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<mods:title id="C0FDE8D137FDF4DC48952B33CF5EEE8E">Two new species of the South African endemic bee genus Rediviva Friese (Hymenoptera: Apoidea: Melittidae)</mods:title>
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<paragraph id="8BD636CC6221FFEFFF0C46D1FDA0FB1B" blockId="1.[151,517,1155,1182]" box="[151,517,1155,1182]" pageId="1" pageNumber="72">
<heading id="D09E81A06221FFEFFF0C46D1FDA0FB1B" bold="true" box="[151,517,1155,1182]" fontSize="11" level="1" pageId="1" pageNumber="72" reason="1">
<taxonomicName id="4C694D4F6221FFEFFF0C46D1FDA0FB1B" ID-CoL="DQWJ" authority="Friese 1909" authorityName="Friese" authorityYear="1909" box="[151,517,1155,1182]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B91DEADE6221FFEFFF0C46D1FDA0FB1B" bold="true" box="[151,517,1155,1182]" pageId="1" pageNumber="72">
<emphasis id="B91DEADE6221FFEFFF0C46D1FECBFB18" bold="true" box="[151,366,1156,1182]" italics="true" pageId="1" pageNumber="72">Andrena africana</emphasis>
<bibRefCitation id="EFF84B3D6221FFEFFEED46D6FDA0FB1B" author="Friese" box="[374,517,1155,1181]" pageId="1" pageNumber="72" refString="Friese, H. (1909) 8. Hymenoptera. 5. Apidae. In: Sjoestedts, B. Y. (Ed.) Kilimandjaro-Meru Expedition. Stockholm, 8, pp. 119 - 168." type="journal article" year="1909">Friese 1909</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C37365476221FFEFFF0C469FFE5CF80B" pageId="1" pageNumber="72" type="reference_group">
<paragraph id="8BD636CC6221FFEFFF0C469FFDA2FB66" blockId="1.[151,749,1226,1432]" box="[151,519,1226,1248]" pageId="1" pageNumber="72">
<treatmentCitationGroup id="AB7911E26221FFEFFF0C469FFDA2FB66" box="[151,519,1226,1248]" pageId="1" pageNumber="72">
<taxonomicName id="4C694D4F6221FFEFFF0C469FFDA6FB66" ID-CoL="DQWJ" authority="Friese, 1909: 126" authorityName="Friese" authorityPageNumber="126" authorityYear="1909" box="[151,515,1226,1248]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B91DEADE6221FFEFFF0C469FFEECFB66" box="[151,329,1226,1248]" italics="true" pageId="1" pageNumber="72">Andrena africana</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFED4469FFDA6FB66" author="Friese" box="[335,515,1226,1248]" page="126" pageId="1" pageNumber="72" year="1909">
<bibRefCitation id="EFF84B3D6221FFEFFED4469FFE75FB66" author="Friese" box="[335,464,1226,1248]" pageId="1" pageNumber="72" refString="Friese, H. (1909) 8. Hymenoptera. 5. Apidae. In: Sjoestedts, B. Y. (Ed.) Kilimandjaro-Meru Expedition. Stockholm, 8, pp. 119 - 168." type="journal article" year="1909">Friese, 1909</bibRefCitation>
: 126
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</taxonomicName>
.
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</paragraph>
<paragraph id="8BD636CC6221FFEFFF0C46BCFD3AFA9B" blockId="1.[151,749,1226,1432]" pageId="1" pageNumber="72">
<treatmentCitationGroup id="AB7911E26221FFEFFF0C46BCFD3AFA9B" pageId="1" pageNumber="72">
<taxonomicName id="4C694D4F6221FFEFFF0C46BCFDBFFB79" ID-CoL="DRZT" authority="Friese, 1915: 265" authorityName="Friese" authorityPageNumber="265" authorityYear="1915" box="[151,538,1257,1279]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="B91DEADE6221FFEFFF0C46BCFEC4FB79" box="[151,353,1257,1279]" italics="true" pageId="1" pageNumber="72">Andrena kristenseni</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFEFC46BCFDBFFB79" author="Friese" box="[359,538,1257,1279]" page="265" pageId="1" pageNumber="72" year="1915">
<bibRefCitation id="EFF84B3D6221FFEFFEFC46BCFE4DFB79" author="Friese" box="[359,488,1257,1279]" pageId="1" pageNumber="72" refString="Friese, H. (1915) Zur Bienenfauna von Abessinien. (Hym.). Deutsche Entomologische Zeitschrift, 1915, 265 - 298." type="journal article" year="1915">Friese, 1915</bibRefCitation>
: 265
</treatmentCitation>
</taxonomicName>
, 268, 282293.
<taxonomicName id="4C694D4F6221FFEFFF0C475DFDA7FA98" ID-CoL="DS8W" authority="Friese, 1915: 265" authorityName="Friese" authorityPageNumber="265" authorityYear="1915" box="[151,514,1288,1310]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="meneliki">
<emphasis id="B91DEADE6221FFEFFF0C475DFEECFA98" box="[151,329,1288,1310]" italics="true" pageId="1" pageNumber="72">Andrena meneliki</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFED4475DFDA7FA98" author="Friese" box="[335,514,1288,1310]" page="265" pageId="1" pageNumber="72" year="1915">
<bibRefCitation id="EFF84B3D6221FFEFFED4475DFE75FA98" author="Friese" box="[335,464,1288,1310]" pageId="1" pageNumber="72" refString="Friese, H. (1915) Zur Bienenfauna von Abessinien. (Hym.). Deutsche Entomologische Zeitschrift, 1915, 265 - 298." type="journal article" year="1915">Friese, 1915</bibRefCitation>
: 265
</treatmentCitation>
</taxonomicName>
, 269, 282283.
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</paragraph>
<paragraph id="8BD636CC6221FFEFFF0C4772FD48FA1E" blockId="1.[151,749,1226,1432]" pageId="1" pageNumber="72">
<treatmentCitationGroup id="AB7911E26221FFEFFF0C4772FD48FA1E" pageId="1" pageNumber="72">
<emphasis id="B91DEADE6221FFEFFF0C4772FE56FABB" box="[151,499,1319,1341]" italics="true" pageId="1" pageNumber="72">
<taxonomicName id="4C694D4F6221FFEFFF0C4772FE0DFABB" box="[151,424,1319,1341]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="subSpecies" species="abessinica" subSpecies="nomen">Andrena abessinica nomen</taxonomicName>
nudum
</emphasis>
:
<treatmentCitation id="0AC810DD6221FFEFFD9A4773FD0BFABA" author="Friese" box="[513,686,1318,1340]" page="281" pageId="1" pageNumber="72" year="1915">
<bibRefCitation id="EFF84B3D6221FFEFFD9A4773FDDEFABA" author="Friese" box="[513,635,1318,1340]" pageId="1" pageNumber="72" refString="Friese, H. (1915) Zur Bienenfauna von Abessinien. (Hym.). Deutsche Entomologische Zeitschrift, 1915, 265 - 298." type="journal article" year="1915">Friese 1915</bibRefCitation>
: 281
</treatmentCitation>
.
<taxonomicName id="4C694D4F6221FFEFFF0C4710FD04FADD" ID-CoL="DR73" authority="Viereck, 1917: 236" authorityName="Viereck" authorityPageNumber="236" authorityYear="1917" box="[151,673,1349,1371]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="boswendica" subGenus="Andrena">
<emphasis id="B91DEADE6221FFEFFF0C4710FF48FADD" box="[151,237,1349,1371]" italics="true" pageId="1" pageNumber="72">Andrena</emphasis>
(
<emphasis id="B91DEADE6221FFEFFF674710FEF7FADD" box="[252,338,1349,1371]" italics="true" pageId="1" pageNumber="72">Andrena</emphasis>
)
<emphasis id="B91DEADE6221FFEFFEFB4710FE7DFADD" box="[352,472,1349,1371]" italics="true" pageId="1" pageNumber="72">boswendica</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFE454710FD04FADD" author="Viereck" box="[478,673,1349,1371]" page="236" pageId="1" pageNumber="72" year="1917">
<bibRefCitation id="EFF84B3D6221FFEFFE454710FDD5FADD" author="Viereck" box="[478,624,1349,1371]" pageId="1" pageNumber="72" refString="Viereck, H. L. (1917) New species of bees of the genus Andrena, from Equatorial Africa, in the American Museum of Natural History. Bulletin of the American Museum of Natural History, 37, 235 - 239." type="journal article" year="1917">Viereck, 1917</bibRefCitation>
: 236
</treatmentCitation>
</taxonomicName>
239.
<taxonomicName id="4C694D4F6221FFEFFF0C4731FD15FAFF" ID-CoL="DQW9" authority="Viereck, 1917: 235" authorityName="Viereck" authorityPageNumber="235" authorityYear="1917" box="[151,688,1379,1402]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="aequatorialis" subGenus="Andrena">
<emphasis id="B91DEADE6221FFEFFF0C4731FF48FAFC" box="[151,237,1380,1402]" italics="true" pageId="1" pageNumber="72">Andrena</emphasis>
(
<emphasis id="B91DEADE6221FFEFFF674731FEF7FAFC" box="[252,338,1380,1402]" italics="true" pageId="1" pageNumber="72">Andrena</emphasis>
)
<emphasis id="B91DEADE6221FFEFFEFB4731FE42FAFC" box="[352,487,1380,1402]" italics="true" pageId="1" pageNumber="72">aequatorialis</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFE754736FD15FAFF" author="Viereck" box="[494,688,1379,1401]" page="235" pageId="1" pageNumber="72" year="1917">
<bibRefCitation id="EFF84B3D6221FFEFFE754736FDDAFAFF" author="Viereck" box="[494,639,1379,1401]" pageId="1" pageNumber="72" refString="Viereck, H. L. (1917) New species of bees of the genus Andrena, from Equatorial Africa, in the American Museum of Natural History. Bulletin of the American Museum of Natural History, 37, 235 - 239." type="journal article" year="1917">Viereck, 1917</bibRefCitation>
: 235
</treatmentCitation>
</taxonomicName>
236.
<taxonomicName id="4C694D4F6221FFEFFF0C47D7FDF4FA1E" authority="Cockerell, 1934: 450" authorityName="Cockerell" authorityPageNumber="450" authorityYear="1934" box="[151,593,1410,1432]" class="Insecta" family="Melittidae" genus="Notomelitta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B91DEADE6221FFEFFF0C47D7FED0FA1E" box="[151,373,1410,1432]" italics="true" pageId="1" pageNumber="72">Notomelitta tropicalis</emphasis>
<treatmentCitation id="0AC810DD6221FFEFFEE047D7FDF4FA1E" author="Cockerell" box="[379,593,1410,1432]" page="450" pageId="1" pageNumber="72" year="1934">
<bibRefCitation id="EFF84B3D6221FFEFFEE047D7FD85FA1E" author="Cockerell" box="[379,544,1410,1432]" pageId="1" pageNumber="72" refString="Cockerell, T. D. A. (1934) Descriptions and records of bees. - CXLVIII. Annals and Magazine of Natural History, (ser. 10) 13, 444 - 456." type="journal article" year="1934">Cockerell, 1934</bibRefCitation>
: 450
</treatmentCitation>
</taxonomicName>
452.
<emphasis id="B91DEADE6221FFEFFD1447D6FD48FA1E" bold="true" box="[655,749,1411,1432]" pageId="1" pageNumber="72">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD636CC6221FFEFFF0C4790FE4CF917" blockId="1.[151,1436,1477,1934]" pageId="1" pageNumber="72">
<taxonomicName id="4C694D4F6221FFEFFF0C4790FE36FA5B" box="[151,403,1477,1501]" class="Insecta" family="Melittidae" genus="Notomelitta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B91DEADE6221FFEFFF0C4790FE36FA5B" box="[151,403,1477,1501]" italics="true" pageId="1" pageNumber="72">Notomelitta tropicalis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFF84B3D6221FFEFFDE64790FCE7FA5B" author="Cockerell" box="[637,834,1477,1501]" pageId="1" pageNumber="72" refString="Cockerell, T. D. A. (1934) Descriptions and records of bees. - CXLVIII. Annals and Magazine of Natural History, (ser. 10) 13, 444 - 456." type="journal article" year="1934">Cockerell (1934)</bibRefCitation>
from
<collectingCountry id="F37E765C6221FFEFFC084790FAC0FA5B" box="[915,1381,1477,1501]" name="Democratic Republic of the Congo" pageId="1" pageNumber="72">the Democratic Republic of the Congo</collectingCountry>
and
<bibRefCitation id="EFF84B3D6221FFEFFF0C47BCFEF5F987" author="Michener" box="[151,336,1513,1537]" pageId="1" pageNumber="72" refString="Michener, C. D. (1981) Classification of the bee family Melittidae with a review of species of Meganomiinae. Contributions of the American Entomological Institute, 18, 1 - 135." type="journal article" year="1981">Michener (1981</bibRefCitation>
: 47), solely based on the description, transferred it to
<taxonomicName id="4C694D4F6221FFEFFC2347BCFBBFF987" ID-CoL="77GC" box="[952,1050,1513,1537]" class="Insecta" family="Melittidae" genus="Rediviva" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEADE6221FFEFFC2347BCFBBFF987" box="[952,1050,1513,1537]" italics="true" pageId="1" pageNumber="72">Rediviva</emphasis>
</taxonomicName>
without examining the
<typeStatus id="54D2886E6221FFEFFAB147BFFAFBF984" box="[1322,1374,1514,1538]" pageId="1" pageNumber="72">type</typeStatus>
. This record of a
<taxonomicName id="4C694D4F6221FFEFFE874458FEDBF9A3" ID-CoL="77GC" box="[284,382,1549,1573]" class="Insecta" family="Melittidae" genus="Rediviva" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEADE6221FFEFFE874458FEDBF9A3" box="[284,382,1549,1573]" italics="true" pageId="1" pageNumber="72">Rediviva</emphasis>
</taxonomicName>
species is far outside the known range of the genus, that is assumed to be endemic to
<collectingCountry id="F37E765C6221FFEFFAC04458FF7AF9CF" name="South Africa" pageId="1" pageNumber="72">South Africa</collectingCountry>
and
<collectingCountry id="F37E765C6221FFEFFE824464FED6F9CF" box="[281,371,1585,1609]" name="Lesotho" pageId="1" pageNumber="72">Lesotho</collectingCountry>
(
<bibRefCitation id="EFF84B3D6221FFEFFE1F4464FD92F9CF" author="Michener" box="[388,567,1585,1609]" pageId="1" pageNumber="72" refString="Michener, C. D. (2007) The Bees of the World. Johns Hopkins University Press, Baltimore, Maryland, 992 pp." type="book" year="2007">Michener 2007</bibRefCitation>
). Thus, it is surprising that
<taxonomicName id="4C694D4F6221FFEFFCEF4467FC5AF9CF" box="[884,1023,1585,1609]" class="Insecta" family="Melittidae" genus="Notomelitta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B91DEADE6221FFEFFCEF4467FC5AF9CF" box="[884,1023,1585,1609]" italics="true" pageId="1" pageNumber="72">R. tropicalis</emphasis>
</taxonomicName>
was not mentioned when the genus was first revised (
<bibRefCitation id="EFF84B3D6221FFEFFEFD4400FD3BF9EB" author="Whitehead" box="[358,670,1621,1645]" pageId="1" pageNumber="72" refString="Whitehead, V. B. &amp; Steiner, K. E. (2001) Oil-collecting bees of the winter rainfall area of South Africa (Melittidae, Rediviva). Annals of the South African Museum, 108, 143 - 277." type="journal article" year="2001">Whitehead &amp; Steiner 2001</bibRefCitation>
,
<bibRefCitation id="EFF84B3D6221FFEFFD374400FC0BF9EB" author="Whitehead" box="[684,942,1621,1645]" pageId="1" pageNumber="72" refString="Whitehead, V. B., Steiner, K. E. &amp; Eardley, C. D. (2008) Oil-collecting bees mostly of the summer rainfall area of southern Africa (Hymenoptera: Melittidae: Rediviva). Journal of the Kansas Entomological Society, 81, 122 - 141." type="journal article" year="2008">
Whitehead
<emphasis id="B91DEADE6221FFEFFCB44403FCCFF9EB" box="[815,874,1621,1645]" italics="true" pageId="1" pageNumber="72">et al.</emphasis>
2008
</bibRefCitation>
) and it is listed under this combination by
<bibRefCitation id="EFF84B3D6221FFEFFF0C442CFE40F917" author="Eardley" box="[151,485,1657,1681]" pageId="1" pageNumber="72" refString="Eardley, C. D. &amp; Urban, R. (2010) Catalogue of Afrotropical bees (Hymenoptera: Apoidea: Apiformes). Zootaxa, 2455, 1 - 548." type="journal article" year="2010">Eardley &amp; Urban (2010: 197)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD636CC6221FFEFFF5C44C8FE5CF80B" blockId="1.[151,1436,1477,1934]" pageId="1" pageNumber="72">
The occurrence of an oil-collecting
<taxonomicName id="4C694D4F6221FFEFFDCF44C8FD13F933" ID-CoL="77GC" box="[596,694,1693,1717]" class="Insecta" family="Melittidae" genus="Rediviva" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEADE6221FFEFFDCF44C8FD13F933" box="[596,694,1693,1717]" italics="true" pageId="1" pageNumber="72">Rediviva</emphasis>
</taxonomicName>
species outside the range of its presently known host plants would potentially have far reaching consequences for the understanding of the origin and evolution of the genus. To clarify this situation the
<typeStatus id="54D2886E6221FFEFFE2744B3FE49F978" box="[444,492,1766,1790]" pageId="1" pageNumber="72">type</typeStatus>
of
<taxonomicName id="4C694D4F6221FFEFFD8744B0FCB3F97B" box="[540,790,1765,1789]" class="Insecta" family="Melittidae" genus="Notomelitta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B91DEADE6221FFEFFD8744B0FCB3F97B" box="[540,790,1765,1789]" italics="true" pageId="1" pageNumber="72">Notomelitta tropicalis</emphasis>
</taxonomicName>
was examined, that is housed in the Musee Royal de l´Afrique Centrale in Tervuren (
<collectingCountry id="F37E765C6221FFEFFE64455CFDC3F8A7" box="[511,614,1801,1825]" name="Belgium" pageId="1" pageNumber="72">Belgium</collectingCountry>
). The specimen turned out to be a male of
<taxonomicName id="4C694D4F6221FFEFFBD1455CFA39F8A7" ID-CoL="DQWJ" authority="Friese 1909" authorityName="Friese" authorityYear="1909" box="[1098,1436,1801,1825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B91DEADE6221FFEFFBD1455CFAB5F8A7" box="[1098,1296,1801,1825]" italics="true" pageId="1" pageNumber="72">Andrena africana</emphasis>
<bibRefCitation id="EFF84B3D6221FFEFFA83455CFA39F8A7" author="Friese" box="[1304,1436,1801,1825]" pageId="1" pageNumber="72" refString="Friese, H. (1909) 8. Hymenoptera. 5. Apidae. In: Sjoestedts, B. Y. (Ed.) Kilimandjaro-Meru Expedition. Stockholm, 8, pp. 119 - 168." type="journal article" year="1909">Friese 1909</bibRefCitation>
</taxonomicName>
and, thus, is a junior synonym of that species. The
<typeStatus id="54D2886E6221FFEFFD7D457BFCB3F8C0" box="[742,790,1838,1862]" pageId="1" pageNumber="72">type</typeStatus>
is labelled as follows: (orange label):
<typeStatus id="54D2886E6221FFEFFB48457BFAABF8C0" box="[1235,1294,1838,1862]" pageId="1" pageNumber="72">Type</typeStatus>
. / Musee du
<collectingCountry id="F37E765C6221FFEFFF0C4504FF41F8EF" box="[151,228,1873,1897]" name="Republic of the Congo" pageId="1" pageNumber="72">Congo</collectingCountry>
, Beni à Lesse, fin
<date id="FFD7100C6221FFEFFE524504FD8FF8EF" box="[457,554,1873,1897]" pageId="1" pageNumber="72" value="1911-07">vii-1911</date>
, Dr. Murtula / R. DÉT, D, 2499 / (handwritten by Cockerell):
<taxonomicName id="4C694D4F6221FFEFFA834504FEE7F80B" authority="Ckll." authorityName="Ckll." class="Insecta" family="Melittidae" genus="Notomelitta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B91DEADE6221FFEFFA834504FEA4F80B" italics="true" pageId="1" pageNumber="72">Notomelitta tropicalis</emphasis>
Ckll.
</taxonomicName>
,
<typeStatus id="54D2886E6221FFEFFED64523FE21F808" box="[333,388,1910,1934]" pageId="1" pageNumber="72">Type</typeStatus>
(MRAC).
</paragraph>
</subSubSection>
</treatment>
</document>