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<mods:title id="F27051C6DFDEF82013B693030CF03BA0">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<subSubSection id="0A87655562529D21EECCFACEFD91EF78" box="[151,584,1375,1401]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="422236DE62529D21EECCFACEFD91EF78" blockId="51.[151,584,1375,1434]" box="[151,584,1375,1401]" pageId="51" pageNumber="52">
<heading id="196A81B262529D21EECCFACEFD91EF78" bold="true" box="[151,584,1375,1401]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="70E9EACC62529D21EECCFACEFD91EF78" bold="true" box="[151,584,1375,1401]" pageId="51" pageNumber="52">
25.
<taxonomicName id="859D4D5D62529D21EE98FACEFD91EF78" ID-CoL="8CJ3P" authority="Brunetti" authorityName="Brunetti" box="[195,584,1375,1401]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="70E9EACC62529D21EE98FACEFE0EEF78" bold="true" box="[195,471,1375,1401]" italics="true" pageId="51" pageNumber="52">Leptomorphus ornatus</emphasis>
Brunetti
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EECCFA13FE2BEF9B" box="[151,498,1410,1434]" pageId="51" pageNumber="52" type="description">
<paragraph id="422236DE62529D21EECCFA13FE2BEF9B" blockId="51.[151,584,1375,1434]" box="[151,498,1410,1434]" pageId="51" pageNumber="52">
(
<figureCitation id="DAA62A5B62529D21EEC5FA13FEC0EF9B" box="[158,281,1410,1434]" captionStart="FIGURES 3033" captionStartId="89.[151,263,1758,1781]" captionTargetBox="[157,1431,223,1729]" captionTargetId="figure-41@89.[151,1436,217,1737]" captionTargetPageId="89" captionText="FIGURES 3033. Leptomorphus habitus. 30. L. ornatus, female. 31. L. panorpiformis, male. 32. L. perplexus, paratype female. 33. L. quadrimaculatus, male." pageId="51" pageNumber="52">Figures 30</figureCitation>
,
<figureCitation id="DAA62A5B62529D21EF7EFA13FE9AEF9B" box="[293,323,1410,1434]" captionStart="FIGURES 5669" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 5669. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="51" pageNumber="52">67</figureCitation>
,
<figureCitation id="DAA62A5B62529D21EF14FA13FEA2EF9B" box="[335,379,1410,1434]" captionStart="FIGURE 137139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp &amp; Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="51" pageNumber="52">137</figureCitation>
,
<figureCitation id="DAA62A5B62529D21EFDCFA13FE6AEF9B" box="[391,435,1410,1434]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values&gt;50% are shown in bold below the branches." pageId="51" pageNumber="52">150</figureCitation>
,
<figureCitation id="DAA62A5B62529D21EFE4FA13FE33EF9B" box="[447,490,1410,1434]" captionStart="FIGURE 152" captionStartId="114.[151,250,1784,1807]" captionTargetBox="[162,1422,1059,1758]" captionTargetId="figure-112@114.[156,1428,1046,1779]" captionTargetPageId="114" captionText="FIGURE 152. One of three equally parsimonious trees showing the relationships between species in the L. ornatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="51" pageNumber="52">152</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EECCFA5BFB96EC66" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="422236DE62529D21EECCFA5BFD40EFE1" blockId="51.[151,749,1482,1535]" box="[151,665,1482,1504]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="628D11F062529D21EECCFA5BFD40EFE1" box="[151,665,1482,1504]" pageId="51" pageNumber="52">
<treatmentCitation id="C33C10CF62529D21EECCFA5BFDE7EFE1" author="Brunetti, E." box="[151,574,1482,1504]" page="85" pageId="51" pageNumber="52" year="1912">
<taxonomicName id="859D4D5D62529D21EECCFA5BFDE7EFE1" ID-CoL="8CJ3P" authority="Brunetti, 1912: 85" authorityName="Brunetti" authorityPageNumber="85" authorityYear="1912" box="[151,574,1482,1504]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="70E9EACC62529D21EECCFA5BFEA5EFE1" box="[151,380,1482,1504]" italics="true" pageId="51" pageNumber="52">Leptomorphus ornatus</emphasis>
<bibRefCitation id="260C4B2F62529D21EFD8FA5BFDE7EFE1" author="Brunetti, E." box="[387,574,1482,1504]" pageId="51" pageNumber="52" refId="ref92314" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. and Marshall, G. A. K. (eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, 581 pp., 12 pls." type="book" year="1912">Brunetti, 1912: 85</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Plate II.
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</paragraph>
<paragraph id="422236DE62529D21EECCFA78FD34EFFE" blockId="51.[151,749,1482,1535]" box="[151,749,1513,1535]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="628D11F062529D21EECCFA78FD34EFFE" box="[151,749,1513,1535]" pageId="51" pageNumber="52">
<treatmentCitation id="C33C10CF62529D21EECCFA78FD30EFFE" author="Matile, L." box="[151,745,1513,1535]" page="145" pageId="51" pageNumber="52" year="1977">
<taxonomicName id="859D4D5D62529D21EECCFA78FD30EFFE" ID-CoL="8CJ3P" authority=": Matile, 1977: 145" authorityName=": Matile" authorityPageNumber="145" authorityYear="1977" box="[151,745,1513,1535]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC62529D21EECCFA78FDFDEFFE" box="[151,548,1513,1535]" italics="true" pageId="51" pageNumber="52">Leptomorphus (Gymnoscutum) ornatus</emphasis>
:
<bibRefCitation id="260C4B2F62529D21EC69FA78FD30EFFE" author="Matile, L." box="[562,745,1513,1535]" pageId="51" pageNumber="52" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 145</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="422236DE62529D21EECCF9BAFB96EC66" blockId="51.[151,1437,1579,2036]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EECCF9BAFEC0EC45" bold="true" box="[151,281,1579,1604]" pageId="51" pageNumber="52">References</emphasis>
:
<bibRefCitation id="260C4B2F62529D21EF72F9BDFDE5EC45" author="Edwards, F. W." box="[297,572,1580,1604]" pageId="51" pageNumber="52" pagination="223 - 296" refId="ref93098" refString="Edwards, F. W. (1933 a) Diptera Nematocera from Mount Kinabulu. Journal of the Federated Malay States Museums, 17, 223 - 296." type="journal article" year="1933">Edwards 1933a: 22930</bibRefCitation>
(comparison to
<taxonomicName id="859D4D5D62529D21ECA2F9BCFCB4EC45" ID-CoL="8CJ45" authorityName=": Matile" authorityYear="1977" box="[761,877,1580,1604]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="chaseni">
<emphasis id="70E9EACC62529D21ECA2F9BCFCB4EC45" box="[761,877,1580,1604]" italics="true" pageId="51" pageNumber="52">L. chaseni</emphasis>
</taxonomicName>
);
<bibRefCitation id="260C4B2F62529D21EDDCF9BDFB65EC45" author="Colless, D. H. &amp; Liepa Z." box="[903,1212,1580,1604]" pageId="51" pageNumber="52" pagination="1973" refId="ref92524" refString="Colless, D. H. &amp; Liepa Z. (1973) Superfamily Myceptophiloidea, family Mycetophilidae. In: Delfinado, M. D. and Hardy, D. E. (eds.) 1973. A catalog of the Diptera of the Oriental Region. Vol. 1. Suborder Nematocera. University Press of Hawaii, Honolulu, pp. 444 - 463." type="book chapter" year="1973">Colless &amp; Liepa 1973: 454</bibRefCitation>
(catalogue);
<bibRefCitation id="260C4B2F62529D21EB08F9BDFF0AEC66" author="Matile, L." pageId="51" pageNumber="52" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
: 141,145 (subgeneric placement);
<bibRefCitation id="260C4B2F62529D21EC0FF9C1FCAAEC66" author="Papp, L. &amp; Sevcik, J." box="[596,883,1615,1640]" pageId="51" pageNumber="52" pagination="139 - 159" refId="ref95170" refString="Papp, L. &amp; Sevcik, J. (2011) Eight new oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientarum Hungaricae, 57, 139 - 159." type="journal article" year="2011">Papp &amp; Ševčík 2011: 139</bibRefCitation>
(notes on identity).
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D21EE9CF9E2FD62EDAD" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="422236DE62529D21EE9CF9E2FEC9ECF6" blockId="51.[151,1437,1579,2036]" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CF9E2FEBDEC8D" bold="true" box="[199,356,1651,1676]" pageId="51" pageNumber="52">DIAGNOSIS</emphasis>
: The only extant species of
<taxonomicName id="859D4D5D62529D21EC92F9E5FCB2EC8D" authorityName=": Edwards" authorityYear="1925" box="[713,875,1652,1676]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC62529D21EC92F9E5FCB2EC8D" box="[713,875,1652,1676]" italics="true" pageId="51" pageNumber="52">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: laterotergite and anepisternum brown; wing with pale brown apical wing spot reaching to wing tip, pale medial wing spot absent (
<figureCitation id="DAA62A5B62529D21EF36F92DFE1DECD5" box="[365,452,1724,1748]" captionStart="FIGURES 5669" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 5669. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="51" pageNumber="52">Fig. 67</figureCitation>
); male genitalia presumably different from other species with these characters (male unknown).
</paragraph>
<paragraph id="422236DE62529D21EE9CF895FD62EDAD" blockId="51.[151,1437,1579,2036]" pageId="51" pageNumber="52">
This species can be distinguished from most other Oriental and eastern Palaearctic species (including those in
<bibRefCitation id="260C4B2F62529D21EECCF8B9FE50ED3E" author="Papp, L. &amp; Sevcik, J." box="[151,393,1831,1856]" pageId="51" pageNumber="52" pagination="139 - 159" refId="ref95170" refString="Papp, L. &amp; Sevcik, J. (2011) Eight new oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientarum Hungaricae, 57, 139 - 159." type="journal article" year="2011">Papp &amp; Ševčík 2011</bibRefCitation>
) based on the yellow katepisternum and metepisternum (
<figureCitation id="DAA62A5B62529D21EA7CF8B6FBA4ED3E" box="[1063,1149,1831,1855]" captionStart="FIGURES 3033" captionStartId="89.[151,263,1758,1781]" captionTargetBox="[157,1431,223,1729]" captionTargetId="figure-41@89.[151,1436,217,1737]" captionTargetPageId="89" captionText="FIGURES 3033. Leptomorphus habitus. 30. L. ornatus, female. 31. L. panorpiformis, male. 32. L. perplexus, paratype female. 33. L. quadrimaculatus, male." pageId="51" pageNumber="52">Fig. 30</figureCitation>
, both brown in all other species in these regions except
<emphasis id="70E9EACC62529D21EFAFF8DCFD81ED65" box="[500,600,1868,1892]" italics="true" pageId="51" pageNumber="52">
<taxonomicName id="859D4D5D62529D21EFAFF8DCFD8DED65" authorityName=": Matile" authorityYear="1977" box="[500,596,1868,1892]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="babai">L. babai</taxonomicName>
,
</emphasis>
and
<taxonomicName id="859D4D5D62529D21ECD5F8DCFC92ED65" authorityName="Borkent" authorityYear="2012" box="[654,843,1869,1892]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="titiwangsensis">
<emphasis id="70E9EACC62529D21ECD5F8DCFC92ED65" box="[654,843,1869,1892]" italics="true" pageId="51" pageNumber="52">L. titiwangsensis</emphasis>
</taxonomicName>
). It can be distinguished from
<taxonomicName id="859D4D5D62529D21EAF3F8DCFADCED65" authorityName=": Matile" authorityYear="1977" box="[1192,1285,1868,1892]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="70E9EACC62529D21EAF3F8DCFADCED65" box="[1192,1285,1868,1892]" italics="true" pageId="51" pageNumber="52">L. babai</emphasis>
</taxonomicName>
by its yellow scutellum (brown in
<taxonomicName id="859D4D5D62529D21EFDFF8E0FE3DED89" authorityName=": Matile" authorityYear="1977" box="[388,484,1904,1928]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="70E9EACC62529D21EFDFF8E0FE3DED89" box="[388,484,1904,1928]" italics="true" pageId="51" pageNumber="52">L. babai</emphasis>
</taxonomicName>
) and from
<taxonomicName id="859D4D5D62529D21EC3DF8E0FCFDED86" authorityName="Borkent" authorityYear="2012" box="[614,804,1904,1928]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="titiwangsensis">
<emphasis id="70E9EACC62529D21EC3DF8E0FCFDED86" box="[614,804,1904,1928]" italics="true" pageId="51" pageNumber="52">L. titiwangsensis</emphasis>
</taxonomicName>
by
<emphasis id="70E9EACC62529D21ED0AF8E0FC16ED89" box="[849,975,1904,1928]" italics="true" pageId="51" pageNumber="52">L. ornatus</emphasis>
occurrence only in
<collectingCountry id="3A8A764E62529D21EAEFF8FEFB2EED86" box="[1204,1271,1903,1927]" name="Nepal" pageId="51" pageNumber="52">Nepal</collectingCountry>
and
<collectingCountry id="3A8A764E62529D21EB6AF8FEFAB3ED86" box="[1329,1386,1903,1927]" name="India" pageId="51" pageNumber="52">India</collectingCountry>
and presumably different male genitalic morphology.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562529D26EE9CF826FE1BEFCA" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
<paragraph id="422236DE62529D26EE9CF826FCCEEF82" blockId="51.[151,1437,1579,2036]" lastBlockId="52.[151,1437,151,2024]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="70E9EACC62529D21EE9CF826FE50EDD1" bold="true" box="[199,393,1975,2000]" pageId="51" pageNumber="52">DESCRIPTION</emphasis>
:
<emphasis id="70E9EACC62529D21EFC3F829FE35EDD1" box="[408,492,1976,2000]" italics="true" pageId="51" pageNumber="52">Female</emphasis>
. (
<figureCitation id="DAA62A5B62529D21EC59F829FD8AEDD1" box="[514,595,1976,2000]" captionStart="FIGURES 3033" captionStartId="89.[151,263,1758,1781]" captionTargetBox="[157,1431,223,1729]" captionTargetId="figure-41@89.[151,1436,217,1737]" captionTargetPageId="89" captionText="FIGURES 3033. Leptomorphus habitus. 30. L. ornatus, female. 31. L. panorpiformis, male. 32. L. perplexus, paratype female. 33. L. quadrimaculatus, male." pageId="51" pageNumber="52">Fig. 30</figureCitation>
)
<emphasis id="70E9EACC62529D21EC38F829FD7BEDD1" bold="true" box="[611,674,1976,2000]" pageId="51" pageNumber="52">Head</emphasis>
: yellow, circular in anterior view. Antenna with basal flagellomeres lighter brown, darkening apically; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 12 large bristles, few setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base yellow remainder either light brown or brown; flagellomere 6 1.1X as long as broad. Clypeus yellow, dorsoventrally elongate oval; bristles on clypeus yellow, 68 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 1.5X as long as face. Face yellow; shape a just longer than wide triangle, with few bristles ventrolaterally. Frons yellow; with few bristles medioventrally, frontal furrow running 3/4 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus yellow; segment 1 hidden behind eye, segments increasing in length, segment 5 2X length of segment 4 with central half thinner than base and apex, segment 3 appears to have apicolateral patch of fine yellow setae weakly encircled by strong dark setae (difficult to see in specimens). Labellum yellow. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black with electric blue green specks.
<emphasis id="70E9EACC62559D26EABEFDD9FAE7E861" bold="true" box="[1253,1342,584,608]" pageId="52" pageNumber="53">Thorax</emphasis>
: Length
<quantity id="85659B3B62559D26EECCFDFDFF20E882" box="[151,249,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="52" pageNumber="53" unit="mm" value="1.31">1.31 mm</quantity>
(
<quantity id="85659B3B62559D26EF53FDFDFE77E882" box="[264,430,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" metricValueMax="1.35" metricValueMin="1.27" pageId="52" pageNumber="53" unit="mm" value="1.31" valueMax="1.35" valueMin="1.27">1.271.35 mm</quantity>
, n = 3). Mostly brown, with anterolateral area yellow. Scutum dark brown with blue-green specks; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae on posterior two thirds; two thirds row of lateral setae present; patch of setae on scutum at wing base present. Scutellum yellow; with 6 large bristles and many small bristles. Prescutum brown. Mediotergite dark brown with 4 bristles on posterolateral corners, absent. Laterotergite brown; anterior margin of laterotergite abutting katepisternum. Anepimeron brown with posterodorsal corner yellow. Anepisternum brown. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter stem yellow, knob light brown.
<emphasis id="70E9EACC62559D26EECCFC1DFF16E9A5" bold="true" box="[151,207,908,932]" pageId="52" pageNumber="53">Legs</emphasis>
: principally yellow; hind coxa with brown spot along length of anterior surface; hind femur with apex and basal 1/3 light brown; extreme anteroapical corner yellow on forefemur, on other femora dark brown; midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, tibial spurs brown, foretibial spur length 2X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 3/4 of its length, placed basally, shortest midtibial spur 0.5X length of longest, longest midtibial spur 5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 3X apical thickness of hind tibia. Foreleg first tarsomere 1.4X length of foretibia.
<emphasis id="70E9EACC62559D26EF5FFB16FE9CEEA1" bold="true" box="[260,325,1159,1184]" pageId="52" pageNumber="53">Wing</emphasis>
(
<figureCitation id="DAA62A5B62559D26EF0EFB19FE73EEA1" box="[341,426,1160,1184]" captionStart="FIGURES 5669" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 5669. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="52" pageNumber="53">Fig. 67</figureCitation>
): Length
<quantity id="85659B3B62559D26EC4DFB19FDB5EE9E" box="[534,620,1160,1184]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="52" pageNumber="53" unit="mm" value="5.5">5.5 mm</quantity>
(5.0
<quantity id="85659B3B62559D26ECF5FB19FCDEEE9E" box="[686,775,1160,1184]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="52" pageNumber="53" unit="mm" value="5.7">5.7 mm</quantity>
, n = 3). Hyaline; apical macula light, beginning at apex of R
<subScript id="DE19349B62559D26EEF2FB2AFF68EEC8" attach="left" box="[169,177,1211,1225]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
and M
<subScript id="DE19349B62559D26EF59FB2AFED3EEC8" attach="left" box="[258,266,1211,1225]" fontSize="6" pageId="52" pageNumber="53">4</subScript>
and reaching wing tip; medial macula absent. Macrotrichia in all cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r absent, faint, or present, when present joining R at 23X its own length prior to origin of Rs. R
<subScript id="DE19349B62559D26EA56FA97FBCCEF15" attach="left" box="[1037,1045,1286,1300]" fontSize="6" pageId="52" pageNumber="53">4</subScript>
absent. R
<subScript id="DE19349B62559D26EADAFA97FB50EF15" attach="left" box="[1153,1161,1286,1300]" fontSize="6" pageId="52" pageNumber="53">5</subScript>
straight, slight posterior turn near tip. M
<subScript id="DE19349B62559D26EF10FABCFE8AEF3A" attach="left" box="[331,339,1325,1339]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
reaching apex before R
<subScript id="DE19349B62559D26EC38FABCFDB2EF3A" attach="right" box="[611,619,1325,1339]" fontSize="6" pageId="52" pageNumber="53">5</subScript>
, apices of M veins thinning towards wing margin. M
<subScript id="DE19349B62559D26EA90FABCFB0AEF3A" attach="left" box="[1227,1235,1325,1339]" fontSize="6" pageId="52" pageNumber="53">4</subScript>
-CuA fork arising apically of origin of r-m. A
<subScript id="DE19349B62559D26EF97FAC2FE0DEF60" attach="left" box="[460,468,1363,1377]" fontSize="6" pageId="52" pageNumber="53">2</subScript>
absent.
<emphasis id="70E9EACC62559D26EC69FAD5FD7EEF5C" bold="true" box="[562,679,1348,1373]" pageId="52" pageNumber="53">Abdomen</emphasis>
: Tergites 35 and sometime 6 with anterior 1/3 yellow, T7 light to dark brown, remainder dark brown/black. Cercus yellow.
</paragraph>
<paragraph id="422236DE62559D26EE9CFA1EFE5DEFA6" blockId="52.[151,1437,151,2024]" box="[199,388,1423,1447]" pageId="52" pageNumber="53">
<emphasis id="70E9EACC62559D26EE9CFA1EFED9EFA6" box="[199,256,1423,1447]" italics="true" pageId="52" pageNumber="53">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="422236DE62559D26EE9CFA25FE1BEFCA" blockId="52.[151,1437,151,2024]" box="[199,450,1459,1483]" pageId="52" pageNumber="53">
<emphasis id="70E9EACC62559D26EE9CFA25FEE7EFCA" box="[199,318,1460,1483]" italics="true" pageId="52" pageNumber="53">Immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562559D26EE9CFA47FE16EFEE" box="[199,463,1494,1519]" pageId="52" pageNumber="53" type="biology_ecology">
<paragraph id="422236DE62559D26EE9CFA47FE16EFEE" blockId="52.[151,1437,151,2024]" box="[199,463,1494,1519]" pageId="52" pageNumber="53">
<emphasis id="70E9EACC62559D26EE9CFA47FE93EFEE" bold="true" box="[199,330,1494,1519]" pageId="52" pageNumber="53">BIOLOGY</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562559D26EE9CFA6BFBA1EC12" box="[199,1144,1530,1555]" pageId="52" pageNumber="53" type="distribution">
<paragraph id="422236DE62559D26EE9CFA6BFBA1EC12" blockId="52.[151,1437,151,2024]" box="[199,1144,1530,1555]" pageId="52" pageNumber="53">
<emphasis id="70E9EACC62559D26EE9CFA6BFE4FEC12" bold="true" box="[199,406,1530,1555]" pageId="52" pageNumber="53">DISTRIBUTION</emphasis>
:
<collectingCountry id="3A8A764E62559D26EFFFFA6AFDF0EC12" box="[420,553,1531,1555]" name="Bangladesh" pageId="52" pageNumber="53">Bangladesh</collectingCountry>
,
<collectingCountry id="3A8A764E62559D26EC6EFA6AFDB7EC12" box="[565,622,1531,1555]" name="India" pageId="52" pageNumber="53">India</collectingCountry>
(
<collectingRegion id="8059F83C62559D26EC26FA6AFD15EC12" box="[637,716,1531,1555]" country="India" name="Assam" pageId="52" pageNumber="53">Assam</collectingRegion>
) and
<collectingCountry id="3A8A764E62559D26ED50FA6AFC97EC12" box="[779,846,1531,1555]" name="Nepal" pageId="52" pageNumber="53">Nepal</collectingCountry>
(
<figureCitation id="DAA62A5B62559D26ED06FA6AFC67EC12" box="[861,958,1531,1555]" captionStart="FIGURE 137139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp &amp; Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="52" pageNumber="53">Fig. 137</figureCitation>
),
<quantity id="85659B3B62559D26ED88FA6AFBADEC12" box="[979,1140,1531,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="2.07" metricValueMin="0.03" pageId="52" pageNumber="53" unit="m" value="1050.0" valueMax="2070.0" valueMin="30.0">302070 masl</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562559D26EE9CF98FFCE6EDE6" pageId="52" pageNumber="53" type="discussion">
<paragraph id="422236DE62559D26EE9CF98FFA5BED7A" blockId="52.[151,1437,151,2024]" pageId="52" pageNumber="53">
<emphasis id="70E9EACC62559D26EE9CF98FFEABEC36" bold="true" box="[199,370,1566,1591]" pageId="52" pageNumber="53">DISCUSSION</emphasis>
:
<materialsCitation id="F2F53C8362559D26EFDFF98EFAA7ED7A" collectionCode="NZSI, TYPE, FEMALE" collectorName="The &amp; A. Chattopadhyay &amp; This &amp; However, I" country="India" county="The" elevation="30" location="The" municipality="Zoological Survey" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" stateProvince="Sylhet" typeStatus="holotype">
<collectingCounty id="AB434E5262559D26EFDFF98EFE76EC36" box="[388,431,1567,1591]" pageId="52" pageNumber="53">The</collectingCounty>
<typeStatus id="9D26887C62559D26EFE0F98EFDC4EC36" box="[443,541,1567,1591]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
is in the collection of the
<collectingMunicipality id="A246ACA462559D26ED07F98EFBF6EC36" box="[860,1071,1567,1591]" pageId="52" pageNumber="53">Zoological Survey</collectingMunicipality>
of
<collectingCountry id="3A8A764E62559D26EA07F98EFB4CEC36" box="[1116,1173,1567,1591]" name="India" pageId="52" pageNumber="53">India</collectingCountry>
(
<collectionCode id="248CAE1B62559D26EAF3F98EFB35EC36" box="[1192,1260,1567,1591]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="52" pageNumber="53">NZSI</collectionCode>
), though I was unable to borrow the specimen.
<location id="4742600562559D26EC78F9D2FD97EC5A" LSID="urn:lsid:plazi:treatment:CA3487C862529D27EECCFACEFE8EEAF6:4742600562559D26EC78F9D2FD97EC5A" box="[547,590,1603,1627]" country="India" county="The" municipality="Zoological Survey" name="The" pageId="52" pageNumber="53" stateProvince="Sylhet">The</location>
type location given for the female
<typeStatus id="9D26887C62559D26EA50F9D2FBB4EC5A" box="[1035,1133,1603,1627]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
is “[
<collectingCountry id="3A8A764E62559D26EAF5F9D2FAE1EC5A" box="[1198,1336,1603,1627]" name="Bangladesh" pageId="52" pageNumber="53">Bangladesh</collectingCountry>
]
<collectingRegion id="8059F83C62559D26EB16F9D2FA4EEC5A" box="[1357,1431,1603,1627]" country="Bangladesh" name="Sylhet" pageId="52" pageNumber="53">Sylhet</collectingRegion>
, 3.ii.[19]05 (Hall). [
<quantity id="85659B3B62559D26EF30F9F6FE46EC7E" box="[363,415,1639,1663]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="53" unit="m" value="30.0">
<elevation id="C9B0D1ED62559D26EF30F9F6FE46EC7E" box="[363,415,1639,1663]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="53" unit="m" value="30.0">30m</elevation>
</quantity>
elevation]”
<collectorName id="EF68530862559D26EC71F9F6FD8CEC7E" box="[554,597,1639,1663]" pageId="52" pageNumber="53">The</collectorName>
specimen is labelled as
<collectionCode id="248CAE1B62559D26ED3EF9F9FC77EC7E" box="[869,942,1640,1663]" pageId="52" pageNumber="53">TYPE</collectionCode>
,
<collectionCode id="248CAE1B62559D26EDE2F9F6FBF5EC7E" box="[953,1068,1639,1663]" pageId="52" pageNumber="53">FEMALE</collectionCode>
, and has three legs and one wing damaged (
<collectorName id="EF68530862559D26EF4BF91AFE38ECA2" box="[272,481,1675,1699]" pageId="52" pageNumber="53">A. Chattopadhyay</collectorName>
, (
<collectionCode id="248CAE1B62559D26EFA2F91AFDE3ECA2" box="[505,570,1675,1699]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="52" pageNumber="53">NZSI</collectionCode>
) pers. comm.).
<collectorName id="EF68530862559D26ECA3F91AFCF0ECA2" box="[760,809,1675,1699]" pageId="52" pageNumber="53">This</collectorName>
species was previously known only from the female
<typeStatus id="9D26887C62559D26EECCF93EFF25ECC6" box="[151,252,1711,1735]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
. I assigned
<specimenCount id="549BFD5762559D26EFDDF93EFDCAECC6" box="[390,531,1711,1735]" pageId="52" pageNumber="53" type="female">three female</specimenCount>
specimens to this species as they were collected close to the type locality and agreed closely with the original description. The only differences noted were: tergite 7 lighter brown in the Nepalese specimens; one Nepalese specimen with a slightly darker head; one Nepalese and the Indian specimen with vein sc-r absent (presence of this wing vein seems to be variable in the
<location id="4742600562559D26EDB7F88AFB40ED32" LSID="urn:lsid:plazi:treatment:CA3487C862529D27EECCFACEFE8EEAF6:4742600562559D26EDB7F88AFB40ED32" box="[1004,1177,1819,1843]" country="India" county="The" municipality="Zoological Survey" name="Oriental region" pageId="52" pageNumber="53" stateProvince="Sylhet">Oriental region</location>
, see
<taxonomicName id="859D4D5D62559D26EA94F88DFA55ED32" authorityName="Borkent" authorityYear="2012" box="[1231,1420,1820,1843]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="titiwangsensis">
<emphasis id="70E9EACC62559D26EA94F88DFA55ED32" box="[1231,1420,1820,1843]" italics="true" pageId="52" pageNumber="53">L. titiwangsensis</emphasis>
</taxonomicName>
). It is possible that these females are actually different species, as females are typically very similar between related species.
<collectorName id="EF68530862559D26EEADF8F2FEABED7B" box="[246,370,1891,1914]" pageId="52" pageNumber="53">However, I</collectorName>
refrained from naming them, as male specimens are needed to confirm genitalic differences
</materialsCitation>
.
</paragraph>
<paragraph id="422236DE62559D26EE9CF816FCE6EDE6" blockId="52.[151,1437,151,2024]" pageId="52" pageNumber="53">
As discussed below in the phylogeny section, the placement of
<taxonomicName id="859D4D5D62559D26EDF3F816FB7CED9E" box="[936,1189,1927,1951]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="70E9EACC62559D26EDF3F816FB93ED9E" box="[936,1098,1927,1951]" italics="true" pageId="52" pageNumber="53">Leptomorphus</emphasis>
species
</taxonomicName>
in subgenera (
<bibRefCitation id="260C4B2F62559D26EB0AF816FF09EDC2" author="Matile, L." pageId="52" pageNumber="53" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
<taxonomicName id="859D4D5D62559D26EECCF85EFEEEEDE6" box="[151,311,1999,2023]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="subGenus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC62559D26EECCF85EFEEEEDE6" box="[151,311,1999,2023]" italics="true" pageId="52" pageNumber="53">Gymnoscutum</emphasis>
</taxonomicName>
and placed solely in the genus
<emphasis id="70E9EACC62559D26ECCEF85EFCE6EDE6" box="[661,831,1999,2023]" italics="true" pageId="52" pageNumber="53">
<taxonomicName id="859D4D5D62559D26ECCEF85EFCE3EDE6" authorityName=": Edwards" authorityYear="1925" box="[661,826,1999,2023]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0A87655562549D27EE9CFF06FE8EEAF6" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="422236DE62549D27EE9CFF06FE8EEAF6" blockId="53.[151,1436,151,247]" pageId="53" pageNumber="54">
<collectionCode id="248CAE1B62549D27EE9CFF06FE86EAB1" box="[199,351,151,176]" pageId="53" pageNumber="54">
<emphasis id="70E9EACC62549D27EE9CFF06FE86EAB1" bold="true" box="[199,351,151,176]" pageId="53" pageNumber="54">MATERIAL</emphasis>
</collectionCode>
<materialsCitation id="F2F53C8362549D27EF37FF06FE8AEAF6" collectingDate="1917-07" collectingDateMax="1967-07-21" collectingDateMin="1917-07" collectionCode="BMNH, CNC" collectorName="S. Kemp." country="India" location="Assam" pageId="53" pageNumber="54" specimenCount="3" specimenCount-female="3" stateProvince="Assam">
<emphasis id="70E9EACC62549D27EF37FF06FDDEEAB1" bold="true" box="[364,519,151,176]" pageId="53" pageNumber="54">EXAMINED</emphasis>
:
<collectingCountry id="3A8A764E62549D27EC47FF06FDB7EAAE" box="[540,622,151,175]" name="India" pageId="53" pageNumber="54">INDIA</collectingCountry>
,
<collectingRegion id="8059F83C62549D27ECDBFF06FD16EAAE" box="[640,719,151,175]" country="India" name="Assam" pageId="53" pageNumber="54">Assam</collectingRegion>
, Above Tura, Garo hills,
<date id="3623101E62549D27EA51FF06FBB1EAAE" box="[1034,1128,151,175]" pageId="53" pageNumber="54" value="1917-07">
<collectingDate id="2667E9F662549D27EA51FF06FBB1EAAE" box="[1034,1128,151,175]" pageId="53" pageNumber="54" value="1917-07">vii.1917</collectingDate>
</date>
,
<collectorName id="EF68530862549D27EA21FF06FB33EAAE" box="[1146,1258,151,175]" pageId="53" pageNumber="54">S. Kemp.</collectorName>
(
<specimenCount id="549BFD5762549D27EB59FF06FAFAEAAE" box="[1282,1315,151,175]" pageId="53" pageNumber="54" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B62549D27EB6CFF06FA49EAAE" box="[1335,1424,151,175]" pageId="53" pageNumber="54">BMNH</collectionCode>
);
<collectingCountry id="3A8A764E62549D27EECCFF2DFF2BEAD5" box="[151,242,188,212]" name="Nepal" pageId="53" pageNumber="54">NEPAL</collectingCountry>
, Kathmandu, Godavaril [= Godawari?],
<date id="3623101E62549D27EC9AFF2DFC9AEAD5" box="[705,835,188,212]" pageId="53" pageNumber="54" value="1967-07-21">
<collectingDate id="2667E9F662549D27EC9AFF2DFC9AEAD5" box="[705,835,188,212]" pageId="53" pageNumber="54" value="1967-07-21">21.vii.1967</collectingDate>
</date>
, Can. Exp. (
<specimenCount id="549BFD5762549D27ED8DFF2DFC2CEAD2" box="[982,1013,188,212]" pageId="53" pageNumber="54" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B62549D27EA5EFF2DFB9AEAD5" box="[1029,1091,188,212]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="53" pageNumber="54">CNC</collectionCode>
); Pulchauki,
<date id="3623101E62549D27EA83FF2DFA83EAD5" box="[1240,1370,188,212]" pageId="53" pageNumber="54" value="1967-07-14">
<collectingDate id="2667E9F662549D27EA83FF2DFA83EAD5" box="[1240,1370,188,212]" pageId="53" pageNumber="54" value="1967-07-14">14.vii.1967</collectingDate>
</date>
, Can. Exp. (
<specimenCount id="549BFD5762549D27EE86FF4EFF27EAF6" box="[221,254,223,247]" pageId="53" pageNumber="54" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B62549D27EF57FF4EFE93EAF6" box="[268,330,223,247]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="53" pageNumber="54">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>