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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.208.89913" ID-Pensoft-Pub="1314-2003-208-103" ID-Pensoft-UUID="88C0802178725C0F86DB6E2074FDD5AB" ModsDocID="1314-2003-208-103" checkinTime="1663785939964" checkinUser="pensoft" docAuthor="Lobban, Christopher S., Ashworth, Matt P., Camacho, Terance, Lam, Daryl W. &amp; Theriot, Edward C." docDate="2022" docId="DD1993EC7F48516D97613E2302BC05FB" docLanguage="en" docName="PhytoKeys 208: 103-184" docOrigin="PhytoKeys 208" docPubDate="2022-09-21" docSource="http://dx.doi.org/10.3897/phytokeys.208.89913" docTitle="Grunowago pacifica Lobban &amp; Ashworth 2022, sp. nov." docType="treatment" docVersion="1" id="88C0802178725C0F86DB6E2074FDD5AB" lastPageNumber="103" masterDocId="88C0802178725C0F86DB6E2074FDD5AB" masterDocTitle="Revision of Ardissoneaceae (Bacillariophyta, Mediophyceae) from Micronesian populations, with descriptions of two new genera, Ardissoneopsis and Grunowago, and new species in Ardissonea, Synedrosphenia and Climacosphenia" masterLastPageNumber="184" masterPageNumber="103" pageNumber="103" updateTime="1663785939964" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Ardissoneaceae (Bacillariophyta, Mediophyceae) from Micronesian populations, with descriptions of two new genera, Ardissoneopsis and Grunowago, and new species in Ardissonea, Synedrosphenia and Climacosphenia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lobban, Christopher S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1596-0656</mods:nameIdentifier>
<mods:affiliation>Division of Natural Sciences, University of Guam, Mangilao, GU 96923, Guam, USA</mods:affiliation>
<mods:nameIdentifier type="email">clobban@guam.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ashworth, Matt P.</mods:namePart>
<mods:affiliation>Department of Molecular Biosciences, The University of Texas at Austin, Austin, Texas, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Camacho, Terance</mods:namePart>
<mods:affiliation>Division of Natural Sciences, University of Guam, Mangilao, GU 96923, Guam, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lam, Daryl W.</mods:namePart>
<mods:affiliation>LSAMP Program, University of Guam, Mangilao, GU 96923, Guam, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Theriot, Edward C.</mods:namePart>
<mods:affiliation>Department of Molecular Biosciences, The University of Texas at Austin, Austin, Texas, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>208</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>103</mods:start>
<mods:end>184</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.208.89913</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.208.89913</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-208-103</mods:identifier>
<mods:identifier type="Pensoft-UUID">88C0802178725C0F86DB6E2074FDD5AB</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:DD1993EC7F48516D97613E2302BC05FB" httpUri="http://treatment.plazi.org/id/DD1993EC7F48516D97613E2302BC05FB" lastPageNumber="103" pageId="0" pageNumber="103">
<subSubSection pageId="0" pageNumber="103" type="nomenclature">
<paragraph pageId="0" pageNumber="103">
<taxonomicName LSID="DD1993EC-7F48-516D-9761-3E2302BC05FB" authority="Lobban &amp; Ashworth" authorityName="Lobban &amp; Ashworth" authorityYear="2022" family="Ardissoneaceae" genus="Grunowago" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Grunowago pacifica" order="Ardissoneales" pageId="0" pageNumber="103" rank="species" species="pacifica" status="sp. nov.">Grunowago pacifica Lobban &amp; Ashworth</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="103">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">Figs 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">, 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="references">
<paragraph pageId="0" pageNumber="103">References.</paragraph>
<paragraph pageId="0" pageNumber="103">
<bibRefCitation author="Lobban, CS" journalOrPublisher="Micronesica" pageId="0" pageNumber="103" pagination="237 - 479" publicationUrl="https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" refId="B41" refString="Lobban, CS, Schefter, M, Jordan, RW, Arai, Y, Sasaki, A, Theriot, EC, Ashworth, M, Ruck, EC, Pennesi, C, 2012. Coral-reef diatoms (Bacillariophyta) from Guam: New records and preliminary checklist, with emphasis on epiphytic species from farmer-fish territories. Micronesica 43: 237 - 479, https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" title="Coral-reef diatoms (Bacillariophyta) from Guam: New records and preliminary checklist, with emphasis on epiphytic species from farmer-fish territories." url="https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" volume="43" year="2012">Lobban et al. 2012</bibRefCitation>
, pp. 256-257, pl. 13, figs 1, 2 [as
<taxonomicName authorityName="Hustedt" authorityYear="1932" baseAuthorityName="Grunow" class="Bacillariophyceae" family="Ardissoneaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Synedra bacillaris" order="Ardissoneales" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="bacillaris">
<emphasis italics="true" pageId="0" pageNumber="103">Synedra bacillaris</emphasis>
</taxonomicName>
(Grunow) Hustedt].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="diagnosis">
<paragraph pageId="0" pageNumber="103">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="103">
Differing from
<taxonomicName authorityName="Lobban &amp; Ashworth" authorityYear="2022" baseAuthorityName="Lobban &amp; Ashworth &amp; Camacho &amp; Lam &amp; Theriot" baseAuthorityYear="1932" family="Ardissoneaceae" genus="Grunowago" kingdom="Plantae" lsidName="Grunowago (Synedra) bacillaris" order="Ardissoneales" pageId="0" pageNumber="103" rank="species" species="bacillaris" subGenus="Synedra">Grunowago (Synedra) bacillaris</taxonomicName>
in width and lanceolate outline of valve.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">Description.</paragraph>
<paragraph pageId="0" pageNumber="103">
Valves lanceolate, often slightly rostrate, 194-341
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 11-16
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, striae 9-10 in 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20A-D</figureCitation>
), the striae comprising large oval pores; poles with smaller, circular pores, but not organized into polar pore fields (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20E</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A</figureCitation>
). Isopolar but wider in the middle in girdle view (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ardissoneopsis gracilis sp. nov. (GU 44 BJ- 4, except B TK 28) A-C three specimens to scale in LM to show size range: valves from Guam (A) and Chuuk (B), frustule from Guam in girdle view (C) showing heteropolarity D frustule in SEM tilted 60 °, showing both valve and girdle faces E, F pole and mid-section of holotype G-K external views of valve in SEM with other species adjacent for comparison (GU 44 BJ- 4): index image (G) with locations of details marked, apical pole (H) (shown by depth of mantle), compare stria density with Ardissoneopsis appressata (A. g.); top half of center (I), offset areolae in striae within the oval suggest location of annulus; narrow portion below center (J), compare with scattered areolae in adjacent Toxarium undulatum (T. u.); basal pole (K), note spines on both poles L, M internal aspect near middle and at the apical pole, showing lack of costae N, O girdle view of frustule apical and basal poles, showing taper in mantle and girdle bands P possible piece of pleura (pl?) with fimbriae Q, R fimbriate edges of valvocopula and copula, respectively. Scale bars: 50 µm (A-D, G); 10 µm (E, F, H-K); 5 µm (L-R)." figureDoi="10.3897/phytokeys.208.89913.figure18" httpUri="https://binary.pensoft.net/fig/746734" pageId="0" pageNumber="103">18F-H</figureCitation>
). Two small spines usually present on each pole (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20G-I</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21B</figureCitation>
). Rim present on the valve border but not extended into a pseudoseptum (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A</figureCitation>
). Valve single-walled (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20J</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A</figureCitation>
), internally costate with a distinctive central costa the entire length of valve and thickened vimines (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20J</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A</figureCitation>
), no annulus detected either as a break in the striae, or any other longitudinal costae (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A</figureCitation>
). Three girdle bands were present (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20F-H</figureCitation>
). Valvocopula (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20I, J</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A, C</figureCitation>
) without extended lip but having groove on pars interior that fits rim of valve. A single row of pores on advalvar side of pars exterior, circular except elongated near poles (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20G, H</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A, C</figureCitation>
), a short line of pores around poles on pars interior (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20I</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21C</figureCitation>
). Copula has a row of pores on outer edge of pars interior (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21A, B, D</figureCitation>
), often hidden under valvocopula and a well-developed fringe on the pars interior; up to 5 rows of pores on pars exterior (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20G, H</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21B, D</figureCitation>
). Pleura a very thin fimbriate band, continuous along cell length (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20G, H</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Grunowago pacifica sp. nov., cont., all GU 44 BJ- 4, except A = SPN 2022 - 1 - 4 A valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum B frustule at 40 ° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow) C valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead) D frustule in girdle view with fimbriate pars interior of pleura exposed (arrow) E frustule in polar view tilted 60 °, showing girdle bands. Scale bars: 10 µm (A-D); 5 µm (E)." figureDoi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103">21B, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 20.</emphasis>
<taxonomicName authorityName="Lobban &amp; Ashworth" authorityYear="2022" family="Ardissoneaceae" genus="Grunowago" kingdom="Plantae" lsidName="Grunowago pacifica" order="Ardissoneales" pageId="0" pageNumber="103" rank="species" species="pacifica">
<emphasis italics="true" pageId="0" pageNumber="103">Grunowago pacifica</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="103">A-D</emphasis>
LM of specimens from GU44BJ-4 (
<emphasis bold="true" pageId="0" pageNumber="103">A-C</emphasis>
) and GU44Z-15 (
<emphasis bold="true" pageId="0" pageNumber="103">D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="103">D, E</emphasis>
holotype specimen
<emphasis bold="true" pageId="0" pageNumber="103">F-H</emphasis>
frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU44BJ-4)
<emphasis bold="true" pageId="0" pageNumber="103">I</emphasis>
external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow)
<emphasis bold="true" pageId="0" pageNumber="103">J</emphasis>
internal view of pole and abvalvar surface of valvocopula. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="103">A-D, F</emphasis>
); 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="103">E, G-J</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.208.89913.figure21" httpUri="https://binary.pensoft.net/fig/746737" pageId="0" pageNumber="103" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 21.</emphasis>
<taxonomicName authorityName="Lobban &amp; Ashworth" authorityYear="2022" family="Ardissoneaceae" genus="Grunowago" kingdom="Plantae" lsidName="Grunowago pacifica" order="Ardissoneales" pageId="0" pageNumber="103" rank="species" species="pacifica">
<emphasis italics="true" pageId="0" pageNumber="103">Grunowago pacifica</emphasis>
</taxonomicName>
sp. nov., cont., all GU44BJ-4, except
<emphasis bold="true" pageId="0" pageNumber="103">A</emphasis>
= SPN2022-1-4
<emphasis bold="true" pageId="0" pageNumber="103">A</emphasis>
valve apex showing continuous rim, broken away at lower right, not expanded into a pseudoseptum
<emphasis bold="true" pageId="0" pageNumber="103">B</emphasis>
frustule at 40° tilt, showing fimbriate inner margin of copula on the hypotheca (top) and line of pores on pars interior (arrow)
<emphasis bold="true" pageId="0" pageNumber="103">C</emphasis>
valvocopula, showing internal and external aspects, the pars exterior pores (arrow) separated by ridges on the pars interior, and the presence of pores on the pars interior at the pole (arrowhead)
<emphasis bold="true" pageId="0" pageNumber="103">D</emphasis>
frustule in girdle view with fimbriate pars interior of pleura exposed (arrow)
<emphasis bold="true" pageId="0" pageNumber="103">E</emphasis>
frustule in polar view tilted 60°, showing girdle bands. Scale bars: 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="103">A-D</emphasis>
); 5
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="103">E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="holotype">
<paragraph pageId="0" pageNumber="103">Holotype</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">(designated here).</emphasis>
Specimen at 18.3 mm E and 6.4 mm S of the mark on slide 146, deposited at ANSP accession # ANSP-GC20085. Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Grunowago pacifica sp. nov. A-D LM of specimens from GU 44 BJ- 4 (A-C) and GU 44 Z- 15 (D) D, E holotype specimen F-H frustule in girdle view, showing isopolar frustule wider in the middle and three girdle bands (GU 44 BJ- 4) I external view of pole and advalvar surface of valvocopula, the latter showing a short row of small pores on the apical part of the pars interior (arrowhead) and the crenulated edge of the pars interior (arrow) J internal view of pole and abvalvar surface of valvocopula. Scale bars: 50 µm (A-D, F); 10 µm (E, G-J)." figureDoi="10.3897/phytokeys.208.89913.figure20" httpUri="https://binary.pensoft.net/fig/746736" pageId="0" pageNumber="103">20D, E</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="registration">
<paragraph pageId="0" pageNumber="103">Registration.</paragraph>
<paragraph pageId="0" pageNumber="103">Phycobank http://phycobank.org/103245.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="type locality">
<paragraph pageId="0" pageNumber="103">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="103">
GabGab reef, Apra Harbor, Guam,
<geoCoordinate degrees="13.443" direction="north" orientation="latitude" precision="55" value="13.443">13.443°N</geoCoordinate>
,
<geoCoordinate degrees="144.643" direction="east" orientation="longitude" precision="55" value="144.643">144.643°E</geoCoordinate>
, ca. 1 m depth, collection number GU44Z-15, 20 June 2009. C.S. Lobban and M. Schefter leg. Additional slides from this sample have previously been deposited at California Academy of Sciences (CAS) for other species holotypes, but specimens of
<taxonomicName lsidName="G. pacifica" pageId="0" pageNumber="103" rank="species" species="pacifica">
<emphasis italics="true" pageId="0" pageNumber="103">G. pacifica</emphasis>
</taxonomicName>
are not marked. Slides include CAS accession/slide numbers: 627386/223007 (
<taxonomicName authorityName="C.S.Lobban &amp; R.W.Jordan" authorityYear="2011" class="Bacillariophyceae" family="Fragilariaceae" genus="Perideraion" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="Perideraion" order="Fragilariales" pageId="0" pageNumber="103" phylum="Bacillariophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Perideraion</emphasis>
</taxonomicName>
spp.), 627383/223005 (
<taxonomicName authorityName="Lobban &amp; J.N.Navarro" authorityYear="2013" class="Bacillariophyceae" family="Mastogloiaceae" genus="Gato" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="Gato hyalinus" order="Mastogloiales" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="hyalinus">
<emphasis italics="true" pageId="0" pageNumber="103">Gato hyalinus</emphasis>
</taxonomicName>
); 627409/223023 (
<taxonomicName authorityName="Ashworth &amp; Lobban" authorityYear="2012" class="Bacillariophyceae" genus="Astrosyne" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="Astrosyne radiata" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="radiata">
<emphasis italics="true" pageId="0" pageNumber="103">Astrosyne radiata</emphasis>
</taxonomicName>
); and 627396/223010 (
<taxonomicName authorityName="C.S.Lobban &amp; M.P.Ashworth" authorityYear="2014" class="Bacillariophyceae" family="Grammatophoraceae" genus="Hanicella" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="Hanicella moenia" order="Rhabdonematales" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="moenia">
<emphasis italics="true" pageId="0" pageNumber="103">Hanicella moenia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="materials_examined">
<paragraph pageId="0" pageNumber="103">Additional materials examined.</paragraph>
<paragraph pageId="0" pageNumber="103">
<collectingCountry name="Guam">Guam</collectingCountry>
: GU44L-C!, GU44AI-5! (in culture), GU44AP-8!, GU44BH-5!.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="etymology">
<paragraph pageId="0" pageNumber="103">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="103">Adjective (Latin) with reference to its presence in the Western Pacific Ocean.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="taxonomic comments">
<paragraph pageId="0" pageNumber="103">Taxonomic comments.</paragraph>
<paragraph pageId="0" pageNumber="103">
Specimens that we (
<bibRefCitation author="Lobban, CS" journalOrPublisher="Micronesica" pageId="0" pageNumber="103" pagination="237 - 479" publicationUrl="https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" refId="B41" refString="Lobban, CS, Schefter, M, Jordan, RW, Arai, Y, Sasaki, A, Theriot, EC, Ashworth, M, Ruck, EC, Pennesi, C, 2012. Coral-reef diatoms (Bacillariophyta) from Guam: New records and preliminary checklist, with emphasis on epiphytic species from farmer-fish territories. Micronesica 43: 237 - 479, https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" title="Coral-reef diatoms (Bacillariophyta) from Guam: New records and preliminary checklist, with emphasis on epiphytic species from farmer-fish territories." url="https://micronesica.org/sites/default/files/6_lobban.schefter_micronesica_431.pdf" volume="43" year="2012">Lobban et al. 2012</bibRefCitation>
) referred to
<taxonomicName authorityName="Hustedt" authorityYear="1932" baseAuthorityName="Grunow" class="Bacillariophyceae" family="Ardissoneaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Synedra bacillaris" order="Ardissoneales" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="bacillaris">
<emphasis italics="true" pageId="0" pageNumber="103">Synedra bacillaris</emphasis>
</taxonomicName>
were 180-400
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 14
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, with 9-14 striae in 10
<normalizedToken originalValue="µm">µm</normalizedToken>
. This range of stria density is suspiciously large for
<taxonomicName authorityName="Lobban &amp; Ashworth &amp; Camacho &amp; Lam &amp; Theriot" authorityYear="2022" baseAuthorityName="Lobban &amp; Ashworth &amp; Camacho &amp; Lam &amp; Theriot" baseAuthorityYear="2022" class="Bacillariophyceae" family="Ardissoneaceae" kingdom="Plantae" lsidName="" order="Ardissoneales" pageId="0" pageNumber="103" rank="family">Ardissoneaceae</taxonomicName>
and suggests that two or more species are being confounded. Some specimens in GU44BJ-4 population were nearly linear but most likely represent an extreme within
<taxonomicName lsidName="G. pacifica" pageId="0" pageNumber="103" rank="species" species="pacifica">
<emphasis italics="true" pageId="0" pageNumber="103">G. pacifica</emphasis>
</taxonomicName>
rather than an occasional
<taxonomicName lsidName="G. bacillaris" pageId="0" pageNumber="103" rank="species" species="bacillaris">
<emphasis italics="true" pageId="0" pageNumber="103">G. bacillaris</emphasis>
</taxonomicName>
. Both
<bibRefCitation author="Hustedt, F" journalOrPublisher="Hidrobiologica" pageId="0" pageNumber="103" refId="B25" refString="Hustedt, F, 1931-1959. Die Kieselalgen Deutschlands, Oesterreichs und der Schweiz. In: Rabenhorst's Kryptogamenflora, Band 7, Teil 2. Johnson Reprint, New York." title="Die Kieselalgen Deutschlands, Oesterreichs und der Schweiz. In: Rabenhorst's Kryptogamenflora, Band 7, Teil 2. Johnson Reprint, New York." year="1931 - 1959">Hustedt (1931-1959)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.1080/0269249X.1995.9705336" author="Sullivan, MJ" journalOrPublisher="Diatom Research" pageId="0" pageNumber="103" pagination="179 - 190" refId="B75" refString="Sullivan, MJ, Wear, DJ, 1995. A morphological study of the giant diatoms Ardissonea formosa and Synedra bacillaris. Diatom Research 10 (1): 179 - 190, DOI: https://doi.org/10.1080/0269249X.1995.9705336" title="A morphological study of the giant diatoms Ardissonea formosa and Synedra bacillaris." url="https://doi.org/10.1080/0269249X.1995.9705336" volume="10" year="1995">Sullivan and Wear (1995)</bibRefCitation>
remarked on the ease of identifying this species by the strong central costa, compared to other
<taxonomicName class="Bacillariophyceae" family="Ardissoneaceae" genus="Ardissonea" kingdom="Plantae" lsidName="Ardissonea" order="Ardissoneales" pageId="0" pageNumber="103" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Ardissonea</emphasis>
</taxonomicName>
spp., but as with other taxa where a strong identifying character has masked species diversity (e.g.,
<taxonomicName authorityName="Castracane" authorityYear="1878" class="Insecta" family="Geometridae" genus="Cyclophora" kingdom="Animalia" lsidName="Cyclophora tenuis" order="Lepidoptera" pageId="0" pageNumber="103" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis italics="true" pageId="0" pageNumber="103">Cyclophora tenuis</emphasis>
</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.2216/12-004.1" author="Ashworth, M" journalOrPublisher="Phycologia" pageId="0" pageNumber="103" pagination="684 - 699" refId="B4" refString="Ashworth, M, Ruck, EC, Lobban, CS, Romanovicz, DK, Theriot, EC, 2012. A revision of the genus Cyclophora and description of Astrosyne gen. nov. (Bacillariophyta), two genera with the pyrenoids contained within pseudosepta. Phycologia 51 (6): 684 - 699, DOI: https://doi.org/10.2216/12-004.1" title="A revision of the genus Cyclophora and description of Astrosyne gen. nov. (Bacillariophyta), two genera with the pyrenoids contained within pseudosepta." url="https://doi.org/10.2216/12-004.1" volume="51" year="2012">Ashworth et al. 2012</bibRefCitation>
,
<taxonomicName authorityName="Round" authorityYear="1990" baseAuthorityName="Grunow" class="Bacillariophyceae" family="Fragilariaceae" genus="Bleakeleya" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="Bleakeleya notata" order="Fragilariales" pageId="0" pageNumber="103" phylum="Bacillariophyta" rank="species" species="notata">
<emphasis italics="true" pageId="0" pageNumber="103">Bleakeleya notata</emphasis>
</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.1127/nova_hedwigia/2016/0327" author="Lobban, CS" journalOrPublisher="Nova Hedwigia" pageId="0" pageNumber="103" pagination="547 - 552" refId="B38" refString="Lobban, CS, Perez, CO, 2016. A second species of bleakeleyoid diatoms (Bacillariophyta, Fragilariales), Koernerella grunowii, from Grunow's Honduras Sargassum materials. Nova Hedwigia 102 (3-4): 547 - 552, DOI: https://doi.org/10.1127/nova_hedwigia/2016/0327" title="A second species of bleakeleyoid diatoms (Bacillariophyta, Fragilariales), Koernerella grunowii, from Grunow's Honduras Sargassum materials." url="https://doi.org/10.1127/nova_hedwigia/2016/0327" volume="102" year="2016">Lobban and Perez 2016</bibRefCitation>
), we suspect that there is hidden diversity within &quot;
<taxonomicName authorityName="Hustedt" authorityYear="1932" baseAuthorityName="Grunow" class="Bacillariophyceae" family="Ardissoneaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Synedra bacillaris" order="Ardissoneales" pageId="0" pageNumber="103" phylum="Ochrophyta" rank="species" species="bacillaris">
<emphasis italics="true" pageId="0" pageNumber="103">Synedra bacillaris</emphasis>
</taxonomicName>
&quot;. The genetic branches are long between lanceolate samples from Guam (GU44AI-5) and Florida (HK291) (see, below, Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Phylogenetic trees resulting from three analytical methods of the 3 - gene DNA sequence dataset assembled for this manuscript. Only the Ardissoneaceae and their sister group in the molecular phylogeny, Lampriscus, is shown here-the complete trees can be found in Suppl. materials 5 - 7: Figs S 3 - S 5 A Maximum Likelihood phylogeny resolved by RAxML, where values at nodes are bootstrap support values B Maximum Likelihood phylogeny resolved by IQTREE, where values at nodes are bootstrap support values C Bayesian Inference phylogeny resolved by ExaBayes, where values at nodes are posterior probability values. Strain taxonomy (name followed by DNA voucher ID or strain ID) follows currently accepted classification, with proposed genera based on morphology shown to the right of strain ID." figureDoi="10.3897/phytokeys.208.89913.figure33" httpUri="https://binary.pensoft.net/fig/746749" pageId="0" pageNumber="103">33</figureCitation>
). For now, we can only describe
<taxonomicName authorityName="Lobban &amp; Ashworth" authorityYear="2022" family="Ardissoneaceae" genus="Grunowago" kingdom="Plantae" lsidName="Grunowago pacifica" order="Ardissoneales" pageId="0" pageNumber="103" rank="species" species="pacifica">
<emphasis italics="true" pageId="0" pageNumber="103">Grunowago pacifica</emphasis>
</taxonomicName>
and deal with the generic placement of that species cluster.
</paragraph>
</subSubSection>
</treatment>
</document>
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