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<document id="123CCE93CAC46E80D0BB6F595686F93F" ID-CLB-Dataset="8695" ID-DOI="10.11646/zootaxa.4829.1.1" ID-GBIF-Dataset="04b19d7f-1d74-41ec-b657-9b57f47826f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402757" ID-ZooBank="7B34E058-03B4-44D0-AC4E-065B010172E1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609343828179" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2020" docId="03C00F49FF9D6E20FF57F8EB9F94F987" docLanguage="en" docName="zootaxa.4829.1.1.pdf" docOrigin="Zootaxa 4829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Winnertzia tumidoides Jaschhof &amp; Jaschhof 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFF97731FF8B6E38FFC0FFBC9D54FF8C" masterDocTitle="Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia" masterLastPageNumber="72" masterPageNumber="1" pageNumber="23" updateTime="1698849859117" updateUser="ExternalLinkService">
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<mods:title id="D475ABFD1741B6C35DA934F020000082">Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia</mods:title>
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<mods:namePart id="E0119A1936D9B9F1D58469B515B175BF">Jaschhof, Mathias</mods:namePart>
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<paragraph id="8BD6BE5FFF9D6E2EFF57F8EB9F58F8FD" blockId="22.[151,524,1878,1941]" box="[151,524,1878,1905]" pageId="22" pageNumber="23">
<heading id="D09E0933FF9D6E2EFF57F8EB9F58F8FD" bold="true" box="[151,524,1878,1905]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
<emphasis id="B91D624DFF9D6E2EFF57F8EB9F58F8FD" bold="true" box="[151,524,1878,1905]" pageId="22" pageNumber="23">
<taxonomicName id="4C69C5DCFF9D6E2EFF57F8EB9CF3F8FD" ID-CoL="8FMJV" authority="Jaschhof &amp; Jaschhof, 2020" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[151,423,1878,1905]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tumidoides" status="sp. nov.">
<emphasis id="B91D624DFF9D6E2EFF57F8EB9CF3F8FD" bold="true" box="[151,423,1878,1905]" italics="true" pageId="22" pageNumber="23">Winnertzia tumidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EDF36FF9D6E2EFE6EF8EB9F58F8FD" box="[430,524,1879,1905]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C373EDD4FF9D6E2EFF57F8C79C41F819" box="[151,277,1915,1941]" pageId="22" pageNumber="23" type="description">
<paragraph id="8BD6BE5FFF9D6E2EFF57F8C79C41F819" blockId="22.[151,524,1878,1941]" box="[151,277,1915,1941]" pageId="22" pageNumber="23">
<figureCitation id="1352A2DAFF9D6E2EFF57F8C79C41F819" box="[151,277,1915,1941]" captionStart="FIGURES 3540" captionStartId="23.[151,264,1811,1835]" captionTargetBox="[160,1427,391,1778]" captionTargetId="figure-121@23.[151,1436,381,1788]" captionTargetPageId="23" captionText="FIGURES 3540. Male morphology of Winnertzia serri (3537) and Winnertzia tumidoides (3840). 35: Fourth flagellomere, lateral, paratype. 36: Ditto, medial. 37: Genitalia, ventral, holotype. 38: Ditto. 39: Fourth flagellomere, lateral, holotype. 40: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402777" httpUri="https://zenodo.org/record/4402777/files/figure.png" pageId="22" pageNumber="23">Figs 3840</figureCitation>
</paragraph>
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<subSubSection id="C373EDD4FF9D6E20FF57F87E99D5FBF1" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BD6BE5FFF9D6E2FFF57F87E9EC2FEE9" blockId="22.[151,1437,1986,2013]" lastBlockId="23.[151,1437,151,359]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="B91D624DFF9D6E2EFF57F87E9C46F850" bold="true" box="[151,274,1986,2012]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
An inconspicuous, medium-sized, brown
<taxonomicName id="4C69C5DCFF9D6E2EFD2AF87F9E37F850" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2013" box="[746,867,1987,2012]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D624DFF9D6E2EFD2AF87F9E37F850" box="[746,867,1987,2012]" italics="true" pageId="22" pageNumber="23">Winnertzia</emphasis>
</taxonomicName>
, whose genitalia are generally similar to that of
<emphasis id="B91D624DFF9D6E2FFA40F87F9DB1FF3D" italics="true" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<taxonomicName id="4C69C5DCFF9D6E2EFA40F87F98C9F850" box="[1408,1437,1987,2012]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="25" phylum="Arthropoda" rank="species" species="tumida">W.</taxonomicName>
tumida
</emphasis>
(widespread in Northern Europe) and
<taxonomicName id="4C69C5DCFF9C6E2FFD5FFF2B9E0FFF3D" authorityName="Mamaev" authorityYear="1963" box="[671,859,151,177]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tamariciphila">
<emphasis id="B91D624DFF9C6E2FFD5FFF2B9E0FFF3D" box="[671,859,151,177]" italics="true" pageId="23" pageNumber="24">W. tamariciphila</emphasis>
</taxonomicName>
(from
<collectingCountry id="F37EFECFFF9C6E2FFC6CFF2B997BFF3E" box="[940,1071,151,178]" name="Uzbekistan" pageId="23" pageNumber="24">Uzbekistan</collectingCountry>
). A structure diagnostic of these three species is the conspicuously long, almost straight gonostylus, whose apex (i.e. the portion above the pectinate claw) is parallel-sided. A peculiarity of
<taxonomicName id="4C69C5DCFF9C6E2FFD9AFF639FADFF75" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[602,761,223,249]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tumidoides">
<emphasis id="B91D624DFF9C6E2FFD9AFF639FADFF75" box="[602,761,223,249]" italics="true" pageId="23" pageNumber="24">W. tumidoides</emphasis>
</taxonomicName>
is that the gonostylar claw is situated, either entirely or for the greater part, at the distal half of the gonostylar body, so that the apex is relatively short (
<figureCitation id="1352A2DAFF9C6E2FFB72FEBF9852FE91" box="[1202,1286,259,285]" captionStart="FIGURES 3540" captionStartId="23.[151,264,1811,1835]" captionTargetBox="[160,1427,391,1778]" captionTargetId="figure-121@23.[151,1436,381,1788]" captionTargetPageId="23" captionText="FIGURES 3540. Male morphology of Winnertzia serri (3537) and Winnertzia tumidoides (3840). 35: Fourth flagellomere, lateral, paratype. 36: Ditto, medial. 37: Genitalia, ventral, holotype. 38: Ditto. 39: Fourth flagellomere, lateral, holotype. 40: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402777" httpUri="https://zenodo.org/record/4402777/files/figure.png" pageId="23" pageNumber="24">Fig. 38</figureCitation>
, ↓
<superScript id="7C1C1317FF9C6E2FFAE1FEBE987DFE9C" attach="none" box="[1313,1321,258,272]" fontSize="6" pageId="23" pageNumber="24">4</superScript>
). Another distinction is that the gonocoxal emargination, which is U- rather than V-shaped, has a reinforced, sclerotized base (↓
<superScript id="7C1C1317FF9C6E2FFF6FFEF69DE3FED4" attach="right" box="[175,183,330,344]" fontSize="6" pageId="23" pageNumber="24">5</superScript>
). Females and preimaginal stages of
<taxonomicName id="4C69C5DCFF9C6E2FFD98FEF79FA2FEE9" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[600,758,331,357]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tumidoides">
<emphasis id="B91D624DFF9C6E2FFD98FEF79FA2FEE9" box="[600,758,331,357]" italics="true" pageId="23" pageNumber="24">W. tumidoides</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<caption id="DF16EED7FF9C6E2FFF57F8AF9EA3F8FF" ID-DOI="http://doi.org/10.5281/zenodo.4402777" ID-Zenodo-Dep="4402777" httpUri="https://zenodo.org/record/4402777/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,1811,1835]" targetBox="[160,1427,391,1778]" targetPageId="23">
<paragraph id="8BD6BE5FFF9C6E2FFF57F8AF9EA3F8FF" blockId="23.[151,1437,1811,1907]" pageId="23" pageNumber="24">
<emphasis id="B91D624DFF9C6E2FFF57F8AF9C00F8A7" bold="true" box="[151,340,1811,1835]" pageId="23" pageNumber="24">FIGURES 3540.</emphasis>
Male morphology of
<taxonomicName id="4C69C5DCFF9C6E2FFDF2F8A89F82F8A7" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[562,726,1812,1835]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="serri">
<emphasis id="B91D624DFF9C6E2FFDF2F8A89F82F8A7" box="[562,726,1812,1835]" italics="true" pageId="23" pageNumber="24">Winnertzia serri</emphasis>
</taxonomicName>
<emphasis id="B91D624DFF9C6E2FFD1CF8AF9E78F8A7" bold="true" box="[732,812,1811,1835]" pageId="23" pageNumber="24">(3537)</emphasis>
and
<taxonomicName id="4C69C5DCFF9C6E2FFC9EF8A89915F8A7" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[862,1089,1811,1835]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tumidoides">
<emphasis id="B91D624DFF9C6E2FFC9EF8A89915F8A7" box="[862,1089,1811,1835]" italics="true" pageId="23" pageNumber="24">Winnertzia tumidoides</emphasis>
</taxonomicName>
<emphasis id="B91D624DFF9C6E2FFB87F8AF99C3F8A7" bold="true" box="[1095,1175,1811,1835]" pageId="23" pageNumber="24">(3840)</emphasis>
.
<emphasis id="B91D624DFF9C6E2FFB64F8AF9992F8A7" bold="true" box="[1188,1222,1811,1835]" pageId="23" pageNumber="24">35:</emphasis>
Fourth flagellomere, lateral, paratype.
<emphasis id="B91D624DFF9C6E2FFE8AF88B9C3FF8C3" bold="true" box="[330,363,1847,1871]" pageId="23" pageNumber="24">36:</emphasis>
Ditto, medial.
<emphasis id="B91D624DFF9C6E2FFDC8F88B9F7EF8C3" bold="true" box="[520,554,1847,1871]" pageId="23" pageNumber="24">37:</emphasis>
Genitalia, ventral, holotype.
<emphasis id="B91D624DFF9C6E2FFC98F88B9E2EF8C3" bold="true" box="[856,890,1847,1871]" pageId="23" pageNumber="24">38:</emphasis>
Ditto.
<emphasis id="B91D624DFF9C6E2FFC04F88B9EB1F8C3" bold="true" box="[964,997,1847,1871]" pageId="23" pageNumber="24">39:</emphasis>
Fourth flagellomere, lateral, holotype.
<emphasis id="B91D624DFF9C6E2FFABAF88B98C8F8C3" bold="true" box="[1402,1436,1847,1871]" pageId="23" pageNumber="24">40:</emphasis>
Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses.
</paragraph>
</caption>
<paragraph id="8BD6BE5FFF9C6E20FF07F81E992EFE79" blockId="23.[151,1437,1953,2016]" lastBlockId="24.[151,1437,150,1547]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<taxonomicName id="4C69C5DCFF9C6E2FFF07F81E9CC2F837" box="[199,406,1953,1979]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B91D624DFF9C6E2FFF07F81E9CC2F837" box="[199,406,1953,1979]" italics="true" pageId="23" pageNumber="24">Winnertzia tumida</emphasis>
</taxonomicName>
, of which we studied all the six males known from
<collectingCountry id="F37EFECFFF9C6E2FFC1AF81D9961F837" box="[986,1077,1953,1979]" name="Sweden" pageId="23" pageNumber="24">Sweden</collectingCountry>
, differs from
<taxonomicName id="4C69C5DCFF9C6E2FFB0EF81E9838F837" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[1230,1388,1953,1979]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tumidoides">
<emphasis id="B91D624DFF9C6E2FFB0EF81E9838F837" box="[1230,1388,1953,1979]" italics="true" pageId="23" pageNumber="24">W. tumidoides</emphasis>
</taxonomicName>
in a number of characters, as follows. It is a slightly more robust and darker brown species; the fore tibia is somewhat longer (1.5 times as long as T
<subScript id="17EDBC1AFF936E20FE2FFF159CA3FF3B" attach="left" box="[495,503,169,183]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
); the empodia are longer (almost claw-long); the wings are broader (2.2 times as long as broad); the fourth flagellomere has a thicker node (1.5 times as long as broad) and a shorter neck (0.6 times as long as the node); and the number of pronotal setae is considerably larger (1517). As for genitalic characters, the larger part of the pectinate claw is situated in the basal half of the gonostylus; the dorsoposterior portions of the gonocoxites are conspicuously inflated; and the gonocoxal emargination is smaller, V- rather than U-shaped, and unsclerotized basally (
<bibRefCitation id="EFF8C3AEFF936E20FE53FEF79F97FEE9" author="Jaschhof, M. &amp; Jaschhof, C." box="[403,707,331,357]" pageId="24" pageNumber="25" pagination="1 - 392" refId="ref56222" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
: fig. 33). One of the two specimens from
<collectingCountry id="F37EFECFFF936E20FB5DFEF799A9FEE9" box="[1181,1277,331,357]" name="United Kingdom" pageId="24" pageNumber="25">England</collectingCountry>
, on which the original description of
<taxonomicName id="4C69C5DCFF936E20FE5DFED39F44FE05" box="[413,528,367,393]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B91D624DFF936E20FE5DFED39F44FE05" box="[413,528,367,393]" italics="true" pageId="24" pageNumber="25">W. tumida</emphasis>
</taxonomicName>
is based, appears to have the gonostylar claw in a slightly more distal position compared with Swedish specimens (
<bibRefCitation id="EFF8C3AEFF936E20FDF5FE2F9F8FFE21" author="Panelius, S." box="[565,731,402,429]" pageId="24" pageNumber="25" pagination="1 - 157" refId="ref56978" refString="Panelius, S. (1965) A revision of the European gall midges of the subfamily Porricondylinae (Diptera: Itonididae). Acta Zoologica Fennica, 113, 1 - 157." type="journal article" year="1965">Panelius 1965</bibRefCitation>
: fig.
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), which indicates some intraspecific variation in this character.
<taxonomicName id="4C69C5DCFF936E20FECAFE0B9F76FE5D" authorityName="Mamaev" authorityYear="1963" box="[266,546,439,465]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tamariciphila">
<emphasis id="B91D624DFF936E20FECAFE0B9F76FE5D" box="[266,546,439,465]" italics="true" pageId="24" pageNumber="25">Winnertzia tamariciphila</emphasis>
</taxonomicName>
is distinguished from both
<taxonomicName id="4C69C5DCFF936E20FC93FE0B9E97FE5D" box="[851,963,439,465]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B91D624DFF936E20FC93FE0B9E97FE5D" box="[851,963,439,465]" italics="true" pageId="24" pageNumber="25">W. tumida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69C5DCFF936E20FC38FE0B99C1FE5D" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[1016,1173,439,465]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tumidoides">
<emphasis id="B91D624DFF936E20FC38FE0B99C1FE5D" box="[1016,1173,439,465]" italics="true" pageId="24" pageNumber="25">W. tumidoides</emphasis>
</taxonomicName>
by the short, three-segmented palpus and the tarsal claws bearing three hair-like teeth (
<bibRefCitation id="EFF8C3AEFF936E20FCA8FE67995BFE79" author="Mamaev, B. M." box="[872,1039,475,501]" pageId="24" pageNumber="25" pagination="562 - 573" refId="ref56533" refString="Mamaev, B. M. (1963) Gall midges of the USSR. Part 3. New species of the genus Winnertzia Rondani, developing in the soil, in fungal mycelium and under decaying bark of coniferous trees (Itonididae, Diptera). Zoologicheskij Zhurnal, 42, 562 - 573." type="journal article" year="1963">Mamaev 1963</bibRefCitation>
: fig. 2.4).
</paragraph>
<paragraph id="8BD6BE5FFF936E20FF07FE4298C9FCB5" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF07FE429C88FD95" bold="true" box="[199,476,510,537]" pageId="24" pageNumber="25">Other male characters.</emphasis>
Body size
<quantity id="4C9113BAFF936E20FD9BFE439FE1FD95" box="[603,693,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="mm" value="1.8">1.8 mm</quantity>
.
<emphasis id="B91D624DFF936E20FD00FE439E53FD95" bold="true" box="[704,775,511,537]" pageId="24" pageNumber="25">Head.</emphasis>
Eye bridge 34 ommatidia long dorsally. Antenna slightly longer than half body. Scape and pedicel same size, both concolorous with flagellum. 11 flagellomeres, translucent sensilla on flagellomeres 110, long, filiform, mostly linear to slightly sinuous, others markedly bent or furcate. Fourth flagellomere: neck 0.8 times as long as node; node 1.8 times as long as broad; sensory hairs dense (
<figureCitation id="1352A2DAFF936E20FAAAFDD79DB6FD25" captionStart="FIGURES 3540" captionStartId="23.[151,264,1811,1835]" captionTargetBox="[160,1427,391,1778]" captionTargetId="figure-121@23.[151,1436,381,1788]" captionTargetPageId="23" captionText="FIGURES 3540. Male morphology of Winnertzia serri (3537) and Winnertzia tumidoides (3840). 35: Fourth flagellomere, lateral, paratype. 36: Ditto, medial. 37: Genitalia, ventral, holotype. 38: Ditto. 39: Fourth flagellomere, lateral, holotype. 40: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402777" httpUri="https://zenodo.org/record/4402777/files/figure.png" pageId="24" pageNumber="25">Figs 3940</figureCitation>
). Palpus slightly shorter than head height, 4 setae-bearing segments, apical segment longest of all. Labella fully developed, albeit small.
<emphasis id="B91D624DFF936E20FE26FD0F9F12FD41" bold="true" box="[486,582,691,717]" pageId="24" pageNumber="25">Thorax.</emphasis>
Pronotal setae 6. Anepimeral setae absent. Lateral mediotergal microtrichia large. Parascutellar area inconspicuous. Wing slightly shorter than body, 2.6 times as long as broad. Costal cell slightly reinforced. M
<subScript id="17EDBC1AFF936E20FE4EFCB19CC2FC97" attach="left" box="[398,406,781,795]" fontSize="6" pageId="24" pageNumber="25">4</subScript>
long, nearly straight, CuA gently bent, both veins extending to edge of wing.
<emphasis id="B91D624DFF936E20FACEFD479819FC99" bold="true" box="[1294,1357,763,789]" pageId="24" pageNumber="25">Legs.</emphasis>
Scales pointed. Basitarsal spines absent. Fore tibia 1.3 times length T
<subScript id="17EDBC1AFF936E20FC99FC8D9E35FCB3" attach="left" box="[857,865,817,831]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
. Acropods: claws slightly bent, empodia vestigial.
</paragraph>
<paragraph id="8BD6BE5FFF936E20FF57FCFE99D5FBF1" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF57FCFE9C47FCD0" bold="true" box="[151,275,834,860]" pageId="24" pageNumber="25">Abdomen.</emphasis>
Pleural membrane setose.
<emphasis id="B91D624DFF936E20FD87FCFE9FE1FCD0" bold="true" box="[583,693,834,860]" pageId="24" pageNumber="25">Genitalia</emphasis>
(
<figureCitation id="1352A2DAFF936E20FD07FCFF9E49FCD1" box="[711,797,835,861]" captionStart="FIGURES 3540" captionStartId="23.[151,264,1811,1835]" captionTargetBox="[160,1427,391,1778]" captionTargetId="figure-121@23.[151,1436,381,1788]" captionTargetPageId="23" captionText="FIGURES 3540. Male morphology of Winnertzia serri (3537) and Winnertzia tumidoides (3840). 35: Fourth flagellomere, lateral, paratype. 36: Ditto, medial. 37: Genitalia, ventral, holotype. 38: Ditto. 39: Fourth flagellomere, lateral, holotype. 40: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402777" httpUri="https://zenodo.org/record/4402777/files/figure.png" pageId="24" pageNumber="25">Fig. 38</figureCitation>
). Ninth tergite shorter than half gonopodal length; setae confined to lateroposterior portions; posterior edge straight, slightly reinforced medially; anterior edge indistinct. Gonocoxal synsclerite slightly broader than long; a short portion ventrobasally non-setose; ventral emargination large; dorsoposterior portions not markedly inflated, narrowly rounded. Gonostylus 3.5 times as long as broad; pectinate claw obliquely aligned, situated somewhat dorsomedially on a small bulge; basolateral apophysis inconspicuous, not angulated. Aedeagal apodeme broadest beyond solid basal portion, the latter long, suddenly narrowed; distal portion gently tapered towards apex. Aedeagal bulge with closely spaced rows of tiny microtrichia. Tegmen largely membranous, only basally sharply contoured, slightly tapered towards broadly rounded apex; flaps distinct, reinforced and with fine microtrichia along edges; parameral apodemes moderately large.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF936E20FF07FB3B9DB8FB49" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BD6BE5FFF936E20FF07FB3B9DB8FB49" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF07FB3B9C1AFB2D" bold="true" box="[199,334,1159,1185]" pageId="24" pageNumber="25">Etymology.</emphasis>
The name, an adjective meaning
<taxonomicName id="4C69C5DCFF936E20FD09FB3B9E43FB2D" box="[713,791,1159,1185]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B91D624DFF936E20FD09FB3B9E43FB2D" box="[713,791,1159,1185]" italics="true" pageId="24" pageNumber="25">tumida</emphasis>
</taxonomicName>
-like, refers to the close resemblance of this species to
<taxonomicName id="4C69C5DCFF936E20FA40FB3B9DB1FB49" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B91D624DFF936E20FA40FB3B9DB1FB49" italics="true" pageId="24" pageNumber="25">W. tumida</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF936E20FF07FB739EDAFA4D" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BD6BE5FFF936E20FF07FB739CD1FABD" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF07FB739C23FB64" bold="true" box="[199,375,1230,1257]" pageId="24" pageNumber="25">Type material.</emphasis>
<materialsCitation id="3B01B402FF936E20FE41FB739CD1FABD" ID-GBIF-Occurrence="3023445536" collectingDate="2016-06-03" collectingDateMax="2016-07-04" collectingDateMin="2016-06-03" collectionCode="CEC, NHRS" collectorName="M. &amp; C. Jaschhof &amp; E. Gustavsson" country="Sweden" county="Gamla Skogsby" location="Male" municipality="Morbylanga" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<emphasis id="B91D624DFF936E20FE41FB739CA1FB65" bold="true" box="[385,501,1231,1257]" pageId="24" pageNumber="25">
<typeStatus id="54D200FDFF936E20FE41FB739CA4FB65" box="[385,496,1231,1257]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EB6E884FF936E20FE3EFB739F6EFB65" LSID="urn:lsid:plazi:treatment:03C00F49FF9D6E20FF57F8EB9F94F987:8EB6E884FF936E20FE3EFB739F6EFB65" box="[510,570,1231,1257]" country="Sweden" county="Gamla Skogsby" municipality="Morbylanga" name="Male" pageId="24" pageNumber="25">Male</location>
,
<collectingCountry id="F37EFECFFF936E20FD88FB739FF7FB65" box="[584,675,1231,1257]" name="Sweden" pageId="24" pageNumber="25">Sweden</collectingCountry>
,
<location id="8EB6E884FF936E20FD71FB6C9FACFB67" LSID="urn:lsid:plazi:treatment:03C00F49FF9D6E20FF57F8EB9F94F987:8EB6E884FF936E20FD71FB6C9FACFB67" box="[689,760,1232,1259]" country="Sweden" county="Gamla Skogsby" municipality="Morbylanga" name="Oland" pageId="24" pageNumber="25">Öland</location>
,
<collectingMunicipality id="6BB22425FF936E20FCC6FB729EC6FB65" box="[774,914,1230,1257]" pageId="24" pageNumber="25">Mörbylånga</collectingMunicipality>
,
<collectingCounty id="62B7C6D3FF936E20FC5FFB739907FB65" box="[927,1107,1231,1257]" pageId="24" pageNumber="25">Gamla Skogsby</collectingCounty>
(Kalkstad), mixed broadleaf forest with plenty of dead ash trees,
<date id="FFD7989FFF936E20FDFDFB4F9E74FA81" box="[573,800,1267,1293]" pageId="24" pageNumber="25" value="2016-06-03" valueMax="2016-07-04" valueMin="2016-06-03">
<collectingDate id="EF936177FF936E20FDFDFB4F9E74FA81" box="[573,800,1267,1293]" pageId="24" pageNumber="25" value="2016-06-03" valueMax="2016-07-04" valueMin="2016-06-03">3 June4 July 2016</collectingDate>
</date>
,
<collectingMethod id="5228C648FF936E20FCEDFB4F9EE9FA81" box="[813,957,1267,1293]" pageId="24" pageNumber="25">Malaise trap</collectingMethod>
,
<collectorName id="269CDB89FF936E20FC0BFB4F99CCFA81" box="[971,1176,1267,1293]" pageId="24" pageNumber="25">M. &amp; C. Jaschhof</collectorName>
&amp;
<collectorName id="269CDB89FF936E20FB00FB4F9837FA81" box="[1216,1379,1267,1293]" pageId="24" pageNumber="25">E. Gustavsson</collectorName>
(spn
<collectionCode id="ED78269AFF936E20FF57FAAB9D9BFABD" box="[151,207,1303,1329]" pageId="24" pageNumber="25">CEC</collectionCode>
<quantity id="4C9113BAFF936E20FF0FFAAB9C77FABD" box="[207,291,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="4.65582" pageId="24" pageNumber="25" unit="in" value="1833.0">1833 in</quantity>
<collectionCode id="ED78269AFF936E20FEE9FAAB9C2DFABD" box="[297,377,1303,1329]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="24" pageNumber="25">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD6BE5FFF936E20FF07FA869EDAFA4D" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF07FA869C8BFAD8" bold="true" box="[199,479,1338,1364]" pageId="24" pageNumber="25">Other material studied.</emphasis>
<materialsCitation id="3B01B402FF936E20FE26FA879DBBFA11" ID-GBIF-Occurrence="3023445417" collectingDate="2015-05-13" collectingDateMax="2015-06-08" collectingDateMin="2015-05-13" collectionCode="MT, MCJ, NHRS" country="Sweden" county="Gamla Skogsby" location="Oland" municipality="Morbylanga" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F37EFECFFF936E20FE26FA879F17FAD9" box="[486,579,1339,1365]" name="Sweden" pageId="24" pageNumber="25">Sweden</collectingCountry>
:
<specimenCount id="9D6F75D6FF936E20FD8EFA879FCDFAD9" box="[590,665,1339,1365]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(only genitalia slide-mounted),
<location id="8EB6E884FF936E20FBC1FA80991CFADB" LSID="urn:lsid:plazi:treatment:03C00F49FF9D6E20FF57F8EB9F94F987:8EB6E884FF936E20FBC1FA80991CFADB" box="[1025,1096,1340,1367]" country="Sweden" county="Gamla Skogsby" municipality="Morbylanga" name="Oland" pageId="24" pageNumber="25">Öland</location>
,
<collectingMunicipality id="6BB22425FF936E20FB93FA86998BFAD9" box="[1107,1247,1338,1365]" pageId="24" pageNumber="25">Mörbylånga</collectingMunicipality>
,
<collectingCounty id="62B7C6D3FF936E20FB2AFA8798C8FAD9" box="[1258,1436,1339,1365]" pageId="24" pageNumber="25">Gamla Skogsby</collectingCounty>
(Kalkstad), ̒diversity meadow, scrubby grassland,
<date id="FFD7989FFF936E20FD22FAE39E82FAF5" box="[738,982,1374,1401]" pageId="24" pageNumber="25" value="2015-05-13" valueMax="2015-06-08" valueMin="2015-05-13">
<collectingDate id="EF936177FF936E20FD22FAE39E82FAF5" box="[738,982,1374,1401]" pageId="24" pageNumber="25" value="2015-05-13" valueMax="2015-06-08" valueMin="2015-05-13">13 May8 June 2015</collectingDate>
</date>
,
<collectionCode id="ED78269AFF936E20FC22FAE39959FAF5" box="[994,1037,1375,1401]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="24" pageNumber="25">MT</collectionCode>
,
<collectionCode id="ED78269AFF936E20FBDAFAE39904FAF5" box="[1050,1104,1375,1401]" country="USA" lsid="urn:lsid:biocol.org:col:13593" name="Missouri Southern State College" pageId="24" pageNumber="25" type="Herbarium">MCJ</collectionCode>
(spn ZFMK-TIS-
<quantity id="4C9113BAFF936E20FADCFAE298C8FAF5" box="[1308,1436,1374,1401]" metricMagnitude="4" metricUnit="m" metricValue="6.47619228" pageId="24" pageNumber="25" unit="in" value="2549682.0">2549682 in</quantity>
<collectionCode id="ED78269AFF936E20FF57FA3F9DB3FA11" box="[151,231,1411,1437]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="24" pageNumber="25">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B01B402FF936E20FF3CFA3F9CB7FA4D" ID-GBIF-Occurrence="3023445408" collectingDate="2015-06-14" collectingDateMax="2015-07-15" collectingDateMin="2015-06-14" collectionCode="MT, MCJ" country="Sweden" location="Morbylanga" municipality="Ullevi" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D6F75D6FF936E20FF3CFA3F9C1EFA11" box="[252,330,1411,1437]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(only genitalia slide-mounted),
<location id="8EB6E884FF936E20FD7CFA3E9E1CFA11" LSID="urn:lsid:plazi:treatment:03C00F49FF9D6E20FF57F8EB9F94F987:8EB6E884FF936E20FD7CFA3E9E1CFA11" box="[700,840,1410,1437]" country="Sweden" municipality="Ullevi" name="Morbylanga" pageId="24" pageNumber="25">Mörbylånga</location>
,
<collectingMunicipality id="6BB22425FF936E20FC95FA3F9ECAFA11" box="[853,926,1411,1437]" pageId="24" pageNumber="25">Ullevi</collectingMunicipality>
, herb-rich meadow near broadleaf forest,
<date id="FFD7989FFF936E20FA40FA3F9C38FA4D" pageId="24" pageNumber="25" value="2015-06-14" valueMax="2015-07-15" valueMin="2015-06-14">
<collectingDate id="EF936177FF936E20FA40FA3F9C38FA4D" pageId="24" pageNumber="25" value="2015-06-14" valueMax="2015-07-15" valueMin="2015-06-14">14 June15 July 2015</collectingDate>
</date>
,
<collectionCode id="ED78269AFF936E20FEB7FA1B9CF6FA4D" box="[375,418,1447,1473]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="24" pageNumber="25">MT</collectionCode>
,
<collectionCode id="ED78269AFF936E20FE6DFA1B9CB7FA4D" box="[429,483,1447,1473]" country="USA" lsid="urn:lsid:biocol.org:col:13593" name="Missouri Southern State College" pageId="24" pageNumber="25" type="Herbarium">MCJ</collectionCode>
</materialsCitation>
<materialsCitation id="3B01B402FF936E20FE2BFA1B9EDEFA4D" ID-GBIF-Occurrence="3023445412" box="[491,906,1446,1473]" collectingDate="2015-06-14" collectingDateMax="2015-07-15" collectingDateMin="2015-06-14" collectionCode="NHRS" country="Sweden" isTailEnumeration="true" location="Morbylanga" municipality="Ullevi" pageId="24" pageNumber="25" specimenCount="1">
(spn ZFMK-TIS-
<quantity id="4C9113BAFF936E20FD6DFA1A9E7FFA4D" box="[685,811,1446,1473]" metricMagnitude="4" metricUnit="m" metricValue="6.475991619999999" pageId="24" pageNumber="25" unit="in" value="2549603.0">2549603 in</quantity>
<collectionCode id="ED78269AFF936E20FCF2FA1B9ED6FA4D" box="[818,898,1447,1473]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="24" pageNumber="25">NHRS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF936E20FF07FA769F94F987" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BD6BE5FFF936E20FF07FA769F94F987" blockId="24.[151,1437,150,1547]" pageId="24" pageNumber="25">
<emphasis id="B91D624DFF936E20FF07FA769F40FA69" bold="true" box="[199,532,1482,1509]" pageId="24" pageNumber="25">Distribution and phenology.</emphasis>
The three specimens known of this species were collected between mid-May and mid-July in both grass- and woodlands on Öland.
</paragraph>
</subSubSection>
</treatment>
</document>