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<document id="73E65700B6C1F7913B11178437FFC36D" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C51581833FF0EFA08EE571E78" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Luciola tiomana Ho 2019, sp. nov." docType="treatment" docVersion="19" lastPageNumber="102" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="100" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="4195ED853E5F34F587F96E8A1521D023">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="13A4D60570D491BB8C7688FB7249D0F8">Ho, - Z.</mods:namePart>
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<mods:date id="707636B9AEC76BD5013A832251709805">2019</mods:date>
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<mods:identifier id="E62B8CA0326E9448C74AE112BC322D65" type="DOI">10.11646/zootaxa.4687.1.1</mods:identifier>
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<treatment id="6C3DA91C51581833FF0EFA08EE571E78" ID-DOI="http://doi.org/10.5281/zenodo.4688893" ID-GBIF-Taxon="159757392" ID-Zenodo-Dep="4688893" LSID="urn:lsid:plazi:treatment:6C3DA91C51581833FF0EFA08EE571E78" httpUri="http://treatment.plazi.org/id/6C3DA91C51581833FF0EFA08EE571E78" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100">
<subSubSection id="AC8E4B8151581835FF0EFA08ED6A19F5" box="[151,594,1431,1458]" pageId="99" pageNumber="100" type="nomenclature">
<paragraph id="E42B180A51581835FF0EFA08ED6A19F5" blockId="99.[151,594,1431,1494]" box="[151,594,1431,1458]" pageId="99" pageNumber="100">
<heading id="BF63AF6651581835FF0EFA08ED6A19F5" bold="true" box="[151,594,1431,1458]" fontSize="11" level="1" pageId="99" pageNumber="100" reason="1">
<emphasis id="D6E0C41851581835FF0EFA08ED6A19F5" bold="true" box="[151,594,1431,1458]" pageId="99" pageNumber="100">
<taxonomicName id="2394638951581835FF0EFA08EED519F5" ID-CoL="8NBWZ" authority="Ballantyne" authorityName="Ho" authorityYear="2019" box="[151,493,1431,1458]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="tiomana" status="sp. nov.">
<emphasis id="D6E0C41851581835FF0EFA08EE6619F5" bold="true" box="[151,350,1431,1458]" italics="true" pageId="99" pageNumber="100">Luciola tiomana</emphasis>
Ballantyne
</taxonomicName>
<taxonomicNameLabel id="CDD3796351581835FE6CFA07ED6A19F5" box="[501,594,1432,1458]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151581835FF0EFA23EE501A33" pageId="99" pageNumber="100" type="description">
<paragraph id="E42B180A51581835FF0EFA23EE091991" blockId="99.[151,594,1431,1494]" box="[151,305,1468,1494]" pageId="99" pageNumber="100">
<figureCitation id="7CAF048F51581835FF0EFA23EE091991" box="[151,305,1468,1494]" captionStart="FIGURES 309314" captionStartId="100.[151,264,1754,1779]" captionTargetBox="[152,1435,166,1723]" captionTargetId="figure@100.[151,1435,166,1734]" captionTargetPageId="100" captionText="FIGURES 309314. Luciola tiomana sp. nov. type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="99" pageNumber="100">Figs 309314</figureCitation>
</paragraph>
<paragraph id="E42B180A51581835FF0EF99FEE501A33" blockId="99.[151,1436,1535,1652]" pageId="99" pageNumber="100">
<emphasis id="D6E0C41851581835FF0EF99FEFE41A5F" bold="true" box="[151,220,1536,1560]" pageId="99" pageNumber="100">Types.</emphasis>
<materialsCitation id="54FC125751581835FF7EFA60EE501A33" ID-GBIF-Occurrence="2427485588" collectingDate="2009-03-29" collectingDateMax="2016-06-29" collectingDateMin="2009-03-29" collectionCode="FRIM" collectorName="V Hula &amp; Compt. &amp; B. Nada" country="Malaysia" county="Tioman Isl." elevation="305" location="Pasir Raja Selatan Forest Reserve" municipality="Isl. Jungle" pageId="99" pageNumber="100" specimenCode="PS52" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="holotype">
<typeStatus id="3B2FA6A851581835FF7EFA60EE7D1A5F" box="[231,325,1535,1560]" pageId="99" pageNumber="100" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="3B2FA6A851581835FE0BF99FEED11A5F" box="[402,489,1536,1560]" pageId="99" pageNumber="100" type="paratype">paratype</typeStatus>
male.
<emphasis id="D6E0C41851581835FDAFFA60EDFC1A5F" bold="true" box="[566,708,1535,1560]" pageId="99" pageNumber="100">
<collectingCountry id="9C83589A51581835FDAFFA60EDF81A5F" box="[566,704,1535,1560]" name="Malaysia" pageId="99" pageNumber="100">MALAYSIA</collectingCountry>
.
</emphasis>
<emphasis id="D6E0C41851581835FD56F99FEC321A5F" box="[719,778,1536,1560]" italics="true" pageId="99" pageNumber="100">
<collectingRegion id="2650D6E851581835FD56F99FEC321A5F" box="[719,778,1536,1560]" country="Malaysia" name="Johor" pageId="99" pageNumber="100">Johor</collectingRegion>
</emphasis>
:
<collectingCounty id="0D4A608651581835FC82FA60ECAC1A5F" box="[795,916,1535,1560]" pageId="99" pageNumber="100">
Tioman
<collectingMunicipality id="044F827051581835FCEDFA60ECAC1A5F" box="[884,916,1535,1560]" pageId="99" pageNumber="100">Isl.</collectingMunicipality>
</collectingCounty>
Jungle track
<quantity id="236CB5EF51581835FBB1FA60EB971A5F" box="[1064,1199,1535,1560]" metricMagnitude="2" metricUnit="m" metricValue="3.05" metricValueMax="6.0" metricValueMin="0.1" pageId="99" pageNumber="100" unit="m" value="305.0" valueMax="600.0" valueMin="10.0">
<elevation id="6FB9FF3951581835FBB1FA60EB971A5F" box="[1064,1199,1535,1560]" metricMagnitude="2" metricUnit="m" metricValue="3.05" metricValueMax="6.0" metricValueMin="0.1" pageId="99" pageNumber="100" unit="m" value="305.0" valueMax="600.0" valueMin="10.0">10600 masl</elevation>
</quantity>
15
<date id="902A3ECA51581835FB45FA60EA7C1A5F" box="[1244,1348,1535,1560]" pageId="99" pageNumber="100" value="2009-03-29">
<collectingDate id="806EC72251581835FB45FA60EA7C1A5F" box="[1244,1348,1535,1560]" pageId="99" pageNumber="100" value="2009-03-29">29.3.2009</collectingDate>
</date>
<collectorName id="49617DDC51581835FAD7F99FEAA41A5F" box="[1358,1436,1535,1560]" pageId="99" pageNumber="100">V Hula</collectorName>
(Prague).
<emphasis id="D6E0C41851581835FEB5F980EE9B1A71" box="[300,419,1567,1590]" italics="true" pageId="99" pageNumber="100">
<collectingRegion id="2650D6E851581835FEB5F980EE9B1A71" box="[300,419,1567,1590]" country="Malaysia" name="Terengganu" pageId="99" pageNumber="100">Terengganu</collectingRegion>
</emphasis>
:
<location id="E14B4ED151581835FE28F981EC351A71" LSID="urn:lsid:plazi:treatment:6C3DA91C51581833FF0EFA08EE571E78:E14B4ED151581835FE28F981EC351A71" box="[433,781,1566,1591]" country="Malaysia" county="Tioman Isl." municipality="Isl. Jungle" name="Pasir Raja Selatan Forest Reserve" pageId="99" pageNumber="100" stateProvince="Johor">Pasir Raja Selatan Forest Reserve</location>
,
<collectorName id="49617DDC51581835FC81F980EC5D1A70" box="[792,869,1567,1591]" pageId="99" pageNumber="100">Compt.</collectorName>
<date id="902A3ECA51581835FCF2F981ECF11A70" box="[875,969,1566,1591]" pageId="99" pageNumber="100" value="1952-06-29">
52,
<collectingDate id="806EC72251581835FC0BF981ECF11A70" box="[914,969,1566,1591]" pageId="99" pageNumber="100" value="2016-06-29">29.vi.</collectingDate>
</date>
2016,
<collectorName id="49617DDC51581835FB95F980EB5A1A70" box="[1036,1122,1566,1591]" pageId="99" pageNumber="100">B. Nada</collectorName>
,
<specimenCount id="F292D38351581835FBF4F980EBB81A71" box="[1133,1152,1567,1590]" pageId="99" pageNumber="100" type="male"></specimenCount>
(
<specimenCode id="B432B07151581835FB17F981EBFD1A70" box="[1166,1221,1566,1591]" collectionCode="PS" pageId="99" pageNumber="100">
<collectionCode id="828580CF51581835FB17F981EB931A70" box="[1166,1195,1566,1591]" pageId="99" pageNumber="100">PS</collectionCode>
52
</specimenCode>
[4]) (
<collectionCode id="828580CF51581835FA98F980EA7D1A70" box="[1281,1349,1567,1591]" collectionName="Malaysia, Kuala Lumpur, Forest Research Institute" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="99" pageNumber="100">FRIM</collectionCode>
). Nadas collection in forest reserve has compartment number and transect number following in (). # = killed fixed and preserved in 70% ethanol.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151581835FF0EF903ED3C1B64" pageId="99" pageNumber="100" type="diagnosis">
<paragraph id="E42B180A51581835FF0EF903ED3C1B64" blockId="99.[151,1437,1692,2043]" pageId="99" pageNumber="100">
<emphasis id="D6E0C41851581835FF0EF903EE321AF0" bold="true" box="[151,266,1692,1719]" pageId="99" pageNumber="100">Diagnosis</emphasis>
. A small species (4.24.3 mm long) with distinctive dorsal colouration of black head, pronotum, dark brown to black MN and MS, and brownish yellow elytra which have diffuse darker brown markings in apical ¾, with paler basal markings restricted to basal area near suture, and an accumulation of whitish fat body along apex of suture and round elytral apex.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151581833FF5EF8B3EE3F1D20" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100" type="description">
<paragraph id="E42B180A51581833FF5EF8B3EE3F1D20" blockId="99.[151,1437,1692,2043]" lastBlockId="101.[151,1437,153,575]" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100">
<emphasis id="D6E0C41851581835FF5EF8B3EE8B1B00" bold="true" box="[199,435,1836,1863]" pageId="99" pageNumber="100">Description of Male</emphasis>
. 4.24.3 mm long. Colour (
<figureCitation id="7CAF048F51581835FD70F8B2ECB21B00" box="[745,906,1837,1863]" captionStart="FIGURES 309314" captionStartId="100.[151,264,1754,1779]" captionTargetBox="[152,1435,166,1723]" captionTargetId="figure@100.[151,1435,166,1734]" captionTargetPageId="100" captionText="FIGURES 309314. Luciola tiomana sp. nov. type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="99" pageNumber="100">Figs 309313</figureCitation>
): pronotum black with narrow brownish orange band across posterior margin; MN and MS black in type, MS with narrow irregular paler brown lateral margins; MN and MS brownish orange in
<typeStatus id="3B2FA6A851581835FD93F8EAED411BC8" box="[522,633,1909,1935]" pageId="99" pageNumber="100" type="paratype">paratypes</typeStatus>
; elytra of
<typeStatus id="3B2FA6A851581835FD73F8EAEC741BC8" box="[746,844,1909,1935]" pageId="99" pageNumber="100" type="holotype">holotype</typeStatus>
with basal ¼ and suture and lateral margin narrowly brownish orange; rest of elytron in type is diffuse dark brown with apices dark; elytra of
<typeStatus id="3B2FA6A851581835FB1BF806EBDA1BF4" box="[1154,1250,1945,1971]" pageId="99" pageNumber="100" type="paratype">paratype</typeStatus>
with basal paler colour occupying slightly less than basal ¼ and brownish orange colour of suture and lateral margin expanding around apex; head antennae palpi black, scape and pedicel paler on underside; venter of thorax and all ventrites anterior to the LO very dark brown to black; Legs brownish orange; LO in
<date id="902A3ECA515F1832FC54F8DBECC91B19" box="[973,1009,1860,1886]" pageId="100" pageNumber="101">V6</date>
, 7 pale creamy white; T25 semitransparent grey, T6, 7 black, T8 semitransparent, pale. Pronotum: slightly narrower than humeral width; sides subparallel (A = B = C); posterolateral corners acute. Elytron: subparallel sided; interstitial lines not well defined. Head: GHW
<date id="902A3ECA515F1832FF7BF82EEE2D1B8C" box="[226,277,1969,1995]" pageId="100" pageNumber="101" value="2010-10">10 x</date>
SIW; ASD &lt;ASW; small eye emargination visible along posterior margin of eye (not visible clearly from above); eyes very close but not contiguous on ventral surface. Antennae: very slightly longer than GHW, segments simple scape longer than remainder, scape and pedicel thicker than remainder; FS 48 short subequal in length, FS 9 slightly longer than 8. Abdomen (
<figureCitation id="7CAF048F515E1833FD39FF22EC781C90" box="[672,832,189,215]" captionStart="FIGURES 309314" captionStartId="100.[151,264,1754,1779]" captionTargetBox="[152,1435,166,1723]" captionTargetId="figure@100.[151,1435,166,1734]" captionTargetPageId="100" captionText="FIGURES 309314. Luciola tiomana sp. nov. type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="101" pageNumber="102">Figs 312, 313</figureCitation>
): LO occupying all of
<date id="902A3ECA515E1833FBA5FF22EB9B1C90" box="[1084,1187,189,215]" pageId="101" pageNumber="102">V6 and 7</date>
with posterior margin of
<date id="902A3ECA515E1833FF2CFF7EEFEF1CBC" box="[181,215,225,251]" pageId="101" pageNumber="102">V7</date>
rounded, MPP not developed. T8 with median posterior margin slightly produced and rounded; anterolateral corners narrowly produced and apically acute, length less than posterior entire portion of T8. Aedeagal sheath and aedeagus (
<figureCitation id="7CAF048F515E1833FE94FEB6EE521D04" box="[269,362,297,323]" captionStart="FIGURES 309314" captionStartId="100.[151,264,1754,1779]" captionTargetBox="[152,1435,166,1723]" captionTargetId="figure@100.[151,1435,166,1734]" captionTargetPageId="100" captionText="FIGURES 309314. Luciola tiomana sp. nov. type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="101" pageNumber="102">Fig 314</figureCitation>
): sheath sternite with entire rounded apex; median anterior margin of tergite slightly and evenly produced.
</paragraph>
</subSubSection>
<caption id="B0EB4882515F1832FF0EF945EA631B51" ID-DOI="http://doi.org/10.5281/zenodo.3508222" ID-Zenodo-Dep="3508222" httpUri="https://zenodo.org/record/3508222/files/figure.png" pageId="100" pageNumber="101" startId="100.[151,264,1754,1779]" targetBox="[152,1435,166,1723]" targetPageId="100">
<paragraph id="E42B180A515F1832FF0EF945EA631B51" blockId="100.[151,1436,1754,1815]" pageId="100" pageNumber="101">
<emphasis id="D6E0C418515F1832FF0EF945EE571AB4" bold="true" box="[151,367,1754,1779]" pageId="100" pageNumber="101">FIGURES 309314.</emphasis>
<taxonomicName id="23946389515F1832FEEFF945ED251AB5" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[374,541,1754,1778]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="tiomana" status="sp. nov.">
<emphasis id="D6E0C418515F1832FEEFF945ED251AB5" box="[374,541,1754,1778]" italics="true" pageId="100" pageNumber="101">Luciola tiomana</emphasis>
</taxonomicName>
<emphasis id="D6E0C418515F1832FDBDF945ED4C1AB5" bold="true" box="[548,628,1754,1778]" pageId="100" pageNumber="101">
<taxonomicNameLabel id="CDD37963515F1832FDBDF945ED4C1AB5" box="[548,628,1754,1778]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
type males. 309, 310 habitus dorsal (309 Prague; 310 FRIM); 311 habitus ventral; 312, 313 abdomen: 312 ventral; 313 dorsal; 314 tergite 8, aedeagal sheath and aedeagus, dorsal. Scale line is 1 mm.
</paragraph>
</caption>
<subSubSection id="AC8E4B81515E1833FF5EFEEEEBA41DCC" box="[199,1180,369,395]" pageId="101" pageNumber="102" type="etymology">
<paragraph id="E42B180A515E1833FF5EFEEEEBA41DCC" blockId="101.[151,1437,153,575]" box="[199,1180,369,395]" pageId="101" pageNumber="102">
<emphasis id="D6E0C418515E1833FF5EFEEEEE761DCC" bold="true" box="[199,334,369,395]" pageId="101" pageNumber="102">Etymology.</emphasis>
The
<typeStatus id="3B2FA6A8515E1833FE1EFEEEEE8F1DCC" box="[391,439,369,395]" pageId="101" pageNumber="102">type</typeStatus>
locality is latinised thus Tioman to tiomana, a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B81515E1833FF5EFE0AEE571E78" pageId="101" pageNumber="102" type="discussion">
<paragraph id="E42B180A515E1833FF5EFE0AEE571E78" blockId="101.[151,1437,153,575]" pageId="101" pageNumber="102">
<emphasis id="D6E0C418515E1833FF5EFE0AEE031DE8" bold="true" box="[199,315,405,431]" pageId="101" pageNumber="102">Remarks.</emphasis>
The similarly coloured
<taxonomicName id="23946389515E1833FDD1FE0AECB51DE8" authority="Pic 1914" authorityName="Pic" authorityYear="1914" box="[584,909,405,431]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="notaticollis">
<emphasis id="D6E0C418515E1833FDD1FE0AEC1B1DE8" box="[584,803,405,431]" italics="true" pageId="101" pageNumber="102">Luciola notaticollis</emphasis>
<bibRefCitation id="800565FB515E1833FCB3FE0AECB51DE8" author="Pic" box="[810,909,405,431]" firstAuthor="Pic" pageId="101" pageNumber="102" pagination="3" refId="ref121203" refString="Pic, M. (1914) Nouveau genre, especes et varietes nouvelles de diverses familles. Melanges exotico-Entomologiques, 10, 3." type="journal article" year="1914">Pic 1914</bibRefCitation>
</taxonomicName>
from Madura
<collectingCountry id="9C83589A515E1833FBA9FE0AEB521DE8" box="[1072,1130,405,431]" name="India" pageId="101" pageNumber="102">India</collectingCountry>
is 7.0 mm long, with black pronotum, MN, and MS, and the basal area of the elytra (slightly less than half the length) orange, with the apical ½ of the elytron black. The
<typeStatus id="3B2FA6A8515E1833FE63FE42ED121DB0" box="[506,554,477,503]" pageId="101" pageNumber="102">type</typeStatus>
in MNHN was examined but not dissected as it is glued face down on a card.
<bibRefCitation id="800565FB515E1833FF0EFD9EEEF01E5B" author="Ballantyne &amp; Jusoh" box="[151,456,513,540]" firstAuthor="Ballantyne" pageId="101" pageNumber="102" pagination="1 - 7" refId="ref114336" refString="Ballantyne, L. A &amp; Jusoh, W. F. A. (2016) List of genera and species in the Luciolinae from SE Asia and the Australopacific Version 2. Checklist of IndoPacific Luciolinae, 2016, 1 - 7. https: // doi. org / 10.13140 / RG. 2.1.4635.2241" type="journal article" year="2016">Ballantyne &amp; Jusoh (2016)</bibRefCitation>
suggested a closer examination was necessary to determine if
<taxonomicName id="23946389515E1833FB15FD9DEA101E5C" authorityName="Pic" authorityYear="1914" box="[1164,1320,513,539]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="notaticollis">
<emphasis id="D6E0C418515E1833FB15FD9DEA101E5C" box="[1164,1320,513,539]" italics="true" pageId="101" pageNumber="102">L. notaticollis</emphasis>
</taxonomicName>
should be assigned to
<taxonomicName id="23946389515E1833FE82FDBAEE5F1E78" box="[283,359,549,575]" class="Insecta" family="Lampyridae" genus="Curtos" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418515E1833FE82FDBAEE5F1E78" box="[283,359,549,575]" italics="true" pageId="101" pageNumber="102">Curtos</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>