorityName=": Matile" authorityYear="1977" box="[551,651,1804,1828]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="babai">L. babai</taxonomicName>
, 
<taxonomicName id="859D4D5D625E9D2DECC1F89CFCCBED25" authorityName=": Matile" authorityYear="1977" box="[666,786,1805,1828]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ornatus">L. ornatus</taxonomicName>
</emphasis>
and 
<taxonomicName id="859D4D5D625E9D2DED0BF89CFBC8ED25" authorityName="Borkent" authorityYear="2012" box="[848,1041,1805,1828]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="titiwangsensis">
<emphasis id="70E9EACC625E9D2DED0BF89CFBC8ED25" box="[848,1041,1805,1828]" italics="true" pageId="63" pageNumber="64">L. titiwangsensis</emphasis>
</taxonomicName>
), or by the wider than long male genitalia with uniquely shaped tergal evaginations at the posterior margin (
<figureCitation id="DAA62A5B625E9D2DED87F8A1FBE7ED49" box="[988,1086,1840,1864]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev &amp; Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="63" pageNumber="64">Fig. 119</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A876555625E9D52EE9CF8C5FE1BEC54" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="description">
<paragraph id="422236DE625E9D52EE9CF8C5FE53EC0C" blockId="63.[151,1437,1624,2008]" lastBlockId="64.[151,1437,151,1940]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<emphasis id="70E9EACC625E9D2DEE9CF8C5FE50ED6C" bold="true" box="[199,393,1876,1901]" pageId="63" pageNumber="64">DESCRIPTION</emphasis>
: 
<emphasis id="70E9EACC625E9D2DEFC0F8C5FE0DED6D" box="[411,468,1876,1900]" italics="true" pageId="63" pageNumber="64">Male</emphasis>
. (
<figureCitation id="DAA62A5B625E9D2DEFB5F8C5FD9AED6D" box="[494,579,1876,1900]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="63" pageNumber="64">Fig. 40</figureCitation>
) 
<emphasis id="70E9EACC625E9D2DEC0EF8C5FD4CED6D" bold="true" box="[597,661,1876,1900]" pageId="63" pageNumber="64">Head</emphasis>
: yellow, taller than wide in anterior view. Antenna with basal 2–3 flagellomeres lighter brown, darkening apically; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third, entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 2 large bristles, few of setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base yellow remainder yellow; flagellomere 6 1.0X as long as broad. Clypeus light brown, slightly laterally compressed oval; bristles on clypeus light brown, 4 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 1.5X as long as face. Face light brown; shape a just longer than wide triangle, with few bristles ventrolaterally. Frons yellow; bare, frontal furrow running 3/4 distance from dorsal margin towards ventral margin, frontal cleft more than 1X diameter of median ocellus anterior of median ocellus. Palpus yellow; segments increasing in length, segment 3 appears spherical (difficult to tell in specimen). Labellum yellow. Eye with inter-ommatidial setulae absent. Occiput yellow with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black. 
<emphasis id="70E9EACC62219D52EDB8FE29FBE5EBD1" bold="true" box="[995,1084,440,464]" pageId="64" pageNumber="65">Thorax</emphasis>
: Length 
<quantity id="85659B3B62219D52EAFBFE26FADAEBCE" box="[1184,1283,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="64" pageNumber="65" unit="mm" value="1.03">1.03 mm</quantity>
(n = 1). Dark brown. Scutum dark brown with blue-green specks; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; two thirds row of lateral setae present; patch of setae on scutum at wing base present. Scutellum yellow; with 6–8 large bristles and few small bristles. Prescutum brown. Mediotergite dark brown with 6 bristles on posterolateral corners, few medially. Laterotergite brown; anterior margin of laterotergite not reaching katepisternum. Anepimeron brown. Anepisternum dark brown. Katepisternum dark brown. Antepronotum and proepisternum dark brown. Margin of anterior and posterior spiracles yellow with light brown trichia. Metepisternum anterior half brown posterior yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown. 
<emphasis id="70E9EACC62219D52EA87FD49FACDE8F1" bold="true" box="[1244,1300,728,752]" pageId="64" pageNumber="65">Legs</emphasis>
: principally yellow; extreme anteroapical corner light brown on all femora; tarsi brown. Midfemur without apical spine-like process. Tibia with covering of yellow macrotrichia, foretibia without comb of short setae along length of anteroventral surface (
<figureCitation id="DAA62A5B62219D52EFC8FCD5FE3EE95D" box="[403,487,836,860]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="64" pageNumber="65">Fig. 88</figureCitation>
), tibial spurs yellow, foretibial spur length 2X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 3/4 of its length, placed basally, shortest midtibial spur subequal to length of longest, longest midtibial spur 3.5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 3X apical thickness of hind tibia. Foreleg first tarsomere 1.6X length of foretibia. 
<emphasis id="70E9EACC62219D52EF50FC42FE95E9ED" bold="true" box="[267,332,979,1004]" pageId="64" pageNumber="65">Wing</emphasis>
(
<figureCitation id="DAA62A5B62219D52EF38FC45FE66E9ED" box="[355,447,980,1004]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="64" pageNumber="65">Fig. 74</figureCitation>
): Length 4.0 mm (n = 1). Hyaline; apical macula absent; medial macula absent. Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present, apical end joining R at 2X its own length prior to origin of Rs. R 
<subScript id="DE19349B62219D52ECDBFBDEFD51EE5C" attach="left" box="[640,648,1103,1117]" fontSize="6" pageId="64" pageNumber="65">4</subScript>
absent. R 
<subScript id="DE19349B62219D52ECA8FBDEFD22EE5C" attach="left" box="[755,763,1103,1117]" fontSize="6" pageId="64" pageNumber="65">5</subScript>
straight, slight posterior turn near tip. M 
<subScript id="DE19349B62219D52EA99FBDEFB13EE5C" attach="left" box="[1218,1226,1103,1117]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
reaching apex just before R 
<subScript id="DE19349B62219D52EEA2FBE7FED8EE85" attach="left" box="[249,257,1142,1156]" fontSize="6" pageId="64" pageNumber="65">5</subScript>
, apices of M veins fading before wing margin. M 
<subScript id="DE19349B62219D52ED68FBE7FCE2EE85" attach="left" box="[819,827,1142,1156]" fontSize="6" pageId="64" pageNumber="65">4</subScript>
-CuA fork arising apically of origin of r-m. A 
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absent. 
<emphasis id="70E9EACC62219D52EECCFB1CFED5EEA7" bold="true" box="[151,268,1165,1190]" pageId="64" pageNumber="65">Abdomen</emphasis>
: Abdominal segments 3–5 swollen relative to other segments; principally dark brown, T3–5 with anterior 1/3–1/2 yellow. Tergite 8 smaller than other abdominal sclerites and 2 rows of bristles on posterior margin and large patches (~30) laterally. 
<emphasis id="70E9EACC62219D52EC47FB44FD53EEEF" bold="true" box="[540,650,1237,1262]" pageId="64" pageNumber="65">Genitalia</emphasis>
(
<figureCitation id="DAA62A5B62219D52ECC7FB44FCDBEEEC" box="[668,770,1237,1261]" captionStart="FIGURES 117–120" captionStartId="104.[151,263,1874,1897]" captionTargetBox="[189,1396,184,1835]" captionTargetId="figure-113@104.[145,1430,173,1883]" captionTargetPageId="104" captionText="FIGURES 117–120. Leptomorphus male genitalia. 117. L. subcaeruleus, ventral. 118. L. subforcipatus (reprinted with permission from Zaitzev &amp; Ševčík 2002), A. dorsal. B. gonocoxite and gonostylus, ventral. 119. L. tagbanua, ventral. 120. L. tabatius, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="64" pageNumber="65">Fig. 119</figureCitation>
): orangish yellow. Sternite 9 reduced to long thin sclerite attached to aedeagus. Tergite 9 wider than long oval, posterior margin with a shallow medial indentation and an intricate strongly sclerotized process with many points on lateral corner. Gonocoxite strongly fused with T9 but visible due to difference in sclerotization, placed centrally on T9, medial margin not reaching median line, bearing gonostylus apically. Gonostylus a single broad-based lobe tapering to apex which is bifurcated into two short sclerotized points, gonocoxite III fused to gonostylus basally but forming its own lobe. Aedeagus ~1/2 length of genitalia, broad, rounded base tapering to apex, strongly united basally with S9, apodemes highly reduced. Parameres squared off so essentially only apodemes remain (posterior margin flat), apodemes strongly united with gonocoxal apodemes.
</paragraph>
<paragraph id="422236DE62219D52EE9CF988FE06EC30" blockId="64.[151,1437,151,1940]" box="[199,479,1561,1585]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9CF988FE85EC30" box="[199,348,1561,1585]" italics="true" pageId="64" pageNumber="65">Female adult</emphasis>
. Unknown.
</paragraph>
<paragraph id="422236DE62219D52EE9CF9AFFE1BEC54" blockId="64.[151,1437,151,1940]" box="[199,450,1597,1621]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9CF9AFFEE7EC54" box="[199,318,1598,1621]" italics="true" pageId="64" pageNumber="65">Immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562219D52EE9CF9F1FE16EC79" box="[199,463,1632,1657]" pageId="64" pageNumber="65" type="biology_ecology">
<paragraph id="422236DE62219D52EE9CF9F1FE16EC79" blockId="64.[151,1437,151,1940]" box="[199,463,1632,1657]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9CF9F1FE93EC78" bold="true" box="[199,330,1632,1657]" pageId="64" pageNumber="65">BIOLOGY</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562219D52EE9CF912FC62EC9D" box="[199,955,1667,1692]" pageId="64" pageNumber="65" type="distribution">
<paragraph id="422236DE62219D52EE9CF912FC62EC9D" blockId="64.[151,1437,151,1940]" box="[199,955,1667,1692]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9CF912FE4FEC9D" bold="true" box="[199,406,1667,1692]" pageId="64" pageNumber="65">DISTRIBUTION</emphasis>
: Coron Island, 
<collectingCountry id="3A8A764E62219D52EC10F915FD10EC9D" box="[587,713,1668,1692]" name="Philippines" pageId="64" pageNumber="65">Philippines</collectingCountry>
(
<figureCitation id="DAA62A5B62219D52EC83F915FCE0EC9D" box="[728,825,1668,1692]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp &amp; Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="64" pageNumber="65">Fig. 137</figureCitation>
), 
<quantity id="85659B3B62219D52ED15F915FC6EEC9D" box="[846,951,1668,1692]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="64" pageNumber="65" unit="m" value="215.0">215 masl</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562219D52EE9CF936FA4FECE2" pageId="64" pageNumber="65" type="discussion">
<paragraph id="422236DE62219D52EE9CF936FA4FECE2" blockId="64.[151,1437,151,1940]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9CF936FEABECC1" bold="true" box="[199,370,1703,1728]" pageId="64" pageNumber="65">DISCUSSION</emphasis>
: This species was compared to the descriptions and figures of the recently described 
<collectingRegion id="8059F83C62219D52EB1BF936FA45ECBE" box="[1344,1436,1703,1727]" country="Republic of the Congo" name="Orientale" pageId="64" pageNumber="65">Oriental</collectingRegion>
and Australasian species (
<bibRefCitation id="260C4B2F62219D52EFECF95AFD7BECE2" author="Papp, L. &amp; Sevcik, J." box="[439,674,1739,1763]" pageId="64" pageNumber="65" pagination="139 - 159" refId="ref95170" refString="Papp, L. &amp; Sevcik, J. (2011) Eight new oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientarum Hungaricae, 57, 139 - 159." type="journal article" year="2011">Papp &amp; Ševčík 2011</bibRefCitation>
), and is clearly different from all of them and not a junior synonym.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562219D52EE9DF97CFD45ED28" pageId="64" pageNumber="65" type="etymology">
<paragraph id="422236DE62219D52EE9DF97CFD45ED28" blockId="64.[151,1437,151,1940]" pageId="64" pageNumber="65">
<emphasis id="70E9EACC62219D52EE9DF97CFE59ED07" bold="true" box="[198,384,1773,1798]" pageId="64" pageNumber="65">ETYMOLOGY</emphasis>
: This species is named for the Tagbanua people, who are the ancestral and current inhabitants of Coron Island where this species was found.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562219D52EE9CF8A4FE4DED95" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="422236DE62219D52EE9CF8A4FE4DED95" blockId="64.[151,1437,151,1940]" pageId="64" pageNumber="65">
<materialsCitation id="F2F53C8362219D52EE9CF8A4FE4DED95" collectingDate="1945-06-21" collectionCode="MATERIAL, ISUI" collectorName="Coll. J. Laffoon &amp; Det. C. J. Borkent" determinerName="Det. C. J. Borkent" pageId="64" pageNumber="65" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="70E9EACC62219D52EE9CF8A4FDD9ED4F" bold="true" box="[199,512,1845,1870]" pageId="64" pageNumber="65">
<collectionCode id="248CAE1B62219D52EE9CF8A4FE86ED4F" box="[199,351,1845,1870]" pageId="64" pageNumber="65">MATERIAL</collectionCode>
EXAMINED
</emphasis>
: 
<emphasis id="70E9EACC62219D52EC54F8A4FDA6ED4C" box="[527,639,1845,1869]" italics="true" pageId="64" pageNumber="65">
<typeStatus id="9D26887C62219D52EC54F8A4FDA0ED4C" box="[527,633,1845,1869]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
:
</emphasis>
here designated, adult male, pinned, labelled “Coron,/ Busuanga,/ Phil. Is.; 
<collectorName id="EF68530862219D52EE9EF8C9FEA4ED71" box="[197,381,1880,1904]" pageId="64" pageNumber="65">Coll. J. Laffoon</collectorName>
/ 
<date id="3623101E62219D52EFCBF8C9FDCEED71" box="[400,535,1880,1904]" pageId="64" pageNumber="65" value="1945-06-21">
<collectingDate id="2667E9F662219D52EFCBF8C9FDCEED71" box="[400,535,1880,1904]" pageId="64" pageNumber="65" value="1945-06-21">VI-21-1945</collectingDate>
</date>
; 
<typeStatus id="9D26887C62219D52EC7CF8C9FD62ED71" box="[551,699,1880,1904]" pageId="64" pageNumber="65" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="549BFD5762219D52EC9DF8C8FD02ED71" box="[710,731,1881,1904]" pageId="64" pageNumber="65" type="male">♂</specimenCount>
/ 
<taxonomicName id="859D4D5D62219D52ECB6F8C8FBDAED70" authorityName="Borkent" authorityYear="2012" box="[749,1027,1881,1905]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tagbanua">
<emphasis id="70E9EACC62219D52ECB6F8C8FBDAED70" box="[749,1027,1881,1905]" italics="true" pageId="64" pageNumber="65">Leptomorphus tagbanua</emphasis>
</taxonomicName>
/ Borkent, 
<taxonomicNameLabel id="6BDA57B762219D52EAD6F8C8FAC1ED71" box="[1165,1304,1880,1904]" pageId="64" pageNumber="65" rank="species">new species</taxonomicNameLabel>
/ 
<determinerName id="B7FEDA1A62219D52EB6DF8C8FF28ED95" pageId="64" pageNumber="65">
<collectorName id="EF68530862219D52EB6DF8C8FF28ED95" pageId="64" pageNumber="65">Det. C.J. Borkent</collectorName>
</determinerName>
2012” [
<collectionCode id="248CAE1B62219D52EF10F8EDFE50ED95" box="[331,393,1916,1940]" country="USA" httpUri="http://grbio.org/cool/j5ep-a4qz" name="Iowa State University" pageId="64" pageNumber="65">ISUI</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>