351 lines
42 KiB
XML
351 lines
42 KiB
XML
<document id="6EE89813B8D725236F9DEB68F1FB5E7A" ID-CLB-Dataset="27715" ID-DOI="10.11646/zootaxa.4541.1.1" ID-GBIF-Dataset="9c3fecf1-4e11-4c79-a908-e0a5abbaf3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617460" ID-ZooBank="B11E734C-218B-418C-84E6-719AB3C58AFF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927824231" checkinUser="plazi" docAuthor="Arachchige, Gayashan M., Jayakody, Sevvandi, Mooi, Rich & Kroh, Andreas" docDate="2019" docId="038087B4FFE78927FF02FDD872369BE8" docLanguage="en" docName="zootaxa.4541.1.1.pdf" docOrigin="Zootaxa 4541 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Metalia Gray 1855" docType="treatment" docVersion="8" lastPageNumber="83" masterDocId="FFB9FFCCFFAA8975FF95FFB47314996C" masterDocTitle="Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka" masterLastPageNumber="100" masterPageNumber="1" pageNumber="78" updateTime="1698715306146" updateUser="ExternalLinkService">
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<mods:mods id="3E2513AD0A8EABB9A32D5C0AC825FBFC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7C45344DCA62AC00D09A813C30F0E61A">
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<mods:title id="148571287A02EEC3738A6FDE55F9301C">Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka</mods:title>
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</mods:titleInfo>
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<mods:name id="C15D894B72935C9B5E58184023D5CD20" type="personal">
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<mods:namePart id="5557E6BD2C9DB3F5F6F65DCAA0EE9CEF">Arachchige, Gayashan M.</mods:namePart>
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<mods:namePart id="E1BEF6F8BE39D041BF3015656BD5713F">Jayakody, Sevvandi</mods:namePart>
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<mods:namePart id="88658D0B0677006E0A9D52BE97F8C64B">Mooi, Rich</mods:namePart>
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</mods:name>
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<mods:name id="11722C81F0A0F4EBA951668704AFD32D" type="personal">
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<mods:roleTerm id="A91C486788F56172CB355FC7FBC6952B">Author</mods:roleTerm>
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<mods:namePart id="C9C37BAF58919FE15B9F6B0450205C52">Kroh, Andreas</mods:namePart>
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<mods:typeOfResource id="300495906D0C0ACDE84ED4C81F2F771A">text</mods:typeOfResource>
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<mods:title id="FF4AE601379B8149A85E218A866C0C6A">Zootaxa</mods:title>
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<mods:part id="C2C0A8BDE576F355CCF8008E6F2FEBF9">
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<mods:date id="DD91E6649A469B1CE1746349CCB93757">2019</mods:date>
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<mods:number id="5F8BC167AFB50854379625CC672C8B51">2019-01-03</mods:number>
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<mods:number id="4CA8109B94A9448FCBEE2CEF3391AC24">4541</mods:number>
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<mods:identifier id="17D9E638BE5248B3E5D65EBD306575CD" type="DOI">10.11646/zootaxa.4541.1.1</mods:identifier>
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<treatment id="038087B4FFE78927FF02FDD872369BE8" ID-DOI="http://doi.org/10.5281/zenodo.5935468" ID-GBIF-Taxon="154529626" ID-Zenodo-Dep="5935468" LSID="urn:lsid:plazi:treatment:038087B4FFE78927FF02FDD872369BE8" httpUri="http://treatment.plazi.org/id/038087B4FFE78927FF02FDD872369BE8" lastPageId="82" lastPageNumber="83" pageId="77" pageNumber="78">
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<subSubSection id="C3336529FFE78938FF02FDD8720A9BEA" box="[151,286,620,646]" pageId="77" pageNumber="78" type="nomenclature">
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<paragraph id="8B9636A2FFE78938FF02FDD8720A9BEA" blockId="77.[151,312,620,679]" box="[151,286,620,646]" pageId="77" pageNumber="78">
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<heading id="D0DE81CEFFE78938FF02FDD8720A9BEA" bold="true" box="[151,286,620,646]" fontSize="11" level="1" pageId="77" pageNumber="78" reason="1">
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<emphasis id="B95DEAB0FFE78938FF02FDD8720A9BEA" bold="true" box="[151,286,620,646]" pageId="77" pageNumber="78">
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<taxonomicName id="4C294D21FFE78938FF02FDD873E09BEA" ID-CoL="7PNXP" authorityName="Gray" authorityYear="1855" box="[151,244,620,646]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="genus">
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<emphasis id="B95DEAB0FFE78938FF02FDD873E09BEA" bold="true" box="[151,244,620,646]" italics="true" pageId="77" pageNumber="78">Metalia</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFE78938FF02FD3A722C9BCB" box="[151,312,654,679]" pageId="77" pageNumber="78" type="description">
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||
<paragraph id="8B9636A2FFE78938FF02FD3A722C9BCB" blockId="77.[151,312,620,679]" box="[151,312,654,679]" pageId="77" pageNumber="78">
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<figureCitation id="13122A27FFE78938FF02FD3A722C9BCB" box="[151,312,654,679]" captionStart-0="FIGURE 68" captionStart-1="FIGURE 69" captionStart-2="FIGURE 70" captionStart-3="FIGURE 71" captionStartId-0="78.[151,250,1927,1949]" captionStartId-1="79.[151,250,1820,1842]" captionStartId-2="80.[151,250,1820,1842]" captionStartId-3="81.[151,250,1877,1899]" captionTargetBox-0="[151,1436,301,1906]" captionTargetBox-1="[151,1436,193,1798]" captionTargetBox-2="[151,1436,193,1798]" captionTargetBox-3="[182,1426,219,1839]" captionTargetId-0="figure@78.[151,1436,301,1906]" captionTargetId-1="figure@79.[151,1436,193,1798]" captionTargetId-2="figure@80.[151,1436,193,1798]" captionTargetId-3="figure@81.[151,1436,193,1856]" captionTargetPageId-0="78" captionTargetPageId-1="79" captionTargetPageId-2="80" captionTargetPageId-3="81" captionText-0="FIGURE 68. Metalia sp. (WUSL/EI/85): A, aboral view; B, oral view; C, lateral view; D, anterior view; E, posterior view." captionText-1="FIGURE 69. Metalia sp. (WUSL/EI/138), aboral view." captionText-2="FIGURE 70. Metalia sp. (WUSL/EI/85): A, sternal view; B, subanal fasciole view." captionText-3="FIGURE 71. Distribution map of Metalia sp. recorded in this study." httpUri-0="https://zenodo.org/record/2617591/files/figure.png" httpUri-1="https://zenodo.org/record/2617593/files/figure.png" httpUri-2="https://zenodo.org/record/2617595/files/figure.png" httpUri-3="https://zenodo.org/record/2617597/files/figure.png" pageId="77" pageNumber="78">Figures 68–71</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFE78938FF02FD6276109A7F" pageId="77" pageNumber="78" type="materials_examined">
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<paragraph id="8B9636A2FFE78938FF02FD6276109A7F" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF02FD6272709B83" bold="true" box="[151,356,726,751]" pageId="77" pageNumber="78">Material studied.</emphasis>
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Four denuded tests: WUSL/EI/85 and EI/86, from Kamburugamuwa and WUSL/EI/138 and EI/ 139 from Negombo 1,
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<collectingCountry id="F33E7632FFE78938FE02FD4E711C9A7F" box="[407,520,762,787]" name="Sri Lanka" pageId="77" pageNumber="78">Sri Lanka</collectingCountry>
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; one complete specimen, WUSL/EI/85 and three broken specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFE78938FF52FCAA76949FB3" pageId="77" pageNumber="78" type="description">
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<paragraph id="8B9636A2FFE78938FF52FCAA760A9A36" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FCAA724D9A5B" bold="true" box="[199,345,798,823]" pageId="77" pageNumber="78">Description.</emphasis>
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<emphasis id="B95DEAB0FFE78938FEF9FCAA71389A5A" box="[364,556,798,823]" italics="true" pageId="77" pageNumber="78">Shape and size</emphasis>
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—Test large, robust 121.0–
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<quantity id="4CD19B47FFE78938FC11FCAB77169A5A" box="[900,1026,799,823]" metricMagnitude="-1" metricUnit="m" metricValue="1.443" pageId="77" pageNumber="78" unit="mm" value="144.3">144.3 mm</quantity>
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TL; width 89–92% TL; height approximately 50% TL; test oval in outline when viewed aborally; no ambital notch in ambulacrum III.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52FCD277429ACF" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FCD2727C9A12" box="[199,360,870,895]" italics="true" pageId="77" pageNumber="78">Apical system</emphasis>
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—Ethmolytic, with four large circular gonopores; madreporite expanding beyond posterior oculars; length 4–5% TL; situated anteriorly, 40–43% TL away from anterior margin.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52FC1A727E9DAE" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FC1A72579AAB" box="[199,323,942,967]" italics="true" pageId="77" pageNumber="78">Ambulacra</emphasis>
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—Ambulacrum III narrow and flush with test, with pore pairs not differentiated to support respiratory tube feet; paired ambulacra distinctly petaloid; petals deeply sunken towards ambitus, straight; each anterior paired petal extending 92–96% of corresponding test radius; anterior paired petals forming angle of 135–145°; anterior and posterior paired petals about equal, length 39–42% TL, width 5–6% TL; angle between anterior paired petals slightly larger than angle between posterior paired petals; interporiferous zones narrow, of equal width, c. 1% TL; pores large, about equal in size, connected by deep groove; adorally well-developed phyllodes occur, featuring 17–18 pores in the anterior paired ambulacra,
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<quantity id="4CD19B47FFE78938FC55FB3370FE9DF3" box="[960,1002,1158,1183]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="77" pageNumber="78" unit="in" value="8.0">8 in</quantity>
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posterior paired ambulacra, and
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<quantity id="4CD19B47FFE78938FAC2FB3373B99DAF" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="77" pageNumber="78" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
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ambulacrum III.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52FB7A713C9C3F" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FB7A72779D8B" box="[199,355,1230,1255]" italics="true" pageId="77" pageNumber="78">Tuberculation</emphasis>
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—Aboral surface densely packed with primary tubercles; larger tubercles inside peripetalous fasciole, bordering ambulacrum III and scattered over posterior interambulacra, usually arranged in short, oblique bands; primary tubercles on oral side larger than aboral ones; primary tubercles in plastron much smaller than those of oral interambulacra 1, 2, 3 and 4.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52FAEA77B29C8F" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FAEA72339C1B" box="[199,295,1374,1399]" italics="true" pageId="77" pageNumber="78">Fasciole</emphasis>
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—Peripetalous fasciole well-developed, closely outlining petals; fasciole strongly indented between petals; subanal fasciole shield-shaped, wider than long; length 16–18% TL, width 22–27% TL; tapering to one point at posterior edge of plastron; upper part shallowly indented; usually four pore pairs enclosed on each side of subanal fasciole; anal branches extending aborally to points even with midpoint of periproct.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52FA5A708C9F6B" blockId="77.[151,1437,726,2011]" box="[199,920,1518,1543]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52FA5A72329F6B" box="[199,294,1518,1543]" italics="true" pageId="77" pageNumber="78">Plastron</emphasis>
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—Narrow, elongate, width 19–20% TL, slightly keeled.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52F9A0764E9F23" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52F9A0722D9F47" box="[199,313,1556,1579]" italics="true" pageId="77" pageNumber="78">Peristome</emphasis>
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—Kidney-shaped; situated anteriorly, c. 21% TL from anterior margin of test; about twice as wide as long; moderately large, length c. 6% TL, width c. 14% TL; from labrum pointed and overhanging peristome.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52F9E872F19FFA" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52F9E872259F1F" box="[199,305,1628,1651]" italics="true" pageId="77" pageNumber="78">Periproct</emphasis>
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—Vertically elongate; pointed aborally and orally; large, length 10–11% TL, width 5–7% TL; region just oral to periproct concave.
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</paragraph>
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<paragraph id="8B9636A2FFE78938FF52F91676949FB3" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52F91671789FD7" bold="true" box="[199,620,1698,1723]" pageId="77" pageNumber="78">
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Observed occurrence in
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<materialsCitation id="3B413CFFFFE78938FE79F91671789FD7" ID-GBIF-Occurrence="2147524162" box="[492,620,1698,1723]" country="Sri Lanka" elevation="23" location="Specimens" pageId="77" pageNumber="78" specimenCount="1">
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<collectingCountry id="F33E7632FFE78938FE79F91671739FD7" box="[492,615,1698,1723]" name="Sri Lanka" pageId="77" pageNumber="78">Sri Lanka</collectingCountry>
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.
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</materialsCitation>
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</emphasis>
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<location id="8EF66079FFE78938FDE1F91671F99FD7" LSID="urn:lsid:plazi:treatment:038087B4FFE78927FF02FDD872369BE8:8EF66079FFE78938FDE1F91671F99FD7" box="[628,749,1698,1723]" country="Sri Lanka" name="Specimens" pageId="77" pageNumber="78">Specimens</location>
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were collected at
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<elevation id="0004D191FFE78938FC2EF916770A9FD6" box="[955,1054,1698,1723]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="77" pageNumber="78" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">
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<quantity id="4CD19B47FFE78938FC2EF916770A9FD6" box="[955,1054,1698,1723]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="77" pageNumber="78" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">20–25 m</quantity>
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</elevation>
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from sand patches between patch reefs in Kamburugamuwa, southern coast, and from
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<elevation id="0004D191FFE78938FD76F97370529FB2" box="[739,838,1735,1759]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="77" pageNumber="78" unit="m" value="27.5" valueMax="28.0" valueMin="27.0">
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<quantity id="4CD19B47FFE78938FD76F97370529FB2" box="[739,838,1735,1759]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="77" pageNumber="78" unit="m" value="27.5" valueMax="28.0" valueMin="27.0">27–28 m</quantity>
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||
</elevation>
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at Negombo, western coast of Sri Lanka (
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<figureCitation id="13122A27FFE78938FA8AF97276669FB3" box="[1311,1394,1734,1759]" captionStart="FIGURE 71" captionStartId="81.[151,250,1877,1899]" captionTargetBox="[182,1426,219,1839]" captionTargetId="figure@81.[151,1436,193,1856]" captionTargetPageId="81" captionText="FIGURE 71. Distribution map of Metalia sp. recorded in this study." httpUri="https://zenodo.org/record/2617597/files/figure.png" pageId="77" pageNumber="78">Fig. 71</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFE78927FF52F95E72369BE8" lastPageId="82" lastPageNumber="83" pageId="77" pageNumber="78" type="discussion">
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||
<paragraph id="8B9636A2FFE78938FF52F95E73CB9EFF" blockId="77.[151,1437,726,2011]" pageId="77" pageNumber="78">
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<emphasis id="B95DEAB0FFE78938FF52F95E722F9E6F" bold="true" box="[199,315,1770,1795]" pageId="77" pageNumber="78">Remarks.</emphasis>
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As noted by one of us (R. Mooi, unpublished and
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<bibRefCitation id="EFB84B53FFE78938FCFBF95E77479E6F" author="Askin, R. & Mooi, R." box="[878,1107,1770,1795]" pageId="77" pageNumber="78" pagination="7" refId="ref33641" refString="Askin, R. & Mooi, R. (2017) The heart of the problem: Phylogenetic systematics of burrowing sea urchins in the genus Metalia (Echinoidea: Spatangoida). Integrative and Comparative Biology, 57 (Supplement 1), e 7." type="journal article" year="2017">Askin & Mooi 2017</bibRefCitation>
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), these specimens differ from the 10 known, extant congeners,
|
||
<taxonomicName id="4C294D21FFE78938FD80F8BB70709E4B" authority="de Ridder, 1984" authorityName="de Ridder" authorityYear="1984" box="[533,868,1806,1831]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="angustus">
|
||
<emphasis id="B95DEAB0FFE78938FD80F8BB71B59E4A" box="[533,673,1807,1830]" italics="true" pageId="77" pageNumber="78">M. angustus</emphasis>
|
||
de Ridder, 1984
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FCE7F8BB77D59E4B" authority="H. L. Clark, 1917" authorityName="H. L. Clark" authorityYear="1917" box="[882,1217,1806,1831]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="dicrana">
|
||
<emphasis id="B95DEAB0FFE78938FCE7F8BB70E59E4B" box="[882,1009,1806,1831]" italics="true" pageId="77" pageNumber="78">M. dicrana</emphasis>
|
||
<bibRefCitation id="EFB84B53FFE78938FC6EF8BB77D59E4B" author="Clark, H. L." box="[1019,1217,1806,1831]" pageId="77" pageNumber="78" pagination="81 - 283" refId="ref34331" refString="Clark, H. L. (1917) Hawaiian and other Pacific Echini, Echinoneidae, Nucleolitidae, Urechinidae, Echinocorythidae, Calymnidae, Pourtalesiidae, Palaestomatidae, Aeropsidae, Palaeopneustidae, Hemiasteridae, and Spatangidae. Memoirs of the Museum of Comparative Zoology at Harvard College, 46, 81 - 283." type="journal article" year="1917">H.L. Clark, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FB5AF8BB72999E27" authority="Baker & Rowe, 1990" authorityName="Baker & Rowe" authorityYear="1990" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="kermadecensis">
|
||
<emphasis id="B95DEAB0FFE78938FB5AF8BB76889E4B" box="[1231,1436,1806,1831]" italics="true" pageId="77" pageNumber="78">M. kermadecensis</emphasis>
|
||
Baker & Rowe, 1990
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FE0CF88071E69E27" authority="H. L. Clark, 1925" authorityName="H. L. Clark" authorityYear="1925" box="[409,754,1842,1867]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="latissima">
|
||
<emphasis id="B95DEAB0FFE78938FE0CF88071319E27" box="[409,549,1842,1867]" italics="true" pageId="77" pageNumber="78">M. latissima</emphasis>
|
||
H.L. Clark, 1925
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FD6AF88077199E27" authority="Verrill, 1867" authorityName="Verrill" authorityYear="1867" box="[767,1037,1842,1867]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="nobilis">
|
||
<emphasis id="B95DEAB0FFE78938FD6AF88070669E27" box="[767,882,1842,1867]" italics="true" pageId="77" pageNumber="78">M. nobilis</emphasis>
|
||
Verrill, 1867
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FB8FF88076649E27" authority="(Mortensen, 1940)" baseAuthorityName="Mortensen" baseAuthorityYear="1940" box="[1050,1392,1843,1867]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="persica">
|
||
<emphasis id="B95DEAB0FFE78938FB8FF88077879E27" box="[1050,1171,1844,1867]" italics="true" pageId="77" pageNumber="78">M. persica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB84B53FFE78938FB36F887767D9E27" author="Mortensen, T." box="[1187,1385,1843,1867]" pageId="77" pageNumber="78" pagination="55 - 112" refId="ref35321" refString="Mortensen, T. (1940) Echinoderms from the Iranian Gulf. Asteroidea, Ophiuroidea, and Echinoidea. Danish Scientific Investigations in Iran, Part 2, 55 - 112." type="book chapter" year="1940">Mortensen, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FAE8F88072D89E03" authority="(de Loriol, 1876)" baseAuthorityName="de Loriol" baseAuthorityYear="1876" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="robillardi">
|
||
<emphasis id="B95DEAB0FFE78938FAE8F88072169E03" italics="true" pageId="77" pageNumber="78">M. robillardi</emphasis>
|
||
(de Loriol, 1876)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FE4DF8E3703E9E03" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[472,810,1878,1903]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="spatagus">
|
||
<emphasis id="B95DEAB0FFE78938FE4DF8E371769E02" box="[472,610,1879,1902]" italics="true" pageId="77" pageNumber="78">M. spatagus</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FCA3F8E377979E03" authority="(Lamarck, 1816)" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[822,1155,1878,1903]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="sternalis">
|
||
<emphasis id="B95DEAB0FFE78938FCA3F8E370A89E03" box="[822,956,1878,1903]" italics="true" pageId="77" pageNumber="78">M. sternalis</emphasis>
|
||
(Lamarck, 1816)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C294D21FFE78938FB2BF8E373CF9EFF" authority="(Bell, 1904)" baseAuthorityName="Bell" baseAuthorityYear="1904" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="townsendi">
|
||
<emphasis id="B95DEAB0FFE78938FB2BF8E376419E03" box="[1214,1365,1878,1903]" italics="true" pageId="77" pageNumber="78">M. townsendi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB84B53FFE78938FAF0F8E273C79EFF" author="Herdman, W. A. & Herdman, J. B. & Bell, F. J." pageId="77" pageNumber="78" pagination="137 - 150" refId="ref34833" refString="Herdman, W. A., Herdman, J. B. & Bell, F. J. (1904) Report on the Echinoderma collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the government of Ceylon on the pearl oyster fisheries of the Gulf of Mannar. The Royal Society, London, pp. 137 - 150." type="book chapter" year="1904">Bell, 1904</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFE7893BFF52F82A73FB999B" blockId="77.[151,1437,726,2011]" lastBlockId="78.[151,1436,151,248]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
|
||
The new species lacks a notch in ambulacrum III, a condition that strongly differs from that in the following species:
|
||
<taxonomicName id="4C294D21FFE78938FF6EF870729C9EB7" authorityName="de Ridder" authorityYear="1984" box="[251,392,1988,2011]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="angustus">
|
||
<emphasis id="B95DEAB0FFE78938FF6EF870729C9EB7" box="[251,392,1988,2011]" italics="true" pageId="77" pageNumber="78">M. angustus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FE0FF870717C9EB7" authorityName="Baker & Rowe" authorityYear="1990" box="[410,616,1986,2011]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="kermadecensis">
|
||
<emphasis id="B95DEAB0FFE78938FE0FF870717C9EB7" box="[410,616,1986,2011]" italics="true" pageId="77" pageNumber="78">M. kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FDEFF870701D9EB7" authorityName="H. L. Clark" authorityYear="1925" box="[634,777,1986,2011]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="latissima">
|
||
<emphasis id="B95DEAB0FFE78938FDEFF870701D9EB7" box="[634,777,1986,2011]" italics="true" pageId="77" pageNumber="78">M. latissima</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFE78938FC8EF87070839EB7" baseAuthorityName="Mortensen" baseAuthorityYear="1940" box="[795,919,1988,2011]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="persica">
|
||
<emphasis id="B95DEAB0FFE78938FC8EF87070839EB7" box="[795,919,1988,2011]" italics="true" pageId="77" pageNumber="78">M. persica</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C294D21FFE78938FC48F87077659EB7" baseAuthorityName="de Loriol" baseAuthorityYear="1876" box="[989,1137,1986,2011]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="77" pageNumber="78" phylum="Echinodermata" rank="species" species="robillardi">
|
||
<emphasis id="B95DEAB0FFE78938FC48F87077659EB7" box="[989,1137,1986,2011]" italics="true" pageId="77" pageNumber="78">M. robillardi</emphasis>
|
||
</taxonomicName>
|
||
. The lack of coalescence between the aboralmost regions of the posterior paired petals is very different from the situation seen in
|
||
<emphasis id="B95DEAB0FFE4893BFAE8FF2D728899B9" italics="true" pageId="78" pageNumber="79">
|
||
<taxonomicName id="4C294D21FFE4893BFAE8FF2D73EF99B9" baseAuthorityName="Lamarck" baseAuthorityYear="1816" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="78" pageNumber="79" phylum="Echinodermata" rank="species" species="sternalis">M. sternalis</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C294D21FFE4893BFE90FF09728899B9" baseAuthorityName="Bell" baseAuthorityYear="1904" box="[261,412,188,213]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="78" pageNumber="79" phylum="Echinodermata" rank="species" species="townsendi">M. townsendi</taxonomicName>
|
||
</emphasis>
|
||
differs in having a subcentral position of the apical system, and much lower anterior end of the test.
|
||
</paragraph>
|
||
<caption id="DF56662AFFE4893BFF02F83376639EF1" box="[151,1399,1927,1949]" httpUri="https://zenodo.org/record/2617591/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,250,1927,1949]" targetBox="[151,1436,301,1906]" targetPageId="78">
|
||
<paragraph id="8B9636A2FFE4893BFF02F83376639EF1" blockId="78.[151,1399,1927,1949]" box="[151,1399,1927,1949]" pageId="78" pageNumber="79">
|
||
<emphasis id="B95DEAB0FFE4893BFF02F83372349EF0" bold="true" box="[151,288,1927,1949]" pageId="78" pageNumber="79">FIGURE 68.</emphasis>
|
||
<taxonomicName id="4C294D21FFE4893BFEB0F83372609EF1" authorityName="Gray" authorityYear="1855" box="[293,372,1927,1949]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="78" pageNumber="79" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFE4893BFEB0F83372609EF1" box="[293,372,1927,1949]" italics="true" pageId="78" pageNumber="79">Metalia</emphasis>
|
||
</taxonomicName>
|
||
sp. (WUSL/EI/85): A, aboral view; B, oral view; C, lateral view; D, anterior view; E, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF56662AFFE5893AFF02F8A871DF9E5E" box="[151,715,1820,1842]" httpUri="https://zenodo.org/record/2617593/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,250,1820,1842]" targetBox="[151,1436,193,1798]" targetPageId="79">
|
||
<paragraph id="8B9636A2FFE5893AFF02F8A871DF9E5E" blockId="79.[151,715,1820,1842]" box="[151,715,1820,1842]" pageId="79" pageNumber="80">
|
||
<emphasis id="B95DEAB0FFE5893AFF02F8A872349E5D" bold="true" box="[151,288,1820,1842]" pageId="79" pageNumber="80">FIGURE 69.</emphasis>
|
||
<taxonomicName id="4C294D21FFE5893AFEB0F8A872609E5E" authorityName="Gray" authorityYear="1855" box="[293,372,1820,1842]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFE5893AFEB0F8A872609E5E" box="[293,372,1820,1842]" italics="true" pageId="79" pageNumber="80">Metalia</emphasis>
|
||
</taxonomicName>
|
||
sp. (WUSL/EI/138), aboral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF56662AFFFA8925FF02F8A870F39E5E" box="[151,999,1820,1842]" httpUri="https://zenodo.org/record/2617595/files/figure.png" pageId="80" pageNumber="81" startId="80.[151,250,1820,1842]" targetBox="[151,1436,193,1798]" targetPageId="80">
|
||
<paragraph id="8B9636A2FFFA8925FF02F8A870F39E5E" blockId="80.[151,999,1820,1842]" box="[151,999,1820,1842]" pageId="80" pageNumber="81">
|
||
<emphasis id="B95DEAB0FFFA8925FF02F8A8720B9E5D" bold="true" box="[151,287,1820,1842]" pageId="80" pageNumber="81">FIGURE 70.</emphasis>
|
||
<taxonomicName id="4C294D21FFFA8925FEB0F8A872609E5E" authorityName="Gray" authorityYear="1855" box="[293,372,1820,1842]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFFA8925FEB0F8A872609E5E" box="[293,372,1820,1842]" italics="true" pageId="80" pageNumber="81">Metalia</emphasis>
|
||
</taxonomicName>
|
||
sp. (WUSL/EI/85): A, sternal view; B, subanal fasciole view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF56662AFFFB8924FF02F8E170589E07" box="[151,844,1877,1899]" httpUri="https://zenodo.org/record/2617597/files/figure.png" pageId="81" pageNumber="82" startId="81.[151,250,1877,1899]" targetBox="[182,1426,219,1839]" targetPageId="81">
|
||
<paragraph id="8B9636A2FFFB8924FF02F8E170589E07" blockId="81.[151,844,1877,1899]" box="[151,844,1877,1899]" pageId="81" pageNumber="82">
|
||
<emphasis id="B95DEAB0FFFB8924FF02F8E1720B9E06" bold="true" box="[151,287,1877,1899]" pageId="81" pageNumber="82">FIGURE 71.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="4C294D21FFFB8924FE61F8E171579E07" authorityName="Gray" authorityYear="1855" box="[500,579,1877,1899]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="81" pageNumber="82" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFFB8924FE61F8E171579E07" box="[500,579,1877,1899]" italics="true" pageId="81" pageNumber="82">Metalia</emphasis>
|
||
</taxonomicName>
|
||
sp. recorded in this study.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9636A2FFFB8924FF52F82D70A69EB8" blockId="81.[151,1436,1944,2005]" pageId="81" pageNumber="82">
|
||
Forms that lack the anterior notch, lack the coalescence, and have the apical system at least somewhat anterior, as in the new species, include
|
||
<taxonomicName id="4C294D21FFFB8924FE7CF80971719EB9" authorityName="H. L. Clark" authorityYear="1917" box="[489,613,1980,2005]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="81" pageNumber="82" phylum="Echinodermata" rank="species" species="dicrana">
|
||
<emphasis id="B95DEAB0FFFB8924FE7CF80971719EB9" box="[489,613,1980,2005]" italics="true" pageId="81" pageNumber="82">M. dicrana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C294D21FFFB8924FDE1F80971F19EB9" authorityName="Verrill" authorityYear="1867" box="[628,741,1980,2005]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="81" pageNumber="82" phylum="Echinodermata" rank="species" species="nobilis">
|
||
<emphasis id="B95DEAB0FFFB8924FDE1F80971F19EB9" box="[628,741,1980,2005]" italics="true" pageId="81" pageNumber="82">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C294D21FFFB8924FCB7F80970B89EB8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[802,940,1981,2004]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="81" pageNumber="82" phylum="Echinodermata" rank="species" species="spatagus">
|
||
<emphasis id="B95DEAB0FFFB8924FCB7F80970B89EB8" box="[802,940,1981,2004]" italics="true" pageId="81" pageNumber="82">M. spatagus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFF88927FF52FF2D717A9808" blockId="82.[151,1436,151,645]" pageId="82" pageNumber="83">
|
||
<taxonomicName id="4C294D21FFF88927FF52FF2D722D99DC" authorityName="Verrill" authorityYear="1867" box="[199,313,151,176]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="species" species="nobilis">
|
||
<emphasis id="B95DEAB0FFF88927FF52FF2D722D99DC" box="[199,313,151,176]" italics="true" pageId="82" pageNumber="83">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
, the only
|
||
<taxonomicName id="4C294D21FFF88927FE39FF23711799DC" authorityName="Gray" authorityYear="1855" box="[428,515,151,176]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFF88927FE39FF23711799DC" box="[428,515,151,176]" italics="true" pageId="82" pageNumber="83">Metalia</emphasis>
|
||
</taxonomicName>
|
||
known from the Americas, has small petals relative to its congeners. In addition, the periproct in
|
||
<taxonomicName id="4C294D21FFF88927FEDEFF0872B699B9" authorityName="Gray" authorityYear="1855" box="[331,418,188,213]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFF88927FEDEFF0872B699B9" box="[331,418,188,213]" italics="true" pageId="82" pageNumber="83">Metalia</emphasis>
|
||
</taxonomicName>
|
||
n. sp. is directed downwards and is not fully visible from the aboral side. This is not only unlike
|
||
<taxonomicName id="4C294D21FFF88927FF70FF55724C9994" authorityName="Verrill" authorityYear="1867" box="[229,344,223,248]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="species" species="nobilis">
|
||
<emphasis id="B95DEAB0FFF88927FF70FF55724C9994" box="[229,344,223,248]" italics="true" pageId="82" pageNumber="83">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
, it is not like
|
||
<taxonomicName id="4C294D21FFF88927FE62FF5571969994" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[503,642,225,248]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="species" species="spatagus">
|
||
<emphasis id="B95DEAB0FFF88927FE62FF5571969994" box="[503,642,225,248]" italics="true" pageId="82" pageNumber="83">M. spatagus</emphasis>
|
||
</taxonomicName>
|
||
either, which has a more vertical posterior region with at least part of the periproct visible in aboral view. This is also true for
|
||
<taxonomicName id="4C294D21FFF88927FC87FEB1709B9871" authorityName="H. L. Clark" authorityYear="1917" box="[786,911,260,285]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="species" species="dicrana">
|
||
<emphasis id="B95DEAB0FFF88927FC87FEB1709B9871" box="[786,911,260,285]" italics="true" pageId="82" pageNumber="83">M. dicrana</emphasis>
|
||
</taxonomicName>
|
||
, which is distinguished from all other
|
||
<taxonomicName id="4C294D21FFF88927FAD0FEB076889871" authorityName="Gray" authorityYear="1855" box="[1349,1436,260,285]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFF88927FAD0FEB076889871" box="[1349,1436,260,285]" italics="true" pageId="82" pageNumber="83">Metalia</emphasis>
|
||
</taxonomicName>
|
||
by its peculiar double point along the sternum just adoral to the subanal fasciole (Mooi, unpubl. data). Therefore, the specimens are very unlike
|
||
<taxonomicName id="4C294D21FFF88927FE7EFEF971739809" authorityName="H. L. Clark" authorityYear="1917" box="[491,615,332,357]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="species" species="dicrana">
|
||
<emphasis id="B95DEAB0FFF88927FE7EFEF971739809" box="[491,615,332,357]" italics="true" pageId="82" pageNumber="83">M. dicrana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFF88927FF52FEDB77E09B74" blockId="82.[151,1436,151,645]" pageId="82" pageNumber="83">
|
||
The Sri Lankan specimens of this new form have a very distinctive, concave region below the periproct which has not been noted in other
|
||
<taxonomicName id="4C294D21FFF88927FE58FE20713098C1" authorityName="Gray" authorityYear="1855" box="[461,548,404,429]" class="Echinoidea" family="Brissidae" genus="Metalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFF88927FE58FE20713098C1" box="[461,548,404,429]" italics="true" pageId="82" pageNumber="83">Metalia</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<figureCitation id="13122A27FFF88927FD19FE2071CB98C0" box="[652,735,404,429]" captionStart="FIGURE 70" captionStartId="80.[151,250,1820,1842]" captionTargetBox="[151,1436,193,1798]" captionTargetId="figure@80.[151,1436,193,1798]" captionTargetPageId="80" captionText="FIGURE 70. Metalia sp. (WUSL/EI/85): A, sternal view; B, subanal fasciole view." httpUri="https://zenodo.org/record/2617595/files/figure.png" pageId="82" pageNumber="83">Fig. 70</figureCitation>
|
||
), although it does occur to some degree in some of the species (Mooi, unpubl. data). As discovered by
|
||
<bibRefCitation id="EFB84B53FFF88927FDC1FE03705D98BC" author="Askin, R. & Mooi, R." box="[596,841,439,464]" pageId="82" pageNumber="83" pagination="7" refId="ref33641" refString="Askin, R. & Mooi, R. (2017) The heart of the problem: Phylogenetic systematics of burrowing sea urchins in the genus Metalia (Echinoidea: Spatangoida). Integrative and Comparative Biology, 57 (Supplement 1), e 7." type="journal article" year="2017">Askin & Mooi (2017)</bibRefCitation>
|
||
using molecular techniques, it would appear that this is a new species. It was depicted by van
|
||
<bibRefCitation id="EFB84B53FFF88927FDFCFE6870479898" author="van Noordenburg, H." box="[617,851,476,501]" pageId="82" pageNumber="83" refId="ref36274" refString="van Noordenburg, H. (2008) Sea Urchins of the Philippines. Part I. The Irregulars. Artificial Harmonics, Utrecht, 153 pp." type="book" year="2008">Noordenburg (2008)</bibRefCitation>
|
||
, but remains unnamed pending further work on the abundant material at the
|
||
<collectingRegion id="49EDF840FFF88927FE39FE4B710A9B74" box="[428,542,511,536]" country="United States of America" name="California" pageId="82" pageNumber="83">California</collectingRegion>
|
||
<materialsCitation id="3B413CFFFFF88927FDB1FE4B77FB9B74" ID-GBIF-Occurrence="2147524161" box="[548,1263,511,536]" location="Academy of Sciences" pageId="82" pageNumber="83" specimenCount="1" stateProvince="Wien">
|
||
<location id="8EF66079FFF88927FDB1FE4B70039B74" LSID="urn:lsid:plazi:treatment:038087B4FFE78927FF02FDD872369BE8:8EF66079FFF88927FDB1FE4B70039B74" box="[548,791,511,536]" name="Academy of Sciences" pageId="82" pageNumber="83" stateProvince="Wien">Academy of Sciences</location>
|
||
and the Natural History Museum,
|
||
<collectingRegion id="49EDF840FFF88927FB09FE4B77FB9B74" box="[1180,1263,511,536]" country="Austria" name="Wien" pageId="82" pageNumber="83">Vienna</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFF88927FF52FD9072369BE8" blockId="82.[151,1436,151,645]" pageId="82" pageNumber="83">
|
||
Variation in the path of the peripetalous fasciole was noted in collected specimens. WUSL/IE/E85 shows shallow aborally-curved segments in interambulacrum 4, and one sharply defined indentation in interambulacrum 1 (
|
||
<figureCitation id="13122A27FFF88927FF0AFDD873E69BE9" box="[159,242,620,645]" captionStart="FIGURE 68" captionStartId="78.[151,250,1927,1949]" captionTargetBox="[151,1436,301,1906]" captionTargetId="figure@78.[151,1436,301,1906]" captionTargetPageId="78" captionText="FIGURE 68. Metalia sp. (WUSL/EI/85): A, aboral view; B, oral view; C, lateral view; D, anterior view; E, posterior view." httpUri="https://zenodo.org/record/2617591/files/figure.png" pageId="82" pageNumber="83">Fig. 68</figureCitation>
|
||
, A).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |