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<mods:title id="3F4F019F923B9D5E0D727079460B3631">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<subSubSection id="0A87655562539D20EECCFF06FDE6EAB0" box="[151,575,151,177]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="422236DE62539D20EECCFF06FDE6EAB0" blockId="50.[151,575,151,210]" box="[151,575,151,177]" pageId="50" pageNumber="51">
<heading id="196A81B262539D20EECCFF06FDE6EAB0" bold="true" box="[151,575,151,177]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
<emphasis id="70E9EACC62539D20EECCFF06FDE6EAB0" bold="true" box="[151,575,151,177]" pageId="50" pageNumber="51">
24.
<taxonomicName id="859D4D5D62539D20EE98FF06FDE6EAB0" ID-CoL="8CJ3N" authority="Matile" authorityName="Matile" box="[195,575,151,177]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="70E9EACC62539D20EE98FF06FE3EEAB0" bold="true" box="[195,487,151,177]" italics="true" pageId="50" pageNumber="51">Leptomorphus obscurus</emphasis>
Matile
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A87655562539D20EECCFF2BFDF0EAD3" box="[151,553,186,210]" pageId="50" pageNumber="51" type="description">
<paragraph id="422236DE62539D20EECCFF2BFDF0EAD3" blockId="50.[151,575,151,210]" box="[151,553,186,210]" pageId="50" pageNumber="51">
(
<figureCitation id="DAA62A5B62539D20EEC5FF2BFEC0EAD3" box="[158,281,186,210]" captionStart="FIGURES 2429" captionStartId="88.[151,263,1982,2005]" captionTargetBox="[153,1435,183,1959]" captionTargetId="figure-54@88.[151,1436,181,1961]" captionTargetPageId="88" captionText="FIGURES 2429. Leptomorphus habitus. 24. L. nebulosus, female, dark form. 25. L. nebulosus, male, dark form. 26. L. nebulosus, female, common colour. 27. L. nebulosus, male, light form. 28. L. obscurus, male. 29. L. neivai, male." pageId="50" pageNumber="51">Figures 28</figureCitation>
,
<figureCitation id="DAA62A5B62539D20EF7EFF2BFE9AEAD3" box="[293,323,186,210]" captionStart="FIGURES 5669" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 5669. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="50" pageNumber="51">66</figureCitation>
,
<figureCitation id="DAA62A5B62539D20EF14FF2BFEA3EAD3" box="[335,378,186,210]" captionStart="FIGURES 113116" captionStartId="103.[151,263,1908,1931]" captionTargetBox="[170,1416,182,1897]" captionTargetId="figure-91@103.[143,1428,179,1924]" captionTargetPageId="103" captionText="FIGURES 113116. Leptomorphus male genitalia, ventral. 113. L. obscurus. 114. L. panorpiformis. 115. L. stigmatus. 116. L. quadrimaculatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="50" pageNumber="51">113</figureCitation>
,
<figureCitation id="DAA62A5B62539D20EFDDFF2BFE6BEAD3" box="[390,434,186,210]" captionStart="FIGURE 134136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="50" pageNumber="51">136</figureCitation>
,
<figureCitation id="DAA62A5B62539D20EFE5FF2BFE33EAD3" box="[446,490,186,210]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values&gt;50% are shown in bold below the branches." pageId="50" pageNumber="51">150</figureCitation>
,
<figureCitation id="DAA62A5B62539D20EFADFF2BFDF8EAD3" box="[502,545,186,210]" captionStart="FIGURE 153" captionStartId="115.[151,250,801,824]" captionTargetBox="[155,1364,186,781]" captionTargetId="figure-100@115.[151,1436,161,836]" captionTargetPageId="115" captionText="FIGURE 153. One of three equally parsimonious trees showing the relationships between species in the L. grjebinei species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="50" pageNumber="51">153</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0A87655562539D20EECCFE93FE35EB80" pageId="50" pageNumber="51" type="reference_group">
<paragraph id="422236DE62539D20EECCFE93FD2DEB19" blockId="50.[151,756,258,280]" box="[151,756,258,280]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="628D11F062539D20EECCFE93FD2DEB19" box="[151,756,258,280]" pageId="50" pageNumber="51">
<treatmentCitation id="C33C10CF62539D20EECCFE93FD29EB19" author="Matile, L." box="[151,752,258,280]" page="152" pageId="50" pageNumber="51" year="1977">
<taxonomicName id="859D4D5D62539D20EECCFE93FD29EB19" ID-CoL="8CJ3N" authority="Matile, 1977: 152" authorityName="Matile" authorityPageNumber="152" authorityYear="1977" box="[151,752,258,280]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC62539D20EECCFE93FDEBEB19" box="[151,562,258,280]" italics="true" pageId="50" pageNumber="51">Leptomorphus (Gymnoscutum) obscurus</emphasis>
<bibRefCitation id="260C4B2F62539D20EC62FE93FD29EB19" author="Matile, L." box="[569,752,258,280]" pageId="50" pageNumber="51" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 152</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="422236DE62539D20EECCFED4FE35EB80" blockId="50.[151,1437,325,2016]" pageId="50" pageNumber="51">
<emphasis id="70E9EACC62539D20EECCFED4FEC0EB5F" bold="true" box="[151,281,325,350]" pageId="50" pageNumber="51">References</emphasis>
:
<bibRefCitation id="260C4B2F62539D20EF6EFED4FDE4EB5C" author="Crosskey, R. W." box="[309,573,325,349]" pageId="50" pageNumber="51" refId="ref92665" refString="Crosskey, R. W. (ed.) (1980) Catalogue of the Diptera of the Afrotropical Region. British Museum of Natural History, London, 1437 pp." type="book" year="1980">Crosskey 1980: 1221</bibRefCitation>
(catalogue appendix);
<bibRefCitation id="260C4B2F62539D20ED3DFED4FBA1EB5C" author="Matile, L." box="[870,1144,325,349]" pageId="50" pageNumber="51" pagination="143 - 150" refId="ref94722" refString="Matile, L. (1997) Nouvelles donnees sur les Leptomorphus Afrotropicaux (Diptera, Mycetophilidae). Revue Francaise d'Entomologie (Nouvelle Serie) 1996, 18 (4), 143 - 150." type="journal article" year="1997">Matile 1997: 147149</bibRefCitation>
(figures, new records, morphological variation, key).
</paragraph>
</subSubSection>
<subSubSection id="0A87655562539D20EE9CFE1CFDA7E8C4" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="422236DE62539D20EE9CFE1CFB55E834" blockId="50.[151,1437,325,2016]" pageId="50" pageNumber="51">
<emphasis id="70E9EACC62539D20EE9CFE1CFEBDEBA7" bold="true" box="[199,356,397,422]" pageId="50" pageNumber="51">DIAGNOSIS</emphasis>
: The only extant species of
<taxonomicName id="859D4D5D62539D20EC92FE1CFCB2EBA4" authorityName=": Edwards" authorityYear="1925" box="[713,875,397,421]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC62539D20EC92FE1CFCB2EBA4" box="[713,875,397,421]" italics="true" pageId="50" pageNumber="51">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: laterotergite, anepisternum and anepimeron yellow; scutellum brown, male genitalia with tergite 9 gradually tapering on apical quarter (tergal evagination), apex bearing medial spur and often a secondary small bump or point, also with strong, ventrally-produced, thin ridge running across ventral surface at the anterior margin of the tergal evagination, creating a semicircular dorsal margin when viewed caudally (
<figureCitation id="DAA62A5B62539D20EA45FD8CFB59E834" box="[1054,1152,541,565]" captionStart="FIGURES 113116" captionStartId="103.[151,263,1908,1931]" captionTargetBox="[170,1416,182,1897]" captionTargetId="figure-91@103.[143,1428,179,1924]" captionTargetPageId="103" captionText="FIGURES 113116. Leptomorphus male genitalia, ventral. 113. L. obscurus. 114. L. panorpiformis. 115. L. stigmatus. 116. L. quadrimaculatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="50" pageNumber="51">Fig. 113</figureCitation>
).
</paragraph>
<paragraph id="422236DE62539D20EE9CFDD0FDA7E8C4" blockId="50.[151,1437,325,2016]" pageId="50" pageNumber="51">
This species can be distinguished from most other Afrotropical species by the brown scutellum and lack of small setae covering the scutum (except along dorsocentral lines). It can be separated from
<taxonomicName id="859D4D5D62539D20EA9AFDF7FA91E87C" authorityName="Matile, 1980:" authorityYear="1229" box="[1217,1352,613,637]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="crosskeyi">
<emphasis id="70E9EACC62539D20EA9AFDF7FA91E87C" box="[1217,1352,613,637]" italics="true" pageId="50" pageNumber="51">L. crosskeyi</emphasis>
</taxonomicName>
and
<taxonomicName id="859D4D5D62539D20EBDEFDF7FF25E8A0" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="70E9EACC62539D20EBDEFDF7FF25E8A0" italics="true" pageId="50" pageNumber="51">L. obscurus</emphasis>
</taxonomicName>
by the gradually tapering shape of the tergal evaginations (
<figureCitation id="DAA62A5B62539D20ED91FD18FBF7E8A0" box="[970,1070,649,673]" captionStart="FIGURES 113116" captionStartId="103.[151,263,1908,1931]" captionTargetBox="[170,1416,182,1897]" captionTargetId="figure-91@103.[143,1428,179,1924]" captionTargetPageId="103" captionText="FIGURES 113116. Leptomorphus male genitalia, ventral. 113. L. obscurus. 114. L. panorpiformis. 115. L. stigmatus. 116. L. quadrimaculatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="50" pageNumber="51">Fig. 113</figureCitation>
) rather than the almost square posterior margin in these two other species.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562539D21EE9CFD40FE1BE85E" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="422236DE62539D21EE9CFD40FECBEBF2" blockId="50.[151,1437,325,2016]" lastBlockId="51.[151,1436,151,1292]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="70E9EACC62539D20EE9CFD40FE50E8EB" bold="true" box="[199,393,721,746]" pageId="50" pageNumber="51">DESCRIPTION</emphasis>
:
<emphasis id="70E9EACC62539D20EFC0FD40FE0DE8E8" box="[411,468,721,745]" italics="true" pageId="50" pageNumber="51">Male</emphasis>
. (
<figureCitation id="DAA62A5B62539D20EFB6FD40FD9BE8E8" box="[493,578,721,745]" captionStart="FIGURES 2429" captionStartId="88.[151,263,1982,2005]" captionTargetBox="[153,1435,183,1959]" captionTargetId="figure-54@88.[151,1436,181,1961]" captionTargetPageId="88" captionText="FIGURES 2429. Leptomorphus habitus. 24. L. nebulosus, female, dark form. 25. L. nebulosus, male, dark form. 26. L. nebulosus, female, common colour. 27. L. nebulosus, male, light form. 28. L. obscurus, male. 29. L. neivai, male." pageId="50" pageNumber="51">Fig. 28</figureCitation>
)
<emphasis id="70E9EACC62539D20EC0FFD40FD4DE8E8" bold="true" box="[596,660,721,745]" pageId="50" pageNumber="51">Head</emphasis>
: yellow with some brown spots, circular in anterior view. Antenna brown; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae absent; pedicel light brown/yellow, with 2 large bristles, several setae on apicodorsal margin, a few fine setae on apicoventral margin; flagellomere 1 with tapered base either brown or yellow remainder brown; flagellomere 6 1.1X as long as broad. Clypeus yellow, slightly laterally compressed oval; bristles on clypeus brown, 46 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face yellow; shape a just longer than wide triangle, with few bristles ventrolaterally. Frons yellow; with few bristles medioventrally, frontal furrow running 1/2 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus with 13 yellow, remainder brown; segment 1 hidden behind eye, segments increasing in length, segment 5 2X length of segment 4 with central half thinner than base and apex, segment 3 with apicolateral patch of fine yellow setae encircled by strong dark setae. Labellum light brown. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with some adventitious brown spotting anteriorly, with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black with electric blue green specks.
<emphasis id="70E9EACC62539D20EF9CFB7CFDC6EF04" bold="true" box="[455,543,1261,1285]" pageId="50" pageNumber="51">Thorax</emphasis>
: Length 1.08 ±
<quantity id="85659B3B62539D20EC8FFB7CFCE1EF04" box="[724,824,1261,1285]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="50" pageNumber="51" unit="mm" value="0.14">0.14 mm</quantity>
(
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, n = 10). Dark brown dorsally, yellow laterally. Scutum dark brown/black with blue-green specks, pair of yellow mediolateral and smaller posterolateral spots; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; single row of lateral setae present; patch of setae on scutum at wing base present. Scutellum brown; with 810 large bristles and many small bristles. Prescutum yellow. Mediotergite dark brown with 68 bristles on posterolateral corners, anteromedial patch of small bristles. Laterotergite yellow; anterior margin of laterotergite not reaching katepisternum. Anepimeron yellow. Anepisternum yellow. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with light brown trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown.
<emphasis id="70E9EACC62539D20EFB7F9A0FDFDEC48" bold="true" box="[492,548,1585,1609]" pageId="50" pageNumber="51">Legs</emphasis>
: principally yellow; hind femur light brown at very apex; extreme anteroapical corner dark brown on all femora. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, hind tibial spur yellow, remainder brown, foretibial spur length 2X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 1/3 of its length, placed basally, shortest midtibial spur 0.8X length of longest, longest midtibial spur 5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 4X apical thickness of hind tibia. Foreleg first tarsomere 1.8X length of foretibia.
<emphasis id="70E9EACC62539D20EA71F898FBB2ED23" bold="true" box="[1066,1131,1801,1826]" pageId="50" pageNumber="51">Wing</emphasis>
(
<figureCitation id="DAA62A5B62539D20EA20F898FB16ED20" box="[1147,1231,1801,1825]" captionStart="FIGURES 5669" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 5669. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="50" pageNumber="51">Fig. 66</figureCitation>
): Length 5.1 ±
<quantity id="85659B3B62539D20EB22F898FF1BED44" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="50" pageNumber="51" unit="mm" value="0.7">0.7 mm</quantity>
(
<quantity id="85659B3B62539D20EE8CF8BCFEB0ED44" box="[215,361,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.7" metricValueMin="4.5" pageId="50" pageNumber="51" unit="mm" value="5.1" valueMax="5.7" valueMin="4.5">4.55.7 mm</quantity>
, n = 10). Hyaline; apical macula absent or very light at R
<subScript id="DE19349B62539D20EA6DF8ADFBE7ED4B" attach="left" box="[1078,1086,1852,1866]" fontSize="6" pageId="50" pageNumber="51">5</subScript>
apex; medial macula absent. Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with group of macrotrichia. Vein sc-r present, apical end joining R at 2X its own length prior to origin of Rs. R
<subScript id="DE19349B62539D20ECB6F83AFD2CEDB8" attach="left" box="[749,757,1963,1977]" fontSize="6" pageId="50" pageNumber="51">4</subScript>
absent. R
<subScript id="DE19349B62539D20ED3BF83AFCB1EDB8" attach="left" box="[864,872,1963,1977]" fontSize="6" pageId="50" pageNumber="51">5</subScript>
straight, slight posterior turn near tip. M
<subScript id="DE19349B62539D20EB76F83AFAECEDB8" attach="left" box="[1325,1333,1963,1977]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
reaching apex before R
<subScript id="DE19349B62539D20EF6FF843FEE5EDE1" attach="left" box="[308,316,2002,2016]" fontSize="6" pageId="50" pageNumber="51">5</subScript>
, apices of M veins fading before wing margin. M
<subScript id="DE19349B62539D20ED37F843FCADEDE1" attach="right" box="[876,884,2002,2016]" fontSize="6" pageId="50" pageNumber="51">4</subScript>
-CuA fork arising before origin of r-m. A
<subScript id="DE19349B62539D20EB1FF843FA95EDE1" attach="left" box="[1348,1356,2002,2016]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
faintly present as crease.
<emphasis id="70E9EACC62529D21EF32FF06FE07EAB1" bold="true" box="[361,478,151,176]" pageId="51" pageNumber="52">Abdomen</emphasis>
: Tergites principally dark brown, T26 with lateral yellow spots. Tergite 8 smaller than all other abdominal sclerites, with 23 bristles on each apicolateral corner.
<emphasis id="70E9EACC62529D21EA11FF2AFB60EAD5" bold="true" box="[1098,1209,187,212]" pageId="51" pageNumber="52">Genitalia</emphasis>
(
<figureCitation id="DAA62A5B62529D21EA97FF2DFAEBEAD5" box="[1228,1330,188,212]" captionStart="FIGURES 113116" captionStartId="103.[151,263,1908,1931]" captionTargetBox="[170,1416,182,1897]" captionTargetId="figure-91@103.[143,1428,179,1924]" captionTargetPageId="103" captionText="FIGURES 113116. Leptomorphus male genitalia, ventral. 113. L. obscurus. 114. L. panorpiformis. 115. L. stigmatus. 116. L. quadrimaculatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="51" pageNumber="52">Fig. 113</figureCitation>
): yellow. Sternite 9 sclerotized, oval/rounded diamond, 3/5 the width of genitalia at widest point, overlapping medial margin of gonoxite. Tergite 9 longer than wide, lateral margins widening slightly to apex and then tapering sharply into pointed lobes with medial U-shaped indentation, a ventrally extending sclerotized band running along the base of posterior lobes. Gonocoxite placed basally on T9, medial margin just not reaching medial line, bearing gonostylus apically. Gonostylus with two lobes, dorsal lobe a broad based point, ventral lobe half the width of dorsal but 2.5X length and barely tapering until apex, gonocoxite III fused to dorsolateral margin. Aedeagus 2/3 length of gonocoxite, tapering towards apex, apodemes 2/5 total length. Parameres a simple taper, apodemes ~1/2 length of parameres.
</paragraph>
<paragraph id="422236DE62529D21EE9CFD91FC68E83D" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFD91FE85E819" box="[199,348,512,536]" italics="true" pageId="51" pageNumber="52">Female adult</emphasis>
. As for male, except as follows.
<emphasis id="70E9EACC62529D21EC95FD91FCFFE819" bold="true" box="[718,806,512,536]" pageId="51" pageNumber="52">Thorax</emphasis>
: Length 1.32 ±
<quantity id="85659B3B62529D21ED80FE6EFBE6E816" box="[987,1087,511,535]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="51" pageNumber="52" unit="mm" value="0.2">0.20 mm</quantity>
(
<quantity id="85659B3B62529D21EA14FE6EFB2CE816" box="[1103,1269,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.48" metricValueMin="1.19" pageId="51" pageNumber="52" unit="mm" value="1.335" valueMax="1.48" valueMin="1.19">1.191.48 mm</quantity>
, n = 9).
<emphasis id="70E9EACC62529D21EB08FE6EFA4DE819" bold="true" box="[1363,1428,511,536]" pageId="51" pageNumber="52">Wing</emphasis>
: Length 6.1 ±
<quantity id="85659B3B62529D21EF76FDB5FE5BE83A" box="[301,386,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="51" pageNumber="52" unit="mm" value="0.9">0.9 mm</quantity>
(
<quantity id="85659B3B62529D21EFC9FDB5FDF3E83A" box="[402,554,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.8" metricValueMin="5.3" pageId="51" pageNumber="52" unit="mm" value="6.05" valueMax="6.8" valueMin="5.3">5.3 6.8 mm</quantity>
, n = 9).
<emphasis id="70E9EACC62529D21ECDCFDB2FD22E83D" bold="true" box="[647,763,547,572]" pageId="51" pageNumber="52">Abdomen</emphasis>
: Cercus yellow.
</paragraph>
<paragraph id="422236DE62529D21EE9CFDD8FE1BE85E" blockId="51.[151,1436,151,1292]" box="[199,450,583,608]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFDD8FEE7E861" box="[199,318,585,608]" italics="true" pageId="51" pageNumber="52">Immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EE9CFDFAFE16E885" box="[199,463,619,644]" pageId="51" pageNumber="52" type="biology_ecology">
<paragraph id="422236DE62529D21EE9CFDFAFE16E885" blockId="51.[151,1436,151,1292]" box="[199,463,619,644]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFDFAFE93E885" bold="true" box="[199,330,619,644]" pageId="51" pageNumber="52">BIOLOGY</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EE9CFD1EFDD5E8CD" pageId="51" pageNumber="52" type="distribution">
<paragraph id="422236DE62529D21EE9CFD1EFDD5E8CD" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFD1EFE4FE8A9" bold="true" box="[199,406,655,680]" pageId="51" pageNumber="52">DISTRIBUTION</emphasis>
:
<collectingCountry id="3A8A764E62529D21EFFEFD1EFE20E8A6" box="[421,505,655,679]" name="Guinea-Bissau" pageId="51" pageNumber="52">Guinea</collectingCountry>
,
<collectingCountry id="3A8A764E62529D21EC5FFD01FD52E8A6" box="[516,651,655,680]" name="Ivory Coast" pageId="51" pageNumber="52">Ivory Coast</collectingCountry>
,
<collectingCountry id="3A8A764E62529D21ECCCFD1EFD3AE8A6" box="[663,739,655,679]" name="Ghana" pageId="51" pageNumber="52">Ghana</collectingCountry>
,
<collectingCountry id="3A8A764E62529D21ECB5FD1EFCE2E8A6" box="[750,827,655,679]" name="Gabon" pageId="51" pageNumber="52">Gabon</collectingCountry>
, the
<collectingCountry id="3A8A764E62529D21ED2AFD1EFB54E8A6" box="[881,1165,655,679]" name="Central African Republic" pageId="51" pageNumber="52">Central African Republic</collectingCountry>
and the
<collectingCountry id="3A8A764E62529D21EAB5FD1EFF3BE8CD" name="Republic of the Congo" pageId="51" pageNumber="52">Republic of the Congo</collectingCountry>
(
<figureCitation id="DAA62A5B62529D21EEAAFD25FE8BE8CD" box="[241,338,692,716]" captionStart="FIGURE 134136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="51" pageNumber="52">Fig. 136</figureCitation>
),
<quantity id="85659B3B62529D21EF3DFD25FDDEE8CD" box="[358,519,692,716]" metricMagnitude="2" metricUnit="m" metricValue="4.35" metricValueMax="7.5" metricValueMin="1.2" pageId="51" pageNumber="52" unit="m" value="435.0" valueMax="750.0" valueMin="120.0">120750 masl</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EE9CFD46FC77E939" pageId="51" pageNumber="52" type="discussion">
<paragraph id="422236DE62529D21EE9CFD46FC77E939" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFD46FEABE8F1" bold="true" box="[199,370,727,752]" pageId="51" pageNumber="52">DISCUSSION</emphasis>
: As discussed below in the phylogeny section, the placement of
<taxonomicName id="859D4D5D62529D21EA27FD49FAA3E8EE" box="[1148,1402,727,752]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="70E9EACC62529D21EA27FD49FAC7E8F1" box="[1148,1310,728,752]" italics="true" pageId="51" pageNumber="52">Leptomorphus</emphasis>
species
</taxonomicName>
in subgenera (
<bibRefCitation id="260C4B2F62529D21EF43FD6DFE7DE915" author="Matile, L." box="[280,420,764,788]" pageId="51" pageNumber="52" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
<taxonomicName id="859D4D5D62529D21EF5DFCB1FE7FE939" box="[262,422,800,824]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="subGenus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC62529D21EF5DFCB1FE7FE939" box="[262,422,800,824]" italics="true" pageId="51" pageNumber="52">Gymnoscutum</emphasis>
</taxonomicName>
and placed solely in the genus
<emphasis id="70E9EACC62529D21ED5EFCB1FC77E939" box="[773,942,800,824]" italics="true" pageId="51" pageNumber="52">
<taxonomicName id="859D4D5D62529D21ED5EFCB1FC72E939" authorityName=": Edwards" authorityYear="1925" box="[773,939,800,824]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EE9CFCD2FD6BEF0D" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="422236DE62529D21EE9CFCD2FC4EE9C6" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<materialsCitation id="F2F53C8362529D21EE9CFCD2FC4EE9C6" collectingDate="1970-09-29" collectionCode="MATERIAL" collectorName="Det. C. J. Borkent" determinerName="L. Matile &amp; Det. C. J. Borkent" location="Matile" pageId="51" pageNumber="52" specimenCount="2" specimenCount-male="2" stateProvince="Monaghan" typeStatus="holotype">
<emphasis id="70E9EACC62529D21EE9CFCD2FDDBE95D" bold="true" box="[199,514,835,860]" pageId="51" pageNumber="52">
<collectionCode id="248CAE1B62529D21EE9CFCD2FE86E95D" box="[199,351,835,860]" pageId="51" pageNumber="52">MATERIAL</collectionCode>
EXAMINED
</emphasis>
:
<emphasis id="70E9EACC62529D21EC4BFCD5FD59E95D" box="[528,640,836,860]" italics="true" pageId="51" pageNumber="52">
<typeStatus id="9D26887C62529D21EC4BFCD5FDA3E95D" box="[528,634,836,860]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
:
</emphasis>
adult male, pinned on double mount minuten, genitalia in glass vial on pin, labelled “[Blue label] REP. CENTRAFRIC./ LA MABOKE/
<date id="3623101E62529D21EDF0FCF6FBF0E97E" box="[939,1065,871,895]" pageId="51" pageNumber="52" value="1970-09-29">
<collectingDate id="2667E9F662529D21EDF0FCF6FBF0E97E" box="[939,1065,871,895]" pageId="51" pageNumber="52" value="1970-09-29">29.IX.1970</collectingDate>
</date>
/ L. MATILE rec.; [Red label]
<typeStatus id="9D26887C62529D21EECCFC1DFEF7E9A5" box="[151,302,908,932]" pageId="51" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
;
<taxonomicName id="859D4D5D62529D21EF65FC1DFE39E9A5" authorityName=": Edwards" authorityYear="1925" box="[318,480,908,932]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
/ (Afroleptomorphus)/ obscurus
<taxonomicNameLabel id="6BDA57B762529D21ED01FC1DFC42E9A2" box="[858,923,908,931]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
<specimenCount id="549BFD5762529D21EDFCFC1DFC65E9A2" box="[935,956,908,931]" pageId="51" pageNumber="52" type="male"></specimenCount>
ht/
<determinerName id="B7FEDA1A62529D21EDBEFC1DFB8BE9A5" box="[997,1106,908,932]" pageId="51" pageNumber="52">L. Matile</determinerName>
det. 1974;
<typeStatus id="9D26887C62529D21EA84FC1DFAAAE9A5" box="[1247,1395,908,932]" pageId="51" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="549BFD5762529D21EB24FC1DFA4DE9A2" box="[1407,1428,908,931]" pageId="51" pageNumber="52" type="male"></specimenCount>
/
<taxonomicName id="859D4D5D62529D21EECCFC21FE7DE9C9" box="[151,420,944,968]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="70E9EACC62529D21EECCFC21FE7DE9C9" box="[151,420,944,968]" italics="true" pageId="51" pageNumber="52">Leptomorphus obscurus</emphasis>
</taxonomicName>
/
<location id="4742600562529D21EFEFFC3EFE25E9C6" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21EFEFFC3EFE25E9C6" box="[436,508,943,967]" name="Matile" pageId="51" pageNumber="52" stateProvince="Monaghan">Matile</location>
/
<determinerName id="B7FEDA1A62529D21EC57FC21FD08E9C6" box="[524,721,943,967]" pageId="51" pageNumber="52">
<collectorName id="EF68530862529D21EC57FC21FD08E9C6" box="[524,721,943,967]" pageId="51" pageNumber="52">Det. C.J. Borkent</collectorName>
</determinerName>
, 2012” [
<collectingRegion id="8059F83C62529D21ED75FC21FC55E9C6" box="[814,908,944,967]" country="Ireland" name="Monaghan" pageId="51" pageNumber="52">MNHN</collectingRegion>
].
</materialsCitation>
</paragraph>
<paragraph id="422236DE62529D21EE9CFC44FD70EE0E" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFC44FE98E9ED" box="[199,321,981,1004]" italics="true" pageId="51" pageNumber="52">
<typeStatus id="9D26887C62529D21EE9CFC44FEE5E9ED" box="[199,316,981,1004]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
:
</emphasis>
labelled as for
<typeStatus id="9D26887C62529D21EFABFC45FD8BE9ED" box="[496,594,980,1004]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
(
<specimenCount id="549BFD5762529D21EC38FC45FD5FE9EA" box="[611,646,980,1004]" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
,
<specimenCount id="549BFD5762529D21ECCEFC45FD61E9EA" box="[661,696,980,1004]" pageId="51" pageNumber="52" type="female">4♀</specimenCount>
(1 labelled
<typeStatus id="9D26887C62529D21ED65FC45FC44E9ED" box="[830,925,980,1004]" pageId="51" pageNumber="52" type="allotype">allotype</typeStatus>
)); except
<date id="3623101E62529D21EA57FC45FB52E9ED" box="[1036,1163,980,1004]" pageId="51" pageNumber="52" value="1970-09-24">24.IX.1970</date>
(
<specimenCount id="549BFD5762529D21EAC5FC45FB66E9EA" box="[1182,1215,980,1004]" pageId="51" pageNumber="52" type="female">1♀</specimenCount>
);
<materialsCitation id="F2F53C8362529D21EA8CFC45FA4FE9ED" box="[1239,1430,980,1004]" collectingDate="1970-09-26" isTailEnumeration="true" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="3623101E62529D21EA8CFC45FA8EE9ED" box="[1239,1367,980,1004]" pageId="51" pageNumber="52" value="1970-09-26">
<collectingDate id="2667E9F662529D21EA8CFC45FA8EE9ED" box="[1239,1367,980,1004]" pageId="51" pageNumber="52" value="1970-09-26">26.IX.1970</collectingDate>
</date>
(
<specimenCount id="549BFD5762529D21EB32FC45FA52E9EA" box="[1385,1419,980,1004]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
)
</materialsCitation>
;
<date id="3623101E62529D21EECCFC66FECFEE0E" box="[151,278,1015,1039]" pageId="51" pageNumber="52" value="1970-09-27">27.IX.1970</date>
(
<specimenCount id="549BFD5762529D21EF7DFC66FE90EE0E" box="[294,329,1015,1039]" pageId="51" pageNumber="52" type="male">2♂</specimenCount>
,
<specimenCount id="549BFD5762529D21EF03FC66FEA3EE0E" box="[344,378,1015,1039]" pageId="51" pageNumber="52" type="female">1♀</specimenCount>
).
<materialsCitation id="F2F53C8362529D21EFC9FC66FD7CEE0E" box="[402,677,1015,1040]" collectorName="All" location="MNHN" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Monaghan" typeStatus="paratype">
<collectorName id="EF68530862529D21EFC9FC66FE6FEE0E" box="[402,438,1015,1039]" pageId="51" pageNumber="52">All</collectorName>
<typeStatus id="9D26887C62529D21EFE6FC69FDF1EE11" box="[445,552,1016,1040]" pageId="51" pageNumber="52" type="paratype">paratypes</typeStatus>
in
<collectingRegion id="8059F83C62529D21EC17FC69FD7CEE0E" box="[588,677,1016,1039]" country="Ireland" name="Monaghan" pageId="51" pageNumber="52">MNHN</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="422236DE62529D21EE9CFB8DFD6BEF0D" blockId="51.[151,1436,151,1292]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CFB8DFEA0EE35" box="[199,377,1052,1076]" italics="true" pageId="51" pageNumber="52">Other material:</emphasis>
<materialsCitation id="F2F53C8362529D21EFDAFB8DFA8FEE35" box="[385,1366,1052,1076]" collectingDate="1970-09-26" collectionCode="MNHN" collectorName="L. Matile." country="Central African Republic" location="La Maboke" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E62529D21EFDAFB8DFCC1EE35" box="[385,792,1052,1076]" name="Central African Republic" pageId="51" pageNumber="52">CENTRAL AFRICAN REPUBLIC</collectingCountry>
,
<location id="4742600562529D21ED7DFB8DFC73EE35" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21ED7DFB8DFC73EE35" box="[806,938,1052,1076]" country="Central African Republic" name="La Maboke" pageId="51" pageNumber="52">La Maboke</location>
,
<date id="3623101E62529D21EDECFB8DFBE8EE35" box="[951,1073,1052,1076]" pageId="51" pageNumber="52" value="1970-09-26">
<collectingDate id="2667E9F662529D21EDECFB8DFBE8EE35" box="[951,1073,1052,1076]" pageId="51" pageNumber="52" value="1970-09-26">26.ix.1970</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EA65FB8DFB76EE35" box="[1086,1199,1052,1076]" pageId="51" pageNumber="52">L. Matile.</collectorName>
(
<specimenCount id="549BFD5762529D21EA99FB8DFB3DEE32" box="[1218,1252,1052,1076]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EAA8FB8DFA97EE35" box="[1267,1358,1052,1076]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EB39FB8DFDD3EE56" collectingDate="1970-09-27" collectionCode="MNHN" country="Central African Republic" pageId="51" pageNumber="52" specimenCount="2" specimenCount-male="2">
same except
<date id="3623101E62529D21EEBCFBAEFEBBEE56" box="[231,354,1087,1111]" pageId="51" pageNumber="52" value="1970-09-27">
<collectingDate id="2667E9F662529D21EEBCFBAEFEBBEE56" box="[231,354,1087,1111]" pageId="51" pageNumber="52" value="1970-09-27">27.ix.1970</collectingDate>
</date>
. (
<specimenCount id="549BFD5762529D21EF2EFBAEFE40EE56" box="[373,409,1087,1111]" pageId="51" pageNumber="52" type="male">2♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EFFCFBAEFDDBEE56" box="[423,514,1087,1111]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EC4EFBAEFCE1EE56" box="[533,824,1087,1111]" collectingDate="1970-09-29" collectionCode="MNHN" country="Central African Republic" isTailEnumeration="true" pageId="51" pageNumber="52" specimenCount="3" specimenCount-male="3">
<date id="3623101E62529D21EC4EFBAEFD49EE56" box="[533,656,1087,1111]" pageId="51" pageNumber="52" value="1970-09-29">
<collectingDate id="2667E9F662529D21EC4EFBAEFD49EE56" box="[533,656,1087,1111]" pageId="51" pageNumber="52" value="1970-09-29">29.ix.1970</collectingDate>
</date>
. (
<specimenCount id="549BFD5762529D21ECFFFBAEFD1EEE56" box="[676,711,1087,1111]" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EC8EFBAEFCE9EE56" box="[725,816,1087,1111]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21ED18FBAEFE61EE7D" collectingDate="1979-05-07" collectingDateMax="1979-05-16" collectingDateMin="1979-05-07" collectionCode="MNHN" collectorName="J. Legrand." country="Gabon" location="Makokou" municipality="Bale Affl." pageId="51" pageNumber="52" specimenCount="2" specimenCount-male="2">
<collectingCountry id="3A8A764E62529D21ED18FBAEFC73EE56" box="[835,938,1087,1111]" name="Gabon" pageId="51" pageNumber="52">GABON</collectingCountry>
,
<location id="4742600562529D21EDEEFBAEFBF9EE56" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21EDEEFBAEFBF9EE56" box="[949,1056,1087,1111]" country="Gabon" municipality="Bale Affl." name="Makokou" pageId="51" pageNumber="52">Makokou</location>
m'passa,
<collectingMunicipality id="A246ACA462529D21EAD6FBAEFAD9EE56" box="[1165,1280,1087,1111]" pageId="51" pageNumber="52">Bale Affl.</collectingMunicipality>
,
<date id="3623101E62529D21EB50FBAEFA4EEE56" box="[1291,1431,1087,1111]" pageId="51" pageNumber="52" value="1979-05-07" valueMax="1979-05-16" valueMin="1979-05-07">
<collectingDate id="2667E9F662529D21EB50FBAEFA4EEE56" box="[1291,1431,1087,1111]" pageId="51" pageNumber="52" value="1979-05-07" valueMax="1979-05-16" valueMin="1979-05-07">716.v.1979</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EECCFBF5FECDEE7D" box="[151,276,1124,1148]" pageId="51" pageNumber="52">J. Legrand.</collectorName>
(
<specimenCount id="549BFD5762529D21EF7FFBF5FE9EEE7A" box="[292,327,1124,1148]" pageId="51" pageNumber="52" type="male">2♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EF0EFBF5FE69EE7D" box="[341,432,1124,1148]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EF98FBF5FB98EE7D" box="[451,1089,1124,1148]" collectingDate="1994-11-08" collectionCode="ZMUN" country="Ghana" location="Kakum National Park" pageId="51" pageNumber="52" specimenCount="2" specimenCount-male="2">
<collectingCountry id="3A8A764E62529D21EF98FBF5FDF2EE7D" box="[451,555,1124,1148]" name="Ghana" pageId="51" pageNumber="52">GHANA</collectingCountry>
,
<location id="4742600562529D21EC6DFBF5FCF4EE7D" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21EC6DFBF5FCF4EE7D" box="[566,813,1124,1148]" country="Ghana" name="Kakum National Park" pageId="51" pageNumber="52">Kakum National Park</location>
,
<date id="3623101E62529D21ED63FBF5FC7AEE7D" box="[824,931,1124,1148]" pageId="51" pageNumber="52" value="1994-11-08">
<collectingDate id="2667E9F662529D21ED63FBF5FC7AEE7D" box="[824,931,1124,1148]" pageId="51" pageNumber="52" value="1994-11-08">8.xi.1994</collectingDate>
</date>
(
<specimenCount id="549BFD5762529D21EDE9FBF5FC0FEE7A" box="[946,982,1124,1148]" pageId="51" pageNumber="52" type="male">2♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EDBFFBF5FBEFEE7A" box="[996,1078,1124,1147]" pageId="51" pageNumber="52">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EA15FBF5FA4FEE7D" box="[1102,1430,1124,1148]" collectingDate="1994-10-31" collectingDateMax="1994-11-08" collectingDateMin="1994-10-31" collectionCode="ZMUN" country="Ghana" isTailEnumeration="true" location="Kakum National Park" pageId="51" pageNumber="52" specimenCount="2" specimenCount-male="2">
<date id="3623101E62529D21EA15FBF5FB2EEE7D" box="[1102,1271,1124,1148]" pageId="51" pageNumber="52" value="1994-10-31" valueMax="1994-11-08" valueMin="1994-10-31">
<collectingDate id="2667E9F662529D21EA15FBF5FB2EEE7D" box="[1102,1271,1124,1148]" pageId="51" pageNumber="52" value="1994-10-31" valueMax="1994-11-08" valueMin="1994-10-31">31.x8.xi.1994</collectingDate>
</date>
(
<specimenCount id="549BFD5762529D21EB5CFBF5FAF3EE7A" box="[1287,1322,1124,1148]" pageId="51" pageNumber="52" type="male">2♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EB63FBF5FA53EE7A" box="[1336,1418,1124,1147]" pageId="51" pageNumber="52">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EECCFB16FE10EE9E" box="[151,457,1159,1183]" collectingDate="1994-11-08" collectingDateMax="1994-11-15" collectingDateMin="1994-11-08" collectionCode="ZMUN" country="Ghana" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<date id="3623101E62529D21EECCFB16FEF2EE9E" box="[151,299,1159,1183]" pageId="51" pageNumber="52" value="1994-11-08" valueMax="1994-11-15" valueMin="1994-11-08">
<collectingDate id="2667E9F662529D21EECCFB16FEF2EE9E" box="[151,299,1159,1183]" pageId="51" pageNumber="52" value="1994-11-08" valueMax="1994-11-15" valueMin="1994-11-08">815.xi.1994</collectingDate>
</date>
(
<specimenCount id="549BFD5762529D21EF67FB16FE87EE9E" box="[316,350,1159,1183]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EF30FB19FE67EE9E" box="[363,446,1160,1183]" pageId="51" pageNumber="52">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EF8EFB16FCD9EE9E" box="[469,768,1159,1183]" collectingDate="1994-10-08" collectingDateMax="1994-10-15" collectingDateMin="1994-10-08" collectionCode="ZMUN" country="Ghana" pageId="51" pageNumber="52" specimenCount="1" specimenCount-female="1">
<date id="3623101E62529D21EF8EFB16FDB8EE9E" box="[469,609,1159,1183]" pageId="51" pageNumber="52" value="1994-10-08" valueMax="1994-10-15" valueMin="1994-10-08">
<collectingDate id="2667E9F662529D21EF8EFB16FDB8EE9E" box="[469,609,1159,1183]" pageId="51" pageNumber="52" value="1994-10-08" valueMax="1994-10-15" valueMin="1994-10-08">815.x.1994</collectingDate>
</date>
(
<specimenCount id="549BFD5762529D21EC28FB16FD4CEE9E" box="[627,661,1159,1183]" pageId="51" pageNumber="52" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B62529D21ECF9FB19FD2DEE9E" box="[674,756,1160,1183]" pageId="51" pageNumber="52">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21ED57FB19FD7BEEC5" collectingDate="1993-12-08" collectionCode="T, ZMUN" collectorName="J. Kjaerandsen &amp; Anderson" country="Ghana" location="Western Region" municipality="Ankasa" pageId="51" pageNumber="52" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location id="4742600562529D21ED57FB19FC1BEE9E" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21ED57FB19FC1BEE9E" box="[780,962,1159,1183]" country="Ghana" municipality="Ankasa" name="Western Region" pageId="51" pageNumber="52">Western Region</location>
,
<collectingMunicipality id="A246ACA462529D21ED96FB16FBF8EE9E" box="[973,1057,1159,1183]" pageId="51" pageNumber="52">Ankasa</collectingMunicipality>
game prod. reserve,
<date id="3623101E62529D21EB50FB16FAA7EE9E" box="[1291,1406,1159,1183]" pageId="51" pageNumber="52" value="1993-12-08">
<collectingDate id="2667E9F662529D21EB50FB16FAA7EE9E" box="[1291,1406,1159,1183]" pageId="51" pageNumber="52" value="1993-12-08">8.xii.1993</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EBD1FB19FEF9EEC5" pageId="51" pageNumber="52">J. Kjaerandsen</collectorName>
,
<collectionCode id="248CAE1B62529D21EF6AFB3DFE9AEEC2" box="[305,323,1196,1219]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="52">T</collectionCode>
.
<collectorName id="EF68530862529D21EF0FFB3DFE18EEC5" box="[340,449,1196,1220]" pageId="51" pageNumber="52">Anderson</collectorName>
(
<specimenCount id="549BFD5762529D21EF82FB3DFE23EEC2" box="[473,506,1196,1220]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<specimenCount id="549BFD5762529D21EC55FB3DFDE8EEC2" box="[526,561,1196,1220]" pageId="51" pageNumber="52" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B62529D21EC1EFB3DFD4EEEC2" box="[581,663,1196,1219]" pageId="51" pageNumber="52">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21ECEDFB3DFF23EEE6" collectingDate="1991-06-18" collectingDateMax="1991-06-29" collectingDateMin="1991-06-18" collectionCode="MNHN" collectorName="Girard &amp; Legrand." country="Guinea-Bissau" location="Mt. Nimba" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E62529D21ECEDFB3DFCFDEEC5" box="[694,804,1196,1220]" name="Guinea-Bissau" pageId="51" pageNumber="52">GUINEA</collectingCountry>
,
<location id="4742600562529D21ED6EFB3DFC60EEC5" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21ED6EFB3DFC60EEC5" box="[821,953,1196,1220]" country="Guinea-Bissau" name="Mt. Nimba" pageId="51" pageNumber="52">Mt. Nimba</location>
,
<date id="3623101E62529D21ED91FB3DFBB6EEC5" box="[970,1135,1196,1220]" pageId="51" pageNumber="52" value="1991-06-18" valueMax="1991-06-29" valueMin="1991-06-18">
<collectingDate id="2667E9F662529D21ED91FB3DFBB6EEC5" box="[970,1135,1196,1220]" pageId="51" pageNumber="52" value="1991-06-18" valueMax="1991-06-29" valueMin="1991-06-18">1829.vi.1991</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EADBFB3DFB10EEC5" box="[1152,1225,1196,1220]" pageId="51" pageNumber="52">Girard</collectorName>
et
<collectorName id="EF68530862529D21EAACFB3DFA85EEC5" box="[1271,1372,1196,1220]" pageId="51" pageNumber="52">Legrand.</collectorName>
(
<specimenCount id="549BFD5762529D21EB2FFB3DFA4CEEC2" box="[1396,1429,1196,1220]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EECCFB5EFF2BEEE6" box="[151,242,1231,1255]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EF5DFB5EFC74EEE6" box="[262,941,1231,1256]" collectingDate="1980-05-05" collectionCode="MNHN" collectorName="G. Couturier." country="Ivory Coast" location="Tay" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E62529D21EF5DFB5EFE1AEEE6" box="[262,451,1231,1255]" name="Ivory Coast" pageId="51" pageNumber="52">IVORY COAST</collectingCountry>
,
<location id="4742600562529D21EF8BFB41FE2DEEE9" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21EF8BFB41FE2DEEE9" box="[464,500,1232,1256]" country="Ivory Coast" name="Tay" pageId="51" pageNumber="52">Taї</location>
,
<date id="3623101E62529D21EC5FFB5EFDBFEEE6" box="[516,614,1231,1255]" pageId="51" pageNumber="52" value="1980-05-05">
<collectingDate id="2667E9F662529D21EC5FFB5EFDBFEEE6" box="[516,614,1231,1255]" pageId="51" pageNumber="52" value="1980-05-05">5.v.1980</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EC2FFB5EFCDFEEE6" box="[628,774,1231,1255]" pageId="51" pageNumber="52">G. Couturier.</collectorName>
(
<specimenCount id="549BFD5762529D21ED42FB5EFCE2EEE6" box="[793,827,1231,1255]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21ED10FB5EFC7FEEE6" box="[843,934,1231,1255]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C8362529D21EDE1FB5EFD77EF0D" collectingDate="1975-11-14" collectionCode="MNHN" collectorName="L. Matile." country="Republic of the Congo" location="Mayombe Dimonika" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E62529D21EDE1FB5EFAC4EEE6" box="[954,1309,1231,1255]" name="Republic of the Congo" pageId="51" pageNumber="52">REPUBLIC OF THE CONGO</collectingCountry>
,
<location id="4742600562529D21EB71FB5EFED3EF0D" LSID="urn:lsid:plazi:treatment:CA3487C862539D21EECCFF06FD6BEF0D:4742600562529D21EB71FB5EFED3EF0D" country="Republic of the Congo" name="Mayombe Dimonika" pageId="51" pageNumber="52">Mayombe Dimonika</location>
,
<date id="3623101E62529D21EF4EFB65FE57EF0D" box="[277,398,1268,1292]" pageId="51" pageNumber="52" value="1975-11-14">
<collectingDate id="2667E9F662529D21EF4EFB65FE57EF0D" box="[277,398,1268,1292]" pageId="51" pageNumber="52" value="1975-11-14">14.xi.1975</collectingDate>
</date>
,
<collectorName id="EF68530862529D21EFC1FB65FDD3EF0D" box="[410,522,1268,1292]" pageId="51" pageNumber="52">L. Matile.</collectorName>
(
<specimenCount id="549BFD5762529D21EC47FB65FDE4EF0A" box="[540,573,1268,1292]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B62529D21EC10FB65FD7FEF0D" box="[587,678,1268,1292]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="52">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>