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<document id="64CC9E0B92909E3D0A7ACDC84F3F05C3" ID-DOI="10.5252/zoosystema2025v47a4" ID-ISSN="1638-9387" ID-Zenodo-Dep="14831210" ID-ZooBank="3749B35A-48E6-43F6-B889-AC340DBD5CF2" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1738917347317" checkinUser="plazi" docAuthor="Tran, Ngat Thi, Phuong, Lien Thi, Anh, Phuong Nguyen, Nguyen, Anh Duc &amp; Engel, Michael S." docDate="2025" docId="03BA0E159478FFFDFF6F3C5EC13B4DA1" docLanguage="en" docName="zoosystema.2025.47.4.pdf" docOrigin="Zoosystema 47 (4)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a4.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.10:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="10" docTitle="Maculonomia (Maculonomia) ferruginea Engel, Tran, &amp; Nguyen 2025, n. sp." docType="treatment" docVersion="1" lastPageNumber="67" masterDocId="FF83766D9468FFEFFF883A2CC32B4B6D" masterDocTitle="New species and records of the nomiine bee genus Maculonomia Wu, 1982 from Vietnam (Hymenoptera, Halictidae)" masterLastPageNumber="73" masterPageNumber="51" pageNumber="65" updateTime="1739212110286" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B2561AED8D38E3225BDDBF9B7701D0A5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CCE3E34AD13B46377E09AB21A064C250">
<mods:title id="F3967190B5063E7CBAD7034D435B3866">New species and records of the nomiine bee genus Maculonomia Wu, 1982 from Vietnam (Hymenoptera, Halictidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="6C443F73F66F6CFBEE05B5E7074752B9">Tran, Ngat Thi</mods:namePart>
<mods:affiliation id="89F78F922E18376B91647492BFD97F35">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="976BE6AED85E6F9CB3131A3B024C2388" type="email">tranthingat1012@gmail.com</mods:nameIdentifier>
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<mods:role id="C731094D8A6A7F476B2396C2281223BE">
<mods:roleTerm id="BC106F9ADBB0DFA782E381B60A3380AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAD45104E92C6616F11727FBE5D20148">Phuong, Lien Thi</mods:namePart>
<mods:affiliation id="2682ECB7DDA47DA99BDE5033004834AB">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="CCC959759775E75E2A4A53241608FA17" type="email">phuonglientit@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7502BCD0D442C8E18A5194DCBA0F88F8" type="personal">
<mods:role id="F7318274D1FA7C905A22A274B5B5B6E2">
<mods:roleTerm id="5A94ED4AE8B5874B32FEE8694C31574F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E286126ABA4C58EAAF0CCFA7B23694EA">Anh, Phuong Nguyen</mods:namePart>
<mods:affiliation id="BFC98C2E95CF845E06C7F199DF6A70C5">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="2376F9DB9622ECE14DF2FDFD1CB527C0" type="email">phuonglientit@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="101FAEA5D467FDB6AB41DF3F579A1AF6" type="personal">
<mods:role id="8CF8C916C0066255611FA4975021FC10">
<mods:roleTerm id="CE054A134F7FDD102133BB342BF36AB2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B93A9024643CA89EF9E0A34D6C8E476">Nguyen, Anh Duc</mods:namePart>
<mods:affiliation id="82A0D0D93D4A9880CE0B9726DD35E52C">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) and Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="BED568117533F6DA8206735A1C7D5948" type="email">ducanh410@yahoo.com</mods:nameIdentifier>
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<mods:name id="5E40C3C0B9FFD1EA0D46D44FB70010B4" type="personal">
<mods:role id="1602EC5EC9DD3543D9C063DE3A54B478">
<mods:roleTerm id="1E84A8FCC1800EC72ED10714C93ADDED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74FA9BE109ED699A3FBD94CF68EF5268">Engel, Michael S.</mods:namePart>
<mods:affiliation id="0F3C19E024D9212A075737824689128E">Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, New York 10024 (United States)</mods:affiliation>
<mods:nameIdentifier id="864A8BD67DF03C982F9E7FCBE667E0CC" type="email">mengel@amnh.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3AF4D82C18921B0477887FAFB4E11E9E">text</mods:typeOfResource>
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<mods:titleInfo id="203C1C0638CFE4B58F5349049D250540">
<mods:title id="9C914483288EF46D8B54CEB715F3BFBA">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="809F8B92204FCFD93210A967A6DC41F8">
<mods:date id="57B3464EEFE30F1F22CE4DF28A97C05C">2025</mods:date>
<mods:detail id="1E4D8D31EF35BF6160A4A1484CFFD37A" type="pubDate">
<mods:number id="B21E5B6FC17A3A9848FD7A2B1809017F">2025-02-06</mods:number>
</mods:detail>
<mods:detail id="7D81E5635EA7BDE68E973B9A98D1D4FE" type="volume">
<mods:number id="633BABF43E0DC936806B2C2BEB3B2717">47</mods:number>
</mods:detail>
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<mods:number id="02EBB95E8FC4AD73AEDF3A626D0E3DFD">4</mods:number>
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<mods:start id="040ADC0440D57D4EDBB1B5F3BD4248E9">51</mods:start>
<mods:end id="258483B7FA8EF926F0DE9BE3A3F1636B">73</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="94270F8A1C5D536C7296C52808FA4371">
<mods:url id="012F73681D8E97FFBF841B61A5DF063C">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a4.pdf</mods:url>
</mods:location>
<mods:classification id="156F1F8C56A887050FE33C44C9D3240E">journal article</mods:classification>
<mods:identifier id="DB5376BC6D2BD0925D4A32072E1E68F6" type="DOI">10.5252/zoosystema2025v47a4</mods:identifier>
<mods:identifier id="27C7856579B573951B3A55BE3A91A622" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="C69DA457E93DDD9F03A2B08FEA51C06A" type="Zenodo-Dep">14831210</mods:identifier>
<mods:identifier id="76248817D320C94AB5B82ADD8E5EE505" type="ZooBank">3749B35A-48E6-43F6-B889-AC340DBD5CF2</mods:identifier>
</mods:mods>
<treatment id="03BA0E159478FFFDFF6F3C5EC13B4DA1" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1" httpUri="http://treatment.plazi.org/id/03BA0E159478FFFDFF6F3C5EC13B4DA1" lastPageId="18" lastPageNumber="67" pageId="16" pageNumber="65">
<subSubSection id="C309EC889478FFFFFF6F3C5EC1414DC0" pageId="16" pageNumber="65" type="nomenclature">
<paragraph id="8BACBF039478FFFFFF6F3C5EC1414DC0" blockId="16.[231,676,1650,1740]" pageId="16" pageNumber="65">
<heading id="D0E4086F9478FFFFFF6F3C5EC1414DC0" centered="true" fontSize="11" level="2" pageId="16" pageNumber="65" reason="3">
<taxonomicName id="4C13C4809478FFFFFF6F3C5EC10F4DC0" authority="Engel, Tran, &amp; Nguyen" authorityName="Engel, Tran, &amp; Nguyen" authorityYear="2025" class="Insecta" family="Halictidae" genus="Maculonomia" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="ferruginea" status="n. sp." subGenus="Maculonomia">
<emphasis id="B96763119478FFFFFF6F3C5EC2554DE0" bold="true" box="[231,382,1650,1677]" italics="true" pageId="16" pageNumber="65">Maculonomia</emphasis>
(
<emphasis id="B96763119478FFFFFE063C5EC10E4DE0" bold="true" box="[398,549,1650,1677]" italics="true" pageId="16" pageNumber="65">Maculonomia</emphasis>
)
<emphasis id="B96763119478FFFFFDBD3C5EC18F4DE1" bold="true" box="[565,676,1650,1676]" italics="true" pageId="16" pageNumber="65">ferruginea</emphasis>
Engel, Tran, &amp; Nguyen
</taxonomicName>
,
<taxonomicNameLabel id="A254DE6A9478FFFFFDA73CBFC1414DC0" box="[559,618,1683,1709]" pageId="16" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309EC889478FFFFFE093C9EC1C04C65" pageId="16" pageNumber="65" type="description">
<paragraph id="8BACBF039478FFFFFE093C9EC1214DA1" blockId="16.[231,676,1650,1740]" box="[385,522,1714,1740]" pageId="16" pageNumber="65">
(
<figureCitation id="1328A3869478FFFFFE003C9EC2F24DA1" box="[392,473,1714,1740]" captionStart="FIG" captionStartId="17.[156,167,1721,1739]" captionTargetBox="[258,1342,215,1676]" captionTargetId="figure-130@17.[246,1342,215,1677]" captionTargetPageId="17" captionText="FIG. 11. — Maculonomia (Maculonomia) ferruginea Engel, Tran, &amp; Nguyen, n. sp., holotype, female: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831238" httpUri="https://zenodo.org/record/14831238/files/figure.png" pageId="16" pageNumber="65">Figs 11</figureCitation>
;
<figureCitation id="1328A3869478FFFFFE6D3C9EC1294DA1" box="[485,514,1714,1740]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">12</figureCitation>
)
</paragraph>
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<uri id="FF82B3019478FFFFFF283CDAC1C04C65" box="[160,747,1782,1800]" pageId="16" pageNumber="65">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C309EC889478FFFFFF0C3D1EC1834C85" pageId="16" pageNumber="65" type="materials_examined">
<paragraph id="8BACBF039478FFFFFF0C3D1EC1564C13" blockId="16.[131,776,1842,2025]" pageId="16" pageNumber="65">
<smallCapsWord id="8D4A29DF9478FFFFFF0C3D1EC39E4C25" baselines="1859,1860" box="[132,181,1842,1865]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="16" pageNumber="65">
<collectionCode id="ED0227C69478FFFFFF0C3D1EC39E4C25" box="[132,181,1842,1865]" pageId="16" pageNumber="65">TYPE</collectionCode>
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. —
<materialsCitation id="3B7BB55E9478FFFFFEC83D1EC1564C13" collectingDate="2019-05-09" collectionCode="IEBR" collectorName="Nguyen L. T. P. &amp; Nguyen C. Q." country="Vietnam" elevation="1586" latitude="21.384277" location="Y Ty" longLatPrecision="1" longitude="105.71183" municipality="Bat Xat" pageId="16" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="holotype">
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<typeStatus id="54A801A19478FFFFFEC83D1EC2B54C27" box="[320,414,1842,1866]" pageId="16" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
.
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<collectingCountry id="F304FF939478FFFFFE223D1FC12B4C26" box="[426,512,1843,1867]" name="Vietnam" pageId="16" pageNumber="65">Vietnam</collectingCountry>
</emphasis>
•
<specimenCount id="9D15748A9478FFFFFD9B3D19C10F4C24" box="[531,548,1845,1865]" count="1" pageId="16" pageNumber="65" type="female">♀</specimenCount>
;
<collectingRegion id="49D771E19478FFFFFDA73D1EC1564C27" box="[559,637,1842,1866]" country="Vietnam" name="Lao Cai" pageId="16" pageNumber="65">Lao Cai</collectingRegion>
,
<collectingMunicipality id="6BC825799478FFFFFD0D3D1EC1FB4C24" box="[645,720,1842,1865]" pageId="16" pageNumber="65">Bat Xat</collectingMunicipality>
,
<location id="8ECCE9D89478FFFFFD503D1EC02F4C27" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFD503D1EC02F4C27" box="[728,772,1842,1866]" country="Vietnam" latitude="21.384277" longLatPrecision="1" longitude="105.71183" municipality="Bat Xat" name="Y Ty" pageId="16" pageNumber="65" stateProvince="Lao Cai">Y Ty</location>
, way to the red waterfall;
<geoCoordinate id="EE27D9C49478FFFFFEE03D61C2D94C08" box="[360,498,1869,1893]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="16" pageNumber="65" precision="1" seconds="03.4" value="21.384277">21°23’03.4”N</geoCoordinate>
,
<geoCoordinate id="EE27D9C49478FFFFFE723D60C1A14C09" box="[506,650,1868,1892]" degrees="105" direction="east" minutes="42" orientation="longitude" pageId="16" pageNumber="65" precision="1" seconds="42.6" value="105.71183">105°42’42.6”E</geoCoordinate>
; alt.
<quantity id="4CEB12E69478FFFFFD3E3D60C0284C09" box="[694,771,1868,1892]" metricMagnitude="3" metricUnit="m" metricValue="1.5859999999999999" pageId="16" pageNumber="65" unit="m" value="1586.0">
<elevation id="003E58309478FFFFFD3E3D60C0284C09" box="[694,771,1868,1892]" metricMagnitude="3" metricUnit="m" metricValue="1.5859999999999999" pageId="16" pageNumber="65" unit="m" value="1586.0">1586 m</elevation>
</quantity>
;
<date id="FFAD99C39478FFFFFF0B3D4BC3F44C12" box="[131,223,1895,1919]" pageId="16" pageNumber="65" value="2019-05-09">
<collectingDate id="EFE9602B9478FFFFFF0B3D4BC3F44C12" box="[131,223,1895,1919]" pageId="16" pageNumber="65" value="2019-05-09">9.V.2019</collectingDate>
</date>
;
<collectorName id="26E6DAD59478FFFFFF613D4BC2524C13" box="[233,377,1895,1919]" pageId="16" pageNumber="65">Nguyen L.T.P.</collectorName>
,
<collectorName id="26E6DAD59478FFFFFE0B3D4BC1214C12" box="[387,522,1895,1919]" pageId="16" pageNumber="65">Nguyen C.Q.</collectorName>
leg.;
<collectionCode id="ED0227C69478FFFFFDB73D4BC1524C13" box="[575,633,1895,1918]" pageId="16" pageNumber="65">IEBR</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3DADC1834C85" blockId="16.[131,776,1842,2025]" pageId="16" pageNumber="65">
<materialsCitation id="3B7BB55E9478FFFFFF0C3DADC18E4CF7" box="[132,677,1921,1947]" collectingDate="2019-05-09" collectionCode="AMNH" collectorName="Nguyen L. T. P. &amp; Nguyen C. Q." country="Vietnam" elevation="1586" latitude="21.384277" location="Vietnam" longLatPrecision="1" longitude="105.71183" municipality="Bat Xat" pageId="16" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<emphasis id="B96763119478FFFFFF0C3DADC3C94CF4" bold="true" box="[132,226,1921,1945]" pageId="16" pageNumber="65">
<typeStatus id="54A801A19478FFFFFF0C3DADC3C94CF4" box="[132,226,1921,1945]" pageId="16" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="B96763119478FFFFFF653DAFC2694CF6" bold="true" box="[237,322,1923,1947]" pageId="16" pageNumber="65">
<collectingCountry id="F304FF939478FFFFFF653DAFC2694CF6" box="[237,322,1923,1947]" name="Vietnam" pageId="16" pageNumber="65">Vietnam</collectingCountry>
</emphasis>
•
<specimenCount id="9D15748A9478FFFFFEDD3DADC25E4CF5" box="[341,373,1921,1945]" count="1" pageId="16" pageNumber="65" type="female">1♀</specimenCount>
; same data as holotype;
<collectionCode id="ED0227C69478FFFFFDDF3DAEC18E4CF7" box="[599,677,1922,1946]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="16" pageNumber="65">AMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7BB55E9478FFFFFD303DADC2D64CA3" collectingDate="2020-10-27" collectorName="Tran N. T." elevation="1264" latitude="22.328056" location="Sa Pa" longLatPrecision="21" longitude="103.831665" municipality="Sa Pa" pageId="16" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="9D15748A9478FFFFFD303DADC1F34CF5" box="[696,728,1921,1945]" count="1" pageId="16" pageNumber="65" type="female">1♀</specimenCount>
;
<collectingRegion id="49D771E19478FFFFFD6B3DAEC3834CD9" country="Vietnam" name="Lao Cai" pageId="16" pageNumber="65">Lao Cai</collectingRegion>
,
<collectingMunicipality id="6BC825799478FFFFFF383DB0C3CD4CDE" box="[176,230,1948,1972]" pageId="16" pageNumber="65">Sa Pa</collectingMunicipality>
,
<location id="8ECCE9D89478FFFFFF663DB0C20B4CDE" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFF663DB0C20B4CDE" box="[238,288,1948,1972]" latitude="22.328056" longLatPrecision="21" longitude="103.831665" municipality="Sa Pa" name="Sa Pa" pageId="16" pageNumber="65" stateProvince="Lao Cai">Sa Pa</location>
town,
<location id="8ECCE9D89478FFFFFEEA3DB0C2844CD9" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFEEA3DB0C2844CD9" box="[354,431,1948,1972]" latitude="22.328056" longLatPrecision="21" longitude="103.831665" municipality="Sa Pa" name="Cat Cat" pageId="16" pageNumber="65" stateProvince="Lao Cai">Cat Cat</location>
village;
<geoCoordinate id="EE27D9C49478FFFFFE743DB0C15D4CD9" box="[508,630,1948,1972]" degrees="22" direction="north" minutes="19" orientation="latitude" pageId="16" pageNumber="65" precision="15" seconds="41" value="22.328056">22°19’41”N</geoCoordinate>
,
<geoCoordinate id="EE27D9C49478FFFFFDF73DB0C1D54CD9" box="[639,766,1948,1972]" degrees="103" direction="east" minutes="49" orientation="longitude" pageId="16" pageNumber="65" precision="15" seconds="54" value="103.831665">103°49’54”E</geoCoordinate>
; alt.
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;
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</date>
;
<collectorName id="26E6DAD59478FFFFFEFC3D9BC2FE4CA3" box="[372,469,1975,1998]" pageId="16" pageNumber="65">Tran N.T.</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="3B7BB55E9478FFFFFD983D9AC1834C85" collectingDate="2023-09-12" collectionCode="IEBR" collectorName="Mai T. V." location="Sa Pa" pageId="16" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="9D15748A9478FFFFFD983D9AC11A4CA0" box="[528,561,1974,1998]" count="1" pageId="16" pageNumber="65" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49D771E19478FFFFFDB53D9BC1A74CA3" box="[573,652,1974,1998]" country="Vietnam" name="Lao Cai" pageId="16" pageNumber="65">Lao Cai</collectingRegion>
,
<location id="8ECCE9D89478FFFFFD1D3D9AC1E04CA3" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFD1D3D9AC1E04CA3" box="[661,715,1974,1998]" name="Sa Pa" pageId="16" pageNumber="65" stateProvince="Lao Cai">Sa Pa</location>
,
<location id="8ECCE9D89478FFFFFD5C3D9AC02C4CA3" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFD5C3D9AC02C4CA3" box="[724,775,1974,1998]" name="Sa Pa" pageId="16" pageNumber="65" stateProvince="Lao Cai">Sa Pa</location>
town,
<location id="8ECCE9D89478FFFFFF4B3DFDC23A4C84" LSID="urn:lsid:plazi:treatment:03BA0E159478FFFDFF6F3C5EC13B4DA1:8ECCE9D89478FFFFFF4B3DFDC23A4C84" box="[195,273,2001,2025]" name="Cat Cat" pageId="16" pageNumber="65" stateProvince="Lao Cai">Cat Cat</location>
village;
<date id="FFAD99C39478FFFFFEE93DFDC2FE4C84" box="[353,469,2001,2025]" pageId="16" pageNumber="65" value="2023-09-12">
<collectingDate id="EFE9602B9478FFFFFEE93DFDC2FE4C84" box="[353,469,2001,2025]" pageId="16" pageNumber="65" value="2023-09-12">12.IX.2023</collectingDate>
</date>
;
<collectorName id="26E6DAD59478FFFFFE573DFDC11E4C85" box="[479,565,2001,2025]" pageId="16" pageNumber="65">Mai T.V.</collectorName>
leg.;
<collectionCode id="ED0227C69478FFFFFDE13DFDC1884C85" box="[617,675,2001,2024]" pageId="16" pageNumber="65">IEBR</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C309EC889478FFFFFCA53AFBC0F14A4E" pageId="16" pageNumber="65" type="etymology">
<paragraph id="8BACBF039478FFFFFCA53AFBC0F14A4E" blockId="16.[813,1455,214,291]" pageId="16" pageNumber="65">
<smallCapsWord id="8D4A29DF9478FFFFFCA53AFBC0814B80" baselines="232,233" box="[813,938,215,238]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="16" pageNumber="65">ETYMOLOGY</smallCapsWord>
. — The specific epithet is the Latin adjective
<taxonomicName id="4C13C4809478FFFFFA053AFAC05F4A64" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B96763119478FFFFFA053AFAC05F4A64" italics="true" pageId="16" pageNumber="65">ferruginea</emphasis>
</taxonomicName>
, which refers to the metasomal terga with conspicuously ferruginous setae.
</paragraph>
</subSubSection>
<subSubSection id="C309EC889478FFFFFCA53B17C6864931" pageId="16" pageNumber="65" type="diagnosis">
<paragraph id="8BACBF039478FFFFFCA53B17C6864931" blockId="16.[811,1455,315,604]" pageId="16" pageNumber="65">
<smallCapsWord id="8D4A29DF9478FFFFFCA53B17C0BD4A3C" baselines="332,333" box="[813,918,315,338]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="16" pageNumber="65">DIAGNOSIS</smallCapsWord>
. — The female of this new species is similar to the female of
<taxonomicName id="4C13C4809478FFFFFCC03B7AC7B54A00" authority="Niu &amp; Zhu, 2017" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[840,1182,341,365]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="xiongjiuensis">
<emphasis id="B96763119478FFFFFCC03B7AC0CA4A00" box="[840,993,342,365]" italics="true" pageId="16" pageNumber="65">M. xiongjiuensis</emphasis>
<bibRefCitation id="EF82C2F29478FFFFFC613B79C7B54A00" author="NIU Z. Q. &amp; FENG Y. &amp; ASCHER J. S. &amp; ZHU C. D." box="[1001,1182,341,365]" pageId="16" pageNumber="65" pagination="38 - 44" refId="ref12622" refString="NIU Z. Q., FENG Y., ASCHER J. S. &amp; ZHU C. D. 2017. - A new species of non-colored bands nominee [sic] bees in subgenus Maculonomia (Hymenoptera: Apoidea: Halictidae: Nomiinae: Nomia) from China. Journal of Zoological Sciences 5 (2): 38-44." type="journal article" year="2017">Niu &amp; Zhu, 2017</bibRefCitation>
</taxonomicName>
as they have a similar body form, with black integument and lacking enameled bands on the metasomal terga apically, the mesoscutum and mesoscutellum with black setae, and the forewing with an obvious dark infumation at the apical corner. Unlike
<taxonomicName id="4C13C4809478FFFFFC7A3BECC7AE4ABA" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[1010,1157,448,471]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="xiongjiuensis">
<emphasis id="B96763119478FFFFFC7A3BECC7AE4ABA" box="[1010,1157,448,471]" italics="true" pageId="16" pageNumber="65">M. xiongjiuensis</emphasis>
</taxonomicName>
, metasomal T1 has faint, sparse punctures (metasomal T1 with relatively coarser, denser punctures in
<taxonomicName id="4C13C4809478FFFFFCA33BD9C0EB4961" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[811,960,501,524]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="xiongjiuensis">
<emphasis id="B96763119478FFFFFCA33BD9C0EB4961" box="[811,960,501,524]" italics="true" pageId="16" pageNumber="65">M. xiongjiuensis</emphasis>
</taxonomicName>
); antennal F4-F12 reddish brown (antennal F4-F12 black in
<taxonomicName id="4C13C4809478FFFFFCF43823C73F494B" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[892,1044,527,550]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="xiongjiuensis">
<emphasis id="B96763119478FFFFFCF43823C73F494B" box="[892,1044,527,550]" italics="true" pageId="16" pageNumber="65">M. xiongjiuensis</emphasis>
</taxonomicName>
); discs of metasomal T1-T
<quantity id="4CEB12E69478FFFFFA9E3822C617494A" box="[1302,1340,526,551]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="16" pageNumber="65" unit="in" value="5.0">5 in</quantity>
lateral view with obvious ferruginous setae (discs of metasomal T1-T2 disc with yellowish-brown setae, T3-T5 with black setae in
<taxonomicName id="4C13C4809478FFFFFA8F3868C6B44936" authorityName="Niu &amp; Zhu" authorityYear="2017" box="[1287,1439,580,603]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="xiongjiuensis">
<emphasis id="B96763119478FFFFFA8F3868C6B44936" box="[1287,1439,580,603]" italics="true" pageId="16" pageNumber="65">M. xiongjiuensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C309EC889478FFFDFCA538BAC3D24E80" lastPageId="18" lastPageNumber="67" pageId="16" pageNumber="65" type="description">
<paragraph id="8BACBF039478FFFFFCA538BAC09649C3" blockId="16.[810,1456,662,816]" box="[813,957,662,688]" pageId="16" pageNumber="65">
<smallCapsWord id="8D4A29DF9478FFFFFCA538BAC09649C3" baselines="681,681" box="[813,957,662,688]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="16" pageNumber="65">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BACBF039478FFFFFCA53899C05D49A2" blockId="16.[810,1456,662,816]" box="[813,886,693,719]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFCA53899C05D49A2" box="[813,886,693,719]" italics="true" pageId="16" pageNumber="65">Female</emphasis>
</paragraph>
<paragraph id="8BACBF039478FFFFFCA538FAC09D4842" blockId="16.[810,1456,662,816]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFCA538FAC0E4499D" bold="true" box="[813,975,726,752]" pageId="16" pageNumber="65">Measurements.</emphasis>
Total body length
<quantity id="4CEB12E69478FFFFFB0438F9C624499D" box="[1164,1295,725,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="16" pageNumber="65" unit="mm" value="12.25" valueMax="12.5" valueMin="12.0">12-12.5 mm</quantity>
(n=4;
<typeStatus id="54A801A19478FFFFFADC38F9C69B4982" box="[1364,1456,725,751]" pageId="16" pageNumber="65" type="holotype">holotype</typeStatus>
=
<quantity id="4CEB12E69478FFFFFCC838D9C0824862" box="[832,937,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="16" pageNumber="65" unit="mm" value="12.5">12.5 mm</quantity>
), forewing length
<quantity id="4CEB12E69478FFFFFBE738D9C62A487D" box="[1135,1281,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.15" metricValueMin="1.1" pageId="16" pageNumber="65" unit="mm" value="11.25" valueMax="11.5" valueMin="11.0">11- 11.5 mm</quantity>
(n =4;
<typeStatus id="54A801A19478FFFFFAD838D9C6844862" box="[1360,1455,757,783]" pageId="16" pageNumber="65" type="holotype">holotype</typeStatus>
=
<quantity id="4CEB12E69478FFFFFCCA3939C0814842" box="[834,938,789,816]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="65" unit="mm" value="11.5">11.5 mm</quantity>
).
</paragraph>
<paragraph id="8BACBF039478FFFFFCA53979C6864F03" blockId="16.[810,1457,853,1134]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFCA53979C0BD4802" bold="true" box="[813,918,853,879]" pageId="16" pageNumber="65">Structure.</emphasis>
Head broader than long, about 1.3× as broad as long (
<figureCitation id="1328A3869478FFFFFCBA3959C0BB48E2" box="[818,912,885,911]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12A</figureCitation>
); compound eyes about 1.4× genal width; mandible with short preapical tooth; supraclypeal area medially convex; clypeus convex laterally, approximately 1.5× as broad as long and apical margin straight (
<figureCitation id="1328A3869478FFFFFBD839F8C79B4882" box="[1104,1200,980,1007]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12A</figureCitation>
). Mesosoma about 1.2× as long as broad; mesoscutellum round apically (
<figureCitation id="1328A3869478FFFFFAC939D8C68F4F62" box="[1345,1444,1012,1039]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12B</figureCitation>
). Forewing with first and third submarginal cells broader than second submarginal cell, 1m-cu entering anterior apex of second marginal cell.Metabasitibial plate triangular, glabrous (
<figureCitation id="1328A3869478FFFFFACF3E78C68A4F03" box="[1351,1441,1108,1134]" captionStart="FIG" captionStartId="17.[156,167,1721,1739]" captionTargetBox="[258,1342,215,1676]" captionTargetId="figure-130@17.[246,1342,215,1677]" captionTargetPageId="17" captionText="FIG. 11. — Maculonomia (Maculonomia) ferruginea Engel, Tran, &amp; Nguyen, n. sp., holotype, female: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831238" httpUri="https://zenodo.org/record/14831238/files/figure.png" pageId="16" pageNumber="65">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="8BACBF039478FFFFFCA53EB8C61C4C41" blockId="16.[811,1457,1172,1836]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFCA53EB8C0914FC3" bold="true" box="[813,954,1172,1198]" pageId="16" pageNumber="65">Sculpturing.</emphasis>
Clypeus with shiny, coarse, wrinkled, dense (not contiguous) punctures; supraclypeal area with dense, contiguous punctures except punctures sparser medioapically; scape with small, sparse punctures; paraocular area with dense, coarse, wrinkled punctures of unequal sizes (
<figureCitation id="1328A3869478FFFFFCBF3F1FC0B24E20" box="[823,921,1331,1357]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12A</figureCitation>
); frons with dense (not contiguous), round punctures; vertex with dense, wrinkled punctures except outer margin of median ocellus with shiny, impunctate, smooth area. Mesoscutum with dense, small, round punctures of unequal sizes; mesoscutellum as on mesoscutum except punctures laterally and slightly so paramedially (
<figureCitation id="1328A3869478FFFFFAC93FFFC68F4E80" box="[1345,1444,1491,1517]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12B</figureCitation>
). Anterior-facing surface of metasomal T1 with small, faint, relatively dense punctures except sparser medio-apically; disc of T1 with small, faint, sparse punctures; apical marginal zone impunctate, faintly imbricate to smooth, with a few sparse punctures scattered at base a slightly more in horizontal rows laterally (
<figureCitation id="1328A3869478FFFFFBB33CBEC78B4DC1" box="[1083,1184,1682,1708]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12D</figureCitation>
). Metasomal T2-T4 with faint, relatively sparse punctures except basally with small, round, denser punctures; marginal zones of metasomal T2-T4 with small, round punctures on basal two-thirds, remainder smooth and impunctate (
<figureCitation id="1328A3869478FFFFFB493D3EC6014C41" box="[1217,1322,1810,1836]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="8BACBF039478FFFFFCA53D7DC61C4C86" blockId="16.[813,1455,1873,2028]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFCA53D7DC05F4C06" bold="true" box="[813,884,1873,1899]" pageId="16" pageNumber="65">Color.</emphasis>
Body black except mandible ferruginous medially (
<figureCitation id="1328A3869478FFFFFCBE3D5DC0BC4CE6" box="[822,919,1905,1931]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12A</figureCitation>
). F4-F12 reddish brown ventrally (
<figureCitation id="1328A3869478FFFFFA8F3D5DC64C4CE6" box="[1287,1383,1905,1931]" captionStart="FIG" captionStartId="18.[132,143,1341,1359]" captionTargetBox="[229,1358,215,1298]" captionTargetId="figure-347@18.[228,1358,215,1298]" captionTargetPageId="18" captionText="FIG. 12. — Maculonomia (Maculonomia) ferruginea Engel,Tran,&amp; Nguyen, n. sp., holotype, female: A, facial view;B, mesosoma in dorsal view;C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="16" pageNumber="65">Fig. 12A</figureCitation>
). Forewing yellow to yellowish brown, often a bit lighter apically beyond the veins, obviously dark infumate at apical corner (
<figureCitation id="1328A3869478FFFFFCBC3DFDC0A84C86" box="[820,899,2001,2027]" captionStart="FIG" captionStartId="17.[156,167,1721,1739]" captionTargetBox="[258,1342,215,1676]" captionTargetId="figure-130@17.[246,1342,215,1677]" captionTargetPageId="17" captionText="FIG. 11. — Maculonomia (Maculonomia) ferruginea Engel, Tran, &amp; Nguyen, n. sp., holotype, female: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831238" httpUri="https://zenodo.org/record/14831238/files/figure.png" pageId="16" pageNumber="65">Fig. 11</figureCitation>
). Metasomal S2-S5 ferruginous apically.
</paragraph>
<caption id="DF6CEF8B9479FFFEFF143C95C6B24DA6" ID-DOI="http://doi.org/10.5281/zenodo.14831238" ID-Zenodo-Dep="14831238" box="[156,1433,1721,1739]" httpUri="https://zenodo.org/record/14831238/files/figure.png" pageId="17" pageNumber="66" startId="17.[156,167,1721,1739]" targetBox="[258,1342,215,1676]" targetPageId="17" targetType="figure">
<paragraph id="8BACBF039479FFFEFF143C95C6B24DA6" blockId="17.[156,1433,1721,1739]" box="[156,1433,1721,1739]" pageId="17" pageNumber="66">
<smallCapsWord id="8D4A29DF9479FFFEFF143C95C39F4DA7" baselines="1734,1735" box="[156,180,1721,1739]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="17" pageNumber="66">FIG</smallCapsWord>
. 11. —
<taxonomicName id="4C13C4809479FFFEFF7E3C95C15F4DA6" authority="Engel" authorityName="Engel" box="[246,628,1721,1739]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="66" phylum="Arthropoda" rank="species" species="ferruginea" subGenus="Maculonomia">
<emphasis id="B96763119479FFFEFF7E3C95C24D4DA6" box="[246,358,1721,1739]" italics="true" pageId="17" pageNumber="66">Maculonomia</emphasis>
(
<emphasis id="B96763119479FFFEFEF83C95C2CB4DA6" box="[368,480,1721,1739]" italics="true" pageId="17" pageNumber="66">Maculonomia</emphasis>
)
<emphasis id="B96763119479FFFEFE623C95C1164DA6" box="[490,573,1721,1739]" italics="true" pageId="17" pageNumber="66">ferruginea</emphasis>
Engel
</taxonomicName>
, Tran, &amp; Nguyen,
<taxonomicNameLabel id="A254DE6A9479FFFEFC8D3C95C01C4DA6" box="[773,823,1721,1739]" pageId="17" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
, holotype, female:
<emphasis id="B96763119479FFFEFC5B3C95C0CB4DA6" bold="true" box="[979,992,1721,1739]" pageId="17" pageNumber="66">A</emphasis>
, lateral habitus;
<emphasis id="B96763119479FFFEFBE13C95C75D4DA6" bold="true" box="[1129,1142,1721,1739]" pageId="17" pageNumber="66">B</emphasis>
, dorsal habitus. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BACBF039479FFFEFF0C3D3EC6214C86" blockId="17.[130,777,1810,2028]" lastBlockId="17.[812,1457,1810,2028]" pageId="17" pageNumber="66">
<emphasis id="B96763119479FFFEFF0C3D3EC2224C41" bold="true" box="[132,265,1810,1836]" pageId="17" pageNumber="66">Pubescence.</emphasis>
Face with short, sparse, blackish setae. Setae of gena softer and denser than those on face. Pronotum basally and outer margin of pronotal lobe with dense, yellowish setae. Mesosoma dorsally and laterally with dense, blackish setae, mesoscutellum covered with dense, blackish setae; mesosoma ventrally with ferruginous setae. Propodeum laterally with tuft of long, ferruginous setae. Metafemur with ferruginous scopa. Outer surface of metatibia with dense, ferruginous, plumose setae. Outer surface of metabasitarsus with bristle-like, ferruginous setae. Inner surfaces of metatibia and metabasitarsus with short, bristle-like, bright-ferruginous setae. Anterior-facing surface of metasomal T1 and discs of T1-T
<quantity id="4CEB12E69479FFFEFA293DBDC0694CA6" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="17" pageNumber="66" unit="in" value="5.0">5 in</quantity>
lateral view with obvious ferruginous setae. Metasomal S2-S5 with bright ferruginous setae apically.
</paragraph>
<caption id="DF6CEF8B947AFFFDFF0C3F11C07C4E08" ID-DOI="http://doi.org/10.5281/zenodo.14831240" ID-Zenodo-Dep="14831240" httpUri="https://zenodo.org/record/14831240/files/figure.png" pageId="18" pageNumber="67" startId="18.[132,143,1341,1359]" targetBox="[229,1358,215,1298]" targetPageId="18" targetType="figure">
<paragraph id="8BACBF03947AFFFDFF0C3F11C07C4E08" blockId="18.[132,1455,1340,1381]" pageId="18" pageNumber="67">
<smallCapsWord id="8D4A29DF947AFFFDFF0C3F11C3B64E20" baselines="1354,1354" box="[132,157,1341,1359]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="18" pageNumber="67">FIG</smallCapsWord>
. 12. —
<taxonomicName id="4C13C480947AFFFDFF553F11C1734E22" authority="Engel" authorityName="Engel" box="[221,600,1341,1359]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="67" phylum="Arthropoda" rank="species" species="ferruginea" subGenus="Maculonomia">
<emphasis id="B9676311947AFFFDFF553F11C2664E22" box="[221,333,1341,1359]" italics="true" pageId="18" pageNumber="67">Maculonomia</emphasis>
(
<emphasis id="B9676311947AFFFDFEDE3F11C2ED4E22" box="[342,454,1341,1359]" italics="true" pageId="18" pageNumber="67">Maculonomia</emphasis>
)
<emphasis id="B9676311947AFFFDFE473F11C1094E22" box="[463,546,1341,1359]" italics="true" pageId="18" pageNumber="67">ferruginea</emphasis>
Engel
</taxonomicName>
,Tran,&amp; Nguyen,
<taxonomicNameLabel id="A254DE6A947AFFFDFD6D3F11C03D4E22" box="[741,790,1341,1359]" pageId="18" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
, holotype, female:
<emphasis id="B9676311947AFFFDFC263F11C0904E22" bold="true" box="[942,955,1341,1359]" pageId="18" pageNumber="67">A</emphasis>
, facial view;
<emphasis id="B9676311947AFFFDFBAB3F11C71B4E22" bold="true" box="[1059,1072,1341,1359]" pageId="18" pageNumber="67">B</emphasis>
, mesosoma in dorsal view;
<emphasis id="B9676311947AFFFDFA9B3F10C60A4E23" bold="true" box="[1299,1313,1340,1358]" pageId="18" pageNumber="67">C</emphasis>
, metatibia showing the basal plate;
<emphasis id="B9676311947AFFFDFEAF3F7FC21E4E08" bold="true" box="[295,309,1363,1381]" pageId="18" pageNumber="67">D</emphasis>
, metasoma in dorsal view. Scale bars: A, B, D, 1 mm; C, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BACBF03947AFFFDFF0C3F9FC3924EA0" blockId="18.[132,249,1459,1517]" box="[132,185,1459,1485]" pageId="18" pageNumber="67">
<emphasis id="B9676311947AFFFDFF0C3F9FC3924EA0" box="[132,185,1459,1485]" italics="true" pageId="18" pageNumber="67">Male</emphasis>
</paragraph>
<paragraph id="8BACBF03947AFFFDFF0C3FFFC3D24E80" blockId="18.[132,249,1459,1517]" box="[132,249,1491,1517]" pageId="18" pageNumber="67">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309EC88947AFFFDFF0C3C3FC13B4DA1" pageId="18" pageNumber="67" type="discussion">
<paragraph id="8BACBF03947AFFFDFF0C3C3FC3C34D46" blockId="18.[130,775,1555,1741]" box="[132,232,1555,1581]" pageId="18" pageNumber="67">
<smallCapsWord id="8D4A29DF947AFFFDFF0C3C3FC3C34D46" baselines="1574,1574" box="[132,232,1555,1581]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="18" pageNumber="67">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BACBF03947AFFFDFF0C3C1FC13B4DA1" blockId="18.[130,775,1555,1741]" pageId="18" pageNumber="67">
At first glance the new species can easily be confused if placed among specimens of
<taxonomicName id="4C13C480947AFFFDFED43C7FC2CF4D01" authorityName="Smith" authorityYear="1875" box="[348,484,1619,1644]" class="Insecta" family="Halictidae" genus="Nomia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="67" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="B9676311947AFFFDFED43C7FC2CF4D01" box="[348,484,1619,1644]" italics="true" pageId="18" pageNumber="67">M. terminata</emphasis>
</taxonomicName>
and indeed we encountered them while entering data for
<taxonomicName id="4C13C480947AFFFDFE3A3C5FC1164DE1" authorityName="Smith" authorityYear="1875" box="[434,573,1651,1676]" class="Insecta" family="Halictidae" genus="Nomia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="67" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="B9676311947AFFFDFE3A3C5FC1164DE1" box="[434,573,1651,1676]" italics="true" pageId="18" pageNumber="67">M. terminata</emphasis>
</taxonomicName>
. However, the species, although seemingly cryptic, has sufficient differentiating factors to validate it as a new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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