743 lines
123 KiB
XML
743 lines
123 KiB
XML
<document id="FD4443B468B0BB250808EFABEADB7E53" ID-CLB-Dataset="29911" ID-DOI="10.11646/zootaxa.4433.3.11" ID-GBIF-Dataset="ce0ab832-46b8-40e9-8aaa-9ca99ab24f32" ID-ISSN="1175-5326" ID-Zenodo-Dep="1290343" ID-ZooBank="A2E322CC-B7CC-4715-8434-B29EECD8F6DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529055656449" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2018" docId="F67D87C5FFCDFFD9FF6CFC53C07BFDD4" docLanguage="en" docName="zootaxa.4433.3.11.pdf" docOrigin="Zootaxa 4433 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus naranjo Anker 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="581" masterDocId="0A44FFBDFFCCFFDEFFFBFFCFC45AFFE9" masterDocTitle="Alpheus naranjo, a new brightly coloured snapping shrimp from the Caribbean coast of Panama (Malacostraca, Decapoda, Alpheidae)" masterLastPageNumber="582" masterPageNumber="574" pageNumber="575" updateTime="1698548336561" updateUser="ExternalLinkService">
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<mods:title id="42AAB9DF80316C91E8FD2847D9043FDF">Alpheus naranjo, a new brightly coloured snapping shrimp from the Caribbean coast of Panama (Malacostraca, Decapoda, Alpheidae)</mods:title>
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<mods:name id="CDD76D4F7873BCA58A1A225EA3779A70" type="personal">
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<mods:roleTerm id="5E393FE6AC7F6ACCA2CBB0CFF07078A2">Author</mods:roleTerm>
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<mods:namePart id="BFF28728ECF1154BA4B319D6692F6A4D">Anker, Arthur</mods:namePart>
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<mods:title id="44823765FDFB02CA4FB64833F04BFE3D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1C2B5AC950668E743FA9CA658D31FF51">
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<mods:date id="372ECB3F9A47F4EF14C752F4A8B20AB0">2018</mods:date>
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<mods:number id="2FC7C4BCAE3E1A5FD703FB4103426DEE">2018-06-13</mods:number>
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<mods:number id="4BE74C9D2E08D586F77485B2FE4C5EA2">4433</mods:number>
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<mods:identifier id="FC36691F91AA828EBC6D172C3D2DFE2D" type="DOI">10.11646/zootaxa.4433.3.11</mods:identifier>
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<mods:identifier id="EAFB072D2071CE37EC7818967B61CB89" type="GBIF-Dataset">ce0ab832-46b8-40e9-8aaa-9ca99ab24f32</mods:identifier>
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<mods:identifier id="748B1E5287484DFB38A3CE183168DD6B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="F67D87C5FFCDFFD9FF6CFC53C07BFDD4" ID-DOI="http://doi.org/10.5281/zenodo.5958274" ID-GBIF-Taxon="145952767" ID-Zenodo-Dep="5958274" LSID="urn:lsid:plazi:treatment:F67D87C5FFCDFFD9FF6CFC53C07BFDD4" httpUri="http://treatment.plazi.org/id/F67D87C5FFCDFFD9FF6CFC53C07BFDD4" lastPageId="7" lastPageNumber="581" pageId="1" pageNumber="575">
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<subSubSection id="36CE6558FFCDFFDFFF6CFC53C518FC31" pageId="1" pageNumber="575" type="nomenclature">
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<paragraph id="7E6B36D3FFCDFFDFFF6CFC53C59CFC5E" blockId="1.[151,454,924,984]" box="[151,454,924,951]" pageId="1" pageNumber="575">
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<heading id="252381BFFFCDFFDFFF6CFC53C59CFC5E" bold="true" box="[151,454,924,951]" fontSize="11" level="1" pageId="1" pageNumber="575" reason="1">
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<emphasis id="4CA0EAC1FFCDFFDFFF6CFC53C59CFC5E" bold="true" box="[151,454,924,951]" pageId="1" pageNumber="575">
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||
<taxonomicName id="B9D44D50FFCDFFDFFF6CFC53C538FC5F" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[151,354,924,950]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="575" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCDFFDFFF6CFC53C538FC5F" bold="true" box="[151,354,924,950]" italics="true" pageId="1" pageNumber="575">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCDFFDFFE92FC52C59CFC5E" box="[361,454,925,951]" pageId="1" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCDFFDFFF6CFC70C518FC31" blockId="1.[151,454,924,984]" box="[151,322,959,984]" pageId="1" pageNumber="575">
|
||
(
|
||
<figureCitation id="E6EF2A56FFCDFFDFFF64FC70C4B6FC31" box="[159,236,959,984]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="E6EF2A56FFCDFFDFFF01FC0FC550FC31" box="[250,266,960,984]" captionStart="FIGURE 2" captionStartId="5.[151,250,1910,1932]" captionTargetBox="[184,1388,351,1872]" captionTargetId="figure@5.[181,1413,336,1897]" captionTargetPageId="5" captionText="FIGURE 2. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—major cheliped, chela, lateral view; B—same, mesial view; C—same, coxa to carpus, lateral view; D—same, ischium, merus and carpus, mesial view, conventional setae omitted; E—same, dactylus, lateral view; F—minor cheliped, lateral view; G—same, chela, mesial view; H—same, ischium, merus and carpus, mesial view, conventional setae omitted." httpUri="https://zenodo.org/record/1290347/files/figure.png" pageId="1" pageNumber="575">2</figureCitation>
|
||
,
|
||
<figureCitation id="E6EF2A56FFCDFFDFFEEDFC0FC560FC31" box="[278,314,960,984]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="1" pageNumber="575">3A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCDFFDFFF6CFBC8C152FB11" pageId="1" pageNumber="575" type="materials_examined">
|
||
<paragraph id="7E6B36D3FFCDFFDFFF6CFBC8C152FB11" blockId="1.[151,1437,1031,2029]" pageId="1" pageNumber="575">
|
||
<materialsCitation id="CEBC3C8EFFCDFFDFFF6CFBC8C08DFBF6" ID-GBIF-Occurrence="1849037850" box="[151,1239,1031,1056]" collectionCode="FLMNH" country="Panama" location="Caribbean" pageId="1" pageNumber="575" specimenCode="UF 44471" specimenCount="2" stateProvince="Panama" typeStatus="holotype">
|
||
<emphasis id="4CA0EAC1FFCDFFDFFF6CFBC8C519FBC9" bold="true" box="[151,323,1031,1056]" pageId="1" pageNumber="575">
|
||
<typeStatus id="A16F8871FFCDFFDFFF6CFBC8C488FBC9" box="[151,210,1031,1056]" pageId="1" pageNumber="575">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="A16F8871FFCDFFDFFEADFBC8C599FBC9" box="[342,451,1031,1056]" pageId="1" pageNumber="575" type="holotype">Holotype</typeStatus>
|
||
: male (cl 9.0 mm),
|
||
<collectionCode id="18C5AE16FFCDFFDFFD4CFBC7C743FBF6" box="[695,793,1032,1055]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="575">FLMNH</collectionCode>
|
||
<specimenCode id="2E729EA8FFCDFFDFFCDDFBC7C7C5FBC9" box="[806,927,1032,1056]" collectionCode="FLMNH" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="575">UF 44471</specimenCode>
|
||
,
|
||
<collectingRegion id="BC10F831FFCDFFDFFC4BFBC7C056FBF6" box="[944,1036,1032,1055]" country="Panama" name="Panama" pageId="1" pageNumber="575">
|
||
<collectingCountry id="06C37643FFCDFFDFFC4BFBC7C056FBF6" box="[944,1036,1032,1055]" name="Panama" pageId="1" pageNumber="575">Panama</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<location id="7B0B6008FFCDFFDFFBE7FBC8C0D5FBC9" LSID="urn:lsid:plazi:treatment:F67D87C5FFCDFFD9FF6CFC53C07BFDD4:7B0B6008FFCDFFDFFBE7FBC8C0D5FBC9" box="[1052,1167,1031,1056]" country="Panama" name="Caribbean" pageId="1" pageNumber="575" stateProvince="Panama">Caribbean</location>
|
||
coast
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="CEBC3C8EFFCDFFDFFB13FBC7C0FDFB64" ID-GBIF-Occurrence="1849038038" collectingDate="2016-05-24" collectionCode="OUMNH" collectorName="Bocas & Toro Archipelago & Isla Popa & Popa Reef & M. Leray & F. Michonneau & R. Lasley" country="Iceland" location="Caribbean" pageId="1" pageNumber="575" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="paratype">
|
||
<collectingRegion id="BC10F831FFCDFFDFFB13FBC7C1C6FBF6" box="[1256,1436,1031,1056]" country="Panama" name="Bocas del Toro" pageId="1" pageNumber="575">
|
||
<collectorName id="D3215305FFCDFFDFFB13FBC7C177FBF6" box="[1256,1325,1032,1055]" pageId="1" pageNumber="575">Bocas</collectorName>
|
||
del
|
||
<collectorName id="D3215305FFCDFFDFFA93FBC7C1C6FBF6" box="[1384,1436,1032,1055]" pageId="1" pageNumber="575">Toro</collectorName>
|
||
</collectingRegion>
|
||
Archipelago, off
|
||
<collectorName id="D3215305FFCDFFDFFEACFBE3C598FBAD" box="[343,450,1068,1093]" pageId="1" pageNumber="575">
|
||
<collectingCountry id="06C37643FFCDFFDFFEACFBE3C5DAFBAC" box="[343,384,1068,1093]" name="Iceland" pageId="1" pageNumber="575">Isla</collectingCountry>
|
||
Popa
|
||
</collectorName>
|
||
,
|
||
<collectorName id="D3215305FFCDFFDFFE35FBE2C61CFBAC" box="[462,582,1068,1093]" pageId="1" pageNumber="575">Popa Reef</collectorName>
|
||
, 9.233° -82.112°, coll.
|
||
<collectorName id="D3215305FFCDFFDFFCB0FBE2C7EEFBAC" box="[843,948,1069,1093]" pageId="1" pageNumber="575">M. Leray</collectorName>
|
||
,
|
||
<collectorName id="D3215305FFCDFFDFFC3BFBE2C032FBAC" box="[960,1128,1068,1093]" pageId="1" pageNumber="575">F. Michonneau</collectorName>
|
||
&
|
||
<collectorName id="D3215305FFCDFFDFFB76FBE2C0A0FBAC" box="[1165,1274,1068,1093]" pageId="1" pageNumber="575">R. Lasley</collectorName>
|
||
,
|
||
<collectingDate id="1A2EE9FBFFCDFFDFFAFDFBE3C1C6FBAC" box="[1286,1436,1068,1093]" pageId="1" pageNumber="575" value="2016-05-24">
|
||
<date id="0A6A1013FFCDFFDFFAFDFBE3C1C6FBAC" box="[1286,1436,1068,1093]" pageId="1" pageNumber="575" value="2016-05-24">24 May 2016</date>
|
||
</collectingDate>
|
||
(fcn BBDT-2166).
|
||
<typeStatus id="A16F8871FFCDFFDFFE95FB9FC5BAFB81" box="[366,480,1104,1128]" pageId="1" pageNumber="575" type="paratype">Paratypes</typeStatus>
|
||
: 1 ovigerous female (cl
|
||
<quantity id="B92C9B36FFCDFFDFFD0FFB9FC715FB81" box="[756,847,1104,1128]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="575" unit="mm" value="9.7">9.7 mm</quantity>
|
||
),
|
||
<collectionCode id="18C5AE16FFCDFFDFFC98FB9FC789FB81" box="[867,979,1104,1128]" pageId="1" pageNumber="575">OUMNH</collectionCode>
|
||
.ZC. 2018.01.0 3, same collection data as for holotype (fcn BBDT-2166); 1 ovigerous female (cl 10.0 mm, missing major cheliped)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="CEBC3C8EFFCDFFDFFB4EFBBAC0BEFB59" ID-GBIF-Occurrence="1849038039" collectionCode="FLMNH" country="Panama" location="Caribbean" pageId="1" pageNumber="575" specimenCode="UF 44448" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
|
||
<collectionCode id="18C5AE16FFCDFFDFFB4EFBBAC14DFB65" box="[1205,1303,1141,1164]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="575">FLMNH</collectionCode>
|
||
<specimenCode id="2E729EA8FFCDFFDFFADAFBBAC1C2FB65" box="[1313,1432,1140,1164]" collectionCode="FLMNH" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="575">UF 44448</specimenCode>
|
||
, same collection data as for holotype (fcn BBDT-2112);
|
||
<specimenCount id="68D2FD5AFFCDFFDFFCFAFB57C711FB59" box="[769,843,1175,1200]" pageId="1" pageNumber="575" type="male">1 male</specimenCount>
|
||
(8.4 mm—dissected and illustrated)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="CEBC3C8EFFCDFFDFFB15FB57C486FB11" ID-GBIF-Occurrence="1849037849" collectionCode="FLMNH" country="Panama" location="Caribbean" pageId="1" pageNumber="575" specimenCount="1" specimenCount-female="1" stateProvince="Panama" typeStatus="paratype">
|
||
<specimenCount id="68D2FD5AFFCDFFDFFB15FB57C115FB59" box="[1262,1359,1175,1200]" pageId="1" pageNumber="575" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="B92C9B36FFCDFFDFFA82FB57C49DFB3D" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="575" unit="mm" value="8.1">8.1 mm</quantity>
|
||
), 1 ovigerous female (
|
||
<quantity id="B92C9B36FFCDFFDFFE35FB73C671FB3D" box="[462,555,1212,1236]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="575" unit="mm" value="8.7">8.7 mm</quantity>
|
||
, missing minor cheliped), same collection data as for holotype (fcn BBDT- 2744)
|
||
</materialsCitation>
|
||
; 1 male (
|
||
<quantity id="B92C9B36FFCDFFDFFEBAFB10C5C7FB11" box="[321,413,1247,1272]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="575" unit="mm" value="8.6">8.6 mm</quantity>
|
||
), MNHN-IU-2017-1547, same collection data for holotype (fcn BBDT-2744).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCDFFDDFF3CFACBC066F964" lastPageId="3" lastPageNumber="577" pageId="1" pageNumber="575" type="description">
|
||
<paragraph id="7E6B36D3FFCDFFDFFF3CFACBC6D6F9FE" blockId="1.[151,1437,1031,2029]" pageId="1" pageNumber="575">
|
||
<emphasis id="4CA0EAC1FFCDFFDFFF3CFACBC50BFAF4" bold="true" box="[199,337,1284,1309]" pageId="1" pageNumber="575">Description</emphasis>
|
||
. Medium-sized species of
|
||
<taxonomicName id="B9D44D50FFCDFFDFFD7AFACBC681FAF5" authorityName="Anker" authorityYear="2018" box="[641,731,1284,1308]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="575" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CA0EAC1FFCDFFDFFD7AFACBC681FAF5" box="[641,731,1284,1308]" italics="true" pageId="1" pageNumber="575">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
(cl reaching 10.0 mm). Carapace smooth, not setose. Rostrum well developed, long, relatively slender, more than twice as long as broad at base, sharp distally, reaching or almost reaching distal margin of first article of antennular peduncle, flattening posteriorly and with base abruptly delimited from adjacent rostro-orbital area; rostral carina not distinct; lateral margin furnished with stiff setae (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFA94FAA0C485FA45" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1A, B</figureCitation>
|
||
). Orbital hoods moderately swollen; anterior margin of each orbital hood armed with anteriorly directed, sharp tooth, latter reaching or almost reaching mid-length of rostrum; margin between tooth and rostrum somewhat flattened (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFEF3FA13C5DBFA1D" box="[264,385,1500,1525]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1A, B</figureCitation>
|
||
). Pterygostomial angle rounded, not protruding (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFC58FA13C7A3FA1D" box="[931,1017,1500,1525]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1B</figureCitation>
|
||
); cardiac notch well developed. Each epistomial sclerite with strong blunt process.
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCDFFDFFF3CF9EBC684F925" blockId="1.[151,1437,1031,2029]" pageId="1" pageNumber="575">
|
||
Telson moderately broad, subrectangular, tapering distally, about twice as long as proximal width, with slightly convex lateral margins; dorsal surface with two pairs of very stout spiniform setae both inserted at some distance from lateral margins, first pair at about telson mid-length, second pair at about 0.7 of telson length; posterior margin broadly rounded, without spiniform setae; each posterolateral angle with pair of stout spiniform setae, lateral about one-third of length of mesial (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFD82F97BC68AF925" box="[633,720,1716,1741]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCDFFDFFF3CF918C57DF805" blockId="1.[151,1437,1031,2029]" pageId="1" pageNumber="575">
|
||
Antennular peduncle with second article short, about 1.2 times as long as wide; stylocerite with sharp point reaching to or beyond mid-length of first article; ventromesial carina with large, broadly triangular tooth, latter with small but well-marked anterior point; lateral antennular flagellum with secondary ramus fused to main ramus over most of its length, except for very short free terminal section, with numerous groups of aesthetascs starting from about fourth article (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFE33F8A8C62AF869" box="[456,624,1895,1920]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1A, B, D</figureCitation>
|
||
). Antenna with stout basicerite bearing large sharp distoventral tooth; scaphocerite relatively stout, with straight lateral margin and well-developed blade, latter not overreaching strong distolateral tooth; carpocerite reaching slightly beyond scaphocerite and exceeding end of antennular peduncle (
|
||
<figureCitation id="E6EF2A56FFCDFFDFFF64F81BC543F805" box="[159,281,2004,2029]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="1" pageNumber="575">Fig. 1A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2AAB665BFFCEFFDCFF6CF8FDC5D5F80B" httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="2" pageNumber="576" startId="2.[151,250,1842,1864]" targetBox="[193,1407,204,1798]" targetPageId="2">
|
||
<paragraph id="7E6B36D3FFCEFFDCFF6CF8FDC5D5F80B" blockId="2.[151,1436,1842,2018]" pageId="2" pageNumber="576">
|
||
<emphasis id="4CA0EAC1FFCEFFDCFF6CF8FDC557F8AE" bold="true" box="[151,269,1842,1864]" pageId="2" pageNumber="576">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="B9D44D50FFCEFFDCFEE2F8FDC59BF8A1" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[281,449,1842,1864]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="576" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCEFFDCFEE2F8FDC59BF8A1" box="[281,449,1842,1864]" italics="true" pageId="2" pageNumber="576">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CA0EAC1FFCEFFDCFE33F8FCC645F8A1" bold="true" box="[456,543,1843,1864]" pageId="2" pageNumber="576">
|
||
<taxonomicNameLabel id="579357BAFFCEFFDCFE33F8FCC641F8A1" box="[456,539,1843,1864]" pageId="2" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CFF58C7A0FEF2" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
Mouthparts typical for genus in external observation. Third maxilliped slender, pediform; coxa with distally subacute lateral plate furnished with some long setae; antepenultimate article slightly flattened ventrolaterally; penultimate article about 3.5 times as long as wide; ultimate article with numerous rows of short serrulate setae and longer simple setae, unarmed distally; arthrobranch well developed (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFC6CFECCC7B6FEF2" box="[919,1004,259,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CFEE9C57CFDB4" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
Major cheliped similar in males and females, not significantly larger in males; ischium very short, stout, with row of short spiniform setae (typically four) along mesial margin; merus relatively stout, about 2.2 times as long as wide, distodorsal margin ending in very strong, somewhat curved, sharp tooth; mesial margin rugose, with row of short stout spiniform setae (typically at least five, proximal ones more robust); distomesial tooth very strong, sharp, with broad dorsal lobe; carpus very short, cup-shaped, with broadly rounded distal lobes; chela elongate, with palm subcylindrical in cross-section and about twice as long as fingers; palm surface mostly smooth, except for some low setiferous tubercles on ventral surface, extending to pollex, without any grooves, crests or notches; dactylus slightly longer than pollex, distally rounded, not twisted, with large stout plunger; adhesive disks well developed (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFF64FD8BC543FDB4" box="[159,281,580,605]" captionStart="FIGURE 2" captionStartId="5.[151,250,1910,1932]" captionTargetBox="[184,1388,351,1872]" captionTargetId="figure@5.[181,1413,336,1897]" captionTargetPageId="5" captionText="FIGURE 2. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—major cheliped, chela, lateral view; B—same, mesial view; C—same, coxa to carpus, lateral view; D—same, ischium, merus and carpus, mesial view, conventional setae omitted; E—same, dactylus, lateral view; F—minor cheliped, lateral view; G—same, chela, mesial view; H—same, ischium, merus and carpus, mesial view, conventional setae omitted." httpUri="https://zenodo.org/record/1290347/files/figure.png" pageId="3" pageNumber="577">Fig. 2A–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CFDA7C04EFC2B" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
Minor cheliped not sexually dimorphic, slightly stouter in some males, much more slender than major cheliped, but of nearly equal length; ischium very short, stout, with row of short spiniform setae (typically four or five) along mesial margin; merus moderately stout, at least 2.5 times as long as wide, distodorsal margin ending in very strong, somewhat curved, sharp tooth; mesial margin rugose and with row of short stout spiniform setae (usually at least five, on proximal half); distomesial tooth very strong, sharp, with broad dorsal lobe; carpus short, cup-shaped, with strong distal lobe mesially; chela elongate, slender, with palm subcylindrical in cross-section, slightly but noticeably shorter than fingers (about 0.8–0.9 of finger length); palm surface smooth, without any grooves, crests or notches; fingers subequal in length, crossing distally; dactylus very slightly flattened dorsally, more conical distally, with simple blade-like cutting edge; pollex thicker than dactylus, with deep median groove flanked by two blade-like cutting edges; adhesive disks very small (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFC75FC65C052FC2B" box="[910,1032,938,963]" captionStart="FIGURE 2" captionStartId="5.[151,250,1910,1932]" captionTargetBox="[184,1388,351,1872]" captionTargetId="figure@5.[181,1413,336,1897]" captionTargetPageId="5" captionText="FIGURE 2. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—major cheliped, chela, lateral view; B—same, mesial view; C—same, coxa to carpus, lateral view; D—same, ischium, merus and carpus, mesial view, conventional setae omitted; E—same, dactylus, lateral view; F—minor cheliped, lateral view; G—same, chela, mesial view; H—same, ischium, merus and carpus, mesial view, conventional setae omitted." httpUri="https://zenodo.org/record/1290347/files/figure.png" pageId="3" pageNumber="577">Fig. 2F–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CFC02C08CFA86" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
Second pereiopod slender; ischium and merus subequal in length; carpus with five joints, their ratio approximately equal to 2.5/1/0.6/0.5/1; chela as long as two distal-most carpal joints combined (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFB14FC3EC11DFBE3" box="[1263,1351,1009,1034]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">Fig. 1F</figureCitation>
|
||
). Third pereiopod slender; ischium with short spiniform seta on ventrolateral surface; merus more than six times as long as wide, not inflated, unarmed; carpus about half length of merus, somewhat more slender, unarmed; propodus much longer than carpus but shorter than merus, with six or seven irregularly inserted, stout spiniform setae on ventral margin, not including one pair of additional spiniform setae near dactylar base; dactylus about 0.25 length of propodus, fairly slender, curved, conical, simple, with some setae subdistally (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFBC3FB6BC0E6FB54" box="[1080,1212,1188,1213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">Fig. 1G, H</figureCitation>
|
||
). Fourth pereiopod generally similar to third, slightly more slender. Fifth pereiopod much more slender than third and fourth; ischium unarmed; merus not inflated, about six times as long as wide; carpus 0.9 length of merus; propodus with five or so spiniform setae along ventromesial margin, and one pair of (or sometimes three) spiniform setae near dactylus, in additional to several rows of cleaning setae on ventrolateral surface, on distal-most portion; dactylus conical, simple, slightly more curved than that of third pereiopod, with some setae subdistally (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFB9AFA99C093FA86" box="[1121,1225,1366,1391]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">
|
||
Fig.
|
||
<date id="0A6A1013FFCFFFDDFB6DFA98C0F5FA86" box="[1174,1199,1367,1391]" pageId="3" pageNumber="577">1I</date>
|
||
, J
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CFAB5C725F9CB" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
Male second pleopod with appendix masculina subequal in length to appendix interna, with stiff setae on apex and along mesial margin (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFE44FA51C640FA5E" box="[447,538,1438,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">Fig. 1K</figureCitation>
|
||
). Uropod with both mesial and lateral lobes of protopod ending in sharp point; exopod broad, slightly truncate distally, with strong distolateral tooth; diaeresis strongly sinuous, with strong triangular tooth adjacent and mesial to very thick, black-coloured spiniform seta; endopod narrower than exopod, with short row of small spiniform setae on distal margin (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFCE0F9C6C72BF9CB" box="[795,881,1545,1570]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="3" pageNumber="577">Fig. 1L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CF9E2C066F964" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFF3CF9E2C502F9AF" bold="true" box="[199,344,1581,1606]" pageId="3" pageNumber="577">Colouration</emphasis>
|
||
. Body uniformly bright reddish-orange, with some paler or colourless areas on carapace flanks; antennules and antennae reddish-orange, with deeper red flagella; chelipeds bright orange; walking legs reddish with colourless areas near joints and more distally; tail fan orange-red (
|
||
<figureCitation id="E6EF2A56FFCFFFDDFC4FF9BBC074F964" box="[948,1070,1652,1677]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="3" pageNumber="577">Fig. 3A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCFFFDDFF3CF957C581F93C" pageId="3" pageNumber="577" type="etymology">
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CF957C581F93C" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFF3CF957C51CF958" bold="true" box="[199,326,1688,1713]" pageId="3" pageNumber="577">Etymology</emphasis>
|
||
. The new species' name refers to its overall bright orange colour (
|
||
<taxonomicName id="B9D44D50FFCFFFDDFBB2F956C151F958" authority=", Spanish" authorityName="Spanish" box="[1097,1291,1688,1713]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="naranjo">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFBB2F956C0F8F959" box="[1097,1186,1689,1712]" italics="true" pageId="3" pageNumber="577">naranjo</emphasis>
|
||
, Spanish
|
||
</taxonomicName>
|
||
for orange); used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCFFFDDFF3CF910C58AF8F2" pageId="3" pageNumber="577" type="distribution">
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CF910C58AF8F2" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFF3CF910C503F911" bold="true" box="[199,345,1759,1784]" pageId="3" pageNumber="577">Distribution</emphasis>
|
||
. Presently known only from the
|
||
<typeStatus id="A16F8871FFCFFFDDFD0EF92EC77FF910" box="[757,805,1761,1785]" pageId="3" pageNumber="577">type</typeStatus>
|
||
locality in the
|
||
<collectingRegion id="BC10F831FFCFFFDDFC11F92EC0FBF911" box="[1002,1185,1759,1784]" country="Panama" name="Bocas del Toro" pageId="3" pageNumber="577">Bocas del Toro</collectingRegion>
|
||
Archipelago, on the Caribbean coast of
|
||
<collectingRegion id="BC10F831FFCFFFDDFE8BF8CBC596F8F2" box="[368,460,1796,1819]" country="Panama" name="Panama" pageId="3" pageNumber="577">
|
||
<collectingCountry id="06C37643FFCFFFDDFE8BF8CBC596F8F2" box="[368,460,1796,1819]" name="Panama" pageId="3" pageNumber="577">Panama</collectingCountry>
|
||
</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCFFFDDFF3CF8E8C131F88A" pageId="3" pageNumber="577" type="biology_ecology">
|
||
<paragraph id="7E6B36D3FFCFFFDDFF3CF8E8C131F88A" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="577">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFF3CF8E8C57EF8A9" bold="true" box="[199,292,1831,1856]" pageId="3" pageNumber="577">Ecology</emphasis>
|
||
. All specimens of
|
||
<taxonomicName id="B9D44D50FFCFFFDDFDF8F8E7C627F8D6" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[515,637,1832,1855]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFDF8F8E7C627F8D6" box="[515,637,1832,1855]" italics="true" pageId="3" pageNumber="577">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCFFFDDFD7CF8E8C6BEF8A9" box="[647,740,1831,1856]" pageId="3" pageNumber="577" rank="species">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFD7CF8E8C6BEF8A9" bold="true" box="[647,740,1831,1856]" pageId="3" pageNumber="577">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were collected by carefully breaking apart dead colonies of thin-leaf lettuce coral,
|
||
<taxonomicName id="B9D44D50FFCFFFDDFE6FF884C710F88D" authority="Dana" authorityName="Dana" box="[404,842,1867,1892]" class="Anthozoa" family="Agariciidae" genus="Agaricia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="3" pageNumber="577" phylum="Cnidaria" rank="species" species="tenuifolia">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFE6FF884C633F88A" box="[404,617,1867,1891]" italics="true" pageId="3" pageNumber="577">Agaricia tenuifolia</emphasis>
|
||
Dana (Agariciidae)
|
||
</taxonomicName>
|
||
, on a very shallow coral reef (depth: about
|
||
<quantity id="B92C9B36FFCFFFDDFAD4F884C104F88A" box="[1327,1374,1867,1891]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="577" unit="m" value="2.0">2 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFCFFFDAFF3CF8A1C076FED6" lastPageId="4" lastPageNumber="578" pageId="3" pageNumber="577" type="discussion">
|
||
<paragraph id="7E6B36D3FFCFFFDAFF3CF8A1C076FED6" blockId="3.[151,1437,151,2035]" lastBlockId="4.[151,1437,151,2035]" lastPageId="4" lastPageNumber="578" pageId="3" pageNumber="577">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFF3CF8A1C56EF86E" bold="true" box="[199,308,1902,1927]" pageId="3" pageNumber="577">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="B9D44D50FFCFFFDDFEBCF8A0C65CF86E" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[327,518,1903,1927]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFEBCF8A0C65CF86E" box="[327,518,1903,1927]" italics="true" pageId="3" pageNumber="577">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCFFFDDFDE9F8A1C62BF86E" box="[530,625,1902,1927]" pageId="3" pageNumber="577" rank="species">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFDE9F8A1C62BF86E" bold="true" box="[530,625,1902,1927]" pageId="3" pageNumber="577">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is morphologically closest to three species, namely
|
||
<taxonomicName id="B9D44D50FFCFFFDDFB18F8A0C5FDF842" authority="Crosnier & Forest, 1965" authorityName="Crosnier & Forest" authorityYear="1965" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFB18F8A0C1C6F86E" box="[1251,1436,1903,1927]" italics="true" pageId="3" pageNumber="577">Alpheus blachei</emphasis>
|
||
<bibRefCitation id="1A454B22FFCFFFDDFF6CF85DC5FDF842" box="[151,423,1938,1963]" pageId="3" pageNumber="577" refId="ref5121" refString="Crosnier, A. & Forest, J. (1965) Note preliminaire sur les Alpheidae recueillis par la Calypso dans l'Atlantique oriental tropical (Crustacea Decapoda Natantia). Bulletin du Museum National d'Histoire Naturelle, Serie 2, 36, 602 - 610." type="journal article">Crosnier & Forest, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
from the tropical eastern Atlantic (
|
||
<bibRefCitation id="1A454B22FFCFFFDDFCCBF85DC065F842" box="[816,1087,1938,1963]" pageId="3" pageNumber="577" refId="ref5121" refString="Crosnier, A. & Forest, J. (1965) Note preliminaire sur les Alpheidae recueillis par la Calypso dans l'Atlantique oriental tropical (Crustacea Decapoda Natantia). Bulletin du Museum National d'Histoire Naturelle, Serie 2, 36, 602 - 610." type="journal article">Crosnier & Forest 1965</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1A454B22FFCFFFDDFBB1F85DC0DCF842" box="[1098,1158,1938,1963]" pageId="3" pageNumber="577" refId="ref5168" refString="Crosnier, A. & Forest, J. (1966) Crustaces Decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19: Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article">1966</bibRefCitation>
|
||
);
|
||
<taxonomicName id="B9D44D50FFCFFFDDFB61F85CC545F826" authority="Kim & Abele, 1988" authorityName="Kim & Abele" authorityYear="1988" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFB61F85CC11CF842" box="[1178,1350,1939,1963]" italics="true" pageId="3" pageNumber="577">A. felgenhaueri</emphasis>
|
||
<bibRefCitation id="1A454B22FFCFFFDDFAB6F85DC545F826" pageId="3" pageNumber="577" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFCFFFDDFEAEF878C6CAF826" authority="Carvacho, 1989" authorityName="Carvacho" authorityYear="1989" box="[341,656,1974,1999]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="577" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFCFFFDDFEAEF878C58CF826" box="[341,470,1975,1999]" italics="true" pageId="3" pageNumber="577">A. confusus</emphasis>
|
||
<bibRefCitation id="1A454B22FFCFFFDDFE26F879C6CAF826" box="[477,656,1974,1999]" pageId="3" pageNumber="577" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
, both from the tropical eastern Pacific (
|
||
<bibRefCitation id="1A454B22FFCFFFDDFBB1F879C17FF826" box="[1098,1317,1974,1999]" pageId="3" pageNumber="577" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCFFFDDFACAF879C489F81B" pageId="3" pageNumber="577" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho 1989</bibRefCitation>
|
||
). These four species are characterised by the presence of sharp orbital teeth on or very close to the anterior margin of the orbital hoods; the well-developed acute rostrum, not or only slightly flattened posteriorly; the elongate subcylindrical major chela; the enlarged and darkly coloured spiniform seta of the uropod; and the uniform red-orange, red-brown, or orange-yellow body colour (
|
||
<figureCitation id="E6EF2A56FFC8FFDAFCA4FF10C7F8FF11" box="[863,930,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3</figureCitation>
|
||
), although the colouration of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB14FF2FC1C1FF1E" authorityName="Kim & Abele" authorityYear="1988" box="[1263,1435,223,247]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB14FF2FC1C1FF1E" box="[1263,1435,223,247]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
presently remains unknown. Based on these characteristics, these four species together may form a small clade within
|
||
<taxonomicName id="B9D44D50FFC8FFDAFF1EFEE8C565FED6" authorityName="Anker" authorityYear="2018" box="[229,319,295,319]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFF1EFEE8C565FED6" box="[229,319,295,319]" italics="true" pageId="4" pageNumber="578">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
, which will be referred to as the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD4AFEE8C73FFED6" authorityName="Crosnier & Forest" authorityYear="1965" box="[689,869,295,319]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD4AFEE8C73FFED6" box="[689,869,295,319]" italics="true" pageId="4" pageNumber="578">Alpheus blachei</emphasis>
|
||
</taxonomicName>
|
||
species complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFC8FFDAFF3CFE84C0E2F983" pageId="4" pageNumber="578" type="reference_group">
|
||
<paragraph id="7E6B36D3FFC8FFDAFF3CFE84C61EFCBE" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="578">
|
||
<taxonomicName id="B9D44D50FFC8FFDAFF3CFE84C5D8FE8B" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[199,386,331,355]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFF3CFE84C5D8FE8B" box="[199,386,331,355]" italics="true" pageId="4" pageNumber="578">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFE72FE85C5B9FE8A" box="[393,483,330,355]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFE72FE85C5B9FE8A" bold="true" box="[393,483,330,355]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be separated from the remaining three species of the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB97FE84C085FE8A" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[1132,1247,331,355]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB97FE84C085FE8A" box="[1132,1247,331,355]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
complex mainly by the absence of rows of balaeniceps setae on the minor chela, which are present in both sexes of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB26FEA1C114FE6F" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[1245,1358,366,390]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB26FEA1C114FE6F" box="[1245,1358,366,390]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFC8FFDAFA7FFEA1C4A3FE43" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFA7FFEA1C4A3FE43" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
, and at least in males of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDD8FE5DC688FE43" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[547,722,402,426]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDD8FE5DC688FE43" box="[547,722,402,426]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
(possibly also in females). The rows of balaeniceps setae are typically present on both fingers of the minor chela and are particularly well developed in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB7EFE79C0ACFE27" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[1157,1270,438,462]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB7EFE79C0ACFE27" box="[1157,1270,438,462]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
(confirmed by examination of the comparative material from
|
||
<collectingRegion id="BC10F831FFC8FFDAFD54FE15C77EFE1B" box="[687,804,474,498]" country="Sao Tome and Principe" name="Sao Tome" pageId="4" pageNumber="578">
|
||
<collectingCountry id="06C37643FFC8FFDAFD54FE15C77EFE1B" box="[687,804,474,498]" name="Sao Tome and Principe" pageId="4" pageNumber="578">São Tomé</collectingCountry>
|
||
</collectingRegion>
|
||
, MZUSP 33940, see also
|
||
<figureCitation id="E6EF2A56FFC8FFDAFBA9FE16C0F3FE1B" box="[1106,1193,473,498]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3C</figureCitation>
|
||
) and
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB16FE15C1C1FE18" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[1261,1435,473,497]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB16FE15C1C1FE18" box="[1261,1435,473,497]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
(confirmed by re-examination of the
|
||
<typeStatus id="A16F8871FFC8FFDAFDCDFE32C6C1FDFF" box="[566,667,509,534]" pageId="4" pageNumber="578" type="holotype">holotype</typeStatus>
|
||
, USNM 143347, see also
|
||
<bibRefCitation id="1A454B22FFC8FFDAFC44FE32C0C1FDFF" box="[959,1179,509,534]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig.
|
||
<quantity id="B92C9B36FFC8FFDAFB28FE32C15AFDFF" box="[1235,1280,509,534]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="4" pageNumber="578" unit="g" value="16.0">16g</quantity>
|
||
, h).
|
||
<bibRefCitation id="1A454B22FFC8FFDAFACAFE32C4BBFDD0" pageId="4" pageNumber="578" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho (1989)</bibRefCitation>
|
||
also mentioned (although not illustrated) the balaeniceps condition of the minor chela in the
|
||
<typeStatus id="A16F8871FFC8FFDAFAFEFDEDC16FFDD3" box="[1285,1333,546,570]" pageId="4" pageNumber="578">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFF6CFD8AC57BFDB4" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[151,289,581,605]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFF6CFD8AC57BFDB4" box="[151,289,581,605]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
. Corroborating Carvacho’s observation, a relatively small ovigerous female from Coiba here tentatively identified as
|
||
<taxonomicName id="B9D44D50FFC8FFDAFE50FDA6C674FD69" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[427,558,616,640]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFE50FDA6C674FD69" box="[427,558,616,640]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
(OUMNH.ZC.2014-02-028) also bears a well-developed row of balaeniceps setae on the minor chela (visible in
|
||
<figureCitation id="E6EF2A56FFC8FFDAFDD9FD43C623FD4C" box="[546,633,652,677]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3C</figureCitation>
|
||
). Another distinguishing feature of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFBFDFD42C024FD4D" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1030,1150,653,676]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBFDFD42C024FD4D" box="[1030,1150,653,676]" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFB7FFD43C087FD4C" box="[1156,1245,652,677]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB7FFD43C087FD4C" bold="true" box="[1156,1245,652,677]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, which separates it at once from
|
||
<taxonomicName id="B9D44D50FFC8FFDAFEBCFD7FC5AEFD21" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[327,500,688,712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFEBCFD7FC5AEFD21" box="[327,500,688,712]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDFFFD7FC62CFD21" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[516,630,688,712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDFFFD7FC62CFD21" box="[516,630,688,712]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD4BFD7FC768FD21" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[688,818,688,712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD4BFD7FC768FD21" box="[688,818,688,712]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
is the elongation of the minor cheliped fingers, which are noticeably longer than the palm vs. distinctly shorter than the palm in the other three species (cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFAE1FD1CC1C2FD05" box="[1306,1432,723,748]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Figs. 1F, G</figureCitation>
|
||
,
|
||
<figureCitation id="E6EF2A56FFC8FFDAFF6CFD37C484FCF9" box="[151,222,760,784]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">3A–D</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="1A454B22FFC8FFDAFEB5FD38C675FCF9" box="[334,559,759,784]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig.
|
||
<quantity id="B92C9B36FFC8FFDAFD90FD38C6CDFCF9" box="[619,663,759,784]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="4" pageNumber="578" unit="g" value="16.0">16g</quantity>
|
||
, h;
|
||
<bibRefCitation id="1A454B22FFC8FFDAFD38FD38C78CFCF9" box="[707,982,759,784]" pageId="4" pageNumber="578" refId="ref5121" refString="Crosnier, A. & Forest, J. (1965) Note preliminaire sur les Alpheidae recueillis par la Calypso dans l'Atlantique oriental tropical (Crustacea Decapoda Natantia). Bulletin du Museum National d'Histoire Naturelle, Serie 2, 36, 602 - 610." type="journal article">Crosnier & Forest 1965</bibRefCitation>
|
||
: fig. 3e). Thus, the minor chela with its relatively long, non-expanded, non-balaeniceps fingers, characteristic of the new species, represents a unique feature within the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFE98FCF0C58FFCBE" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[355,469,831,855]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFE98FCF0C58FFCBE" box="[355,469,831,855]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFC8FFDAFF3CFCADC0E2F983" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="578">
|
||
Additional morphological features that can be used to separate
|
||
<taxonomicName id="B9D44D50FFC8FFDAFC78FCACC7A0FC93" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[899,1018,867,890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFC78FCACC7A0FC93" box="[899,1018,867,890]" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFBFAFCADC000FC92" box="[1025,1114,866,891]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBFAFCADC000FC92" bold="true" box="[1025,1114,866,891]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from the other species of the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFF6CFC48C550FC77" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[151,266,902,926]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFF6CFC48C550FC77" box="[151,266,902,926]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
complex are found on the frontal margin of the carapace, antennules, antenna, major cheliped, second and third pereiopods, and uropod. For instance, the new species differs from
|
||
<taxonomicName id="B9D44D50FFC8FFDAFBD9FC65C088FC2B" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[1058,1234,938,962]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBD9FC65C088FC2B" box="[1058,1234,938,962]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
by the large and anteriorly directed orbital teeth, which are much smaller and mesially pointed in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFBCEFC01C0B9FC0F" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[1077,1251,974,998]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBCEFC01C0B9FC0F" box="[1077,1251,974,998]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
; the distolateral tooth of the scaphocerite reaching far beyond the blade vs. hardly overreaching it in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFBA4FC3DC156FBE0" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[1119,1292,1009,1033]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBA4FC3DC156FBE0" box="[1119,1292,1009,1033]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFABEFC3EC1C6FBE3" box="[1349,1436,1009,1034]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Fig. 1A</figureCitation>
|
||
and
|
||
<bibRefCitation id="1A454B22FFC8FFDAFF37FBDAC5E1FBC7" box="[204,443,1045,1070]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig. 16a); and the distinctly more slender second and third pereiopods, which are comparatively much more robust in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDD7FBF6C68DFBB8" ID-CoL="85LHQ" authorityName="Kim & Abele" authorityYear="1988" box="[556,727,1081,1105]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDD7FBF6C68DFBB8" box="[556,727,1081,1105]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
, this difference being particularly obvious in the proportions of the merus, propodus and dactylus of the third pereiopod (cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFCB6FB93C799FB9C" box="[845,963,1116,1141]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Fig. 1F–H</figureCitation>
|
||
and
|
||
<bibRefCitation id="1A454B22FFC8FFDAFC07FB93C086FB9C" box="[1020,1244,1116,1141]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig.
|
||
<date id="0A6A1013FFC8FFDAFAE3FB93C166FB9C" box="[1304,1340,1116,1141]" pageId="4" pageNumber="578" value="2016-01">16i</date>
|
||
–k). The new species also differs from
|
||
<taxonomicName id="B9D44D50FFC8FFDAFE04FB4EC62FFB71" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[511,629,1152,1176]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFE04FB4EC62FFB71" box="[511,629,1152,1176]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
by the noticeably more slender third pereiopod, exemplified by the proportions of the merus (about six times as long as wide in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFCCDFB6AC7F4FB55" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[822,942,1189,1212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFCCDFB6AC7F4FB55" box="[822,942,1189,1212]" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFC4FFB6BC057FB54" box="[948,1037,1188,1213]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFC4FFB6BC057FB54" bold="true" box="[948,1037,1188,1213]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
vs. at most five times in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFADFFB6AC1CFFB55" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[1316,1429,1188,1212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFADFFB6AC1CFFB55" box="[1316,1429,1188,1212]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
, cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFF40FB08C54AFB09" box="[187,272,1223,1248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Fig. 1G</figureCitation>
|
||
and
|
||
<bibRefCitation id="1A454B22FFC8FFDAFEBDFB08C609FB09" box="[326,595,1223,1248]" pageId="4" pageNumber="578" refId="ref5168" refString="Crosnier, A. & Forest, J. (1966) Crustaces Decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19: Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article">Crosnier & Forest 1966</bibRefCitation>
|
||
: fig.
|
||
<quantity id="B92C9B36FFC8FFDAFD70FB07C6F3FB09" box="[651,681,1224,1248]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="578" unit="g" value="3.0">3g</quantity>
|
||
); and the much larger spiniform seta on the uropod (cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFAE1FB08C137FB09" box="[1306,1389,1223,1248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Fig. 1L</figureCitation>
|
||
and
|
||
<bibRefCitation id="1A454B22FFC8FFDAFF6CFB24C5FDFAED" box="[151,423,1259,1284]" pageId="4" pageNumber="578" refId="ref5168" refString="Crosnier, A. & Forest, J. (1966) Crustaces Decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19: Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article">Crosnier & Forest 1966</bibRefCitation>
|
||
: fig. 3h).
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDEFFB23C695FAEA" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[532,719,1260,1284]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDEFFB23C695FAEA" box="[532,719,1260,1284]" italics="true" pageId="4" pageNumber="578">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFD2CFB24C76BFAED" box="[727,817,1259,1284]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD2CFB24C76BFAED" bold="true" box="[727,817,1259,1284]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
may be separated specifically from
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB37FB23C114FAED" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[1228,1358,1260,1284]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB37FB23C114FAED" box="[1228,1358,1260,1284]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the frontal margin, e.g., by the larger, more marginal and more anteriorly directed orbital teeth and longer rostrum, which typically is reaching the distal margin of the first article of the antennular peduncle vs. reaching only half-length of this article in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDE4FA98C6FEFA86" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[543,676,1367,1391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDE4FA98C6FEFA86" box="[543,676,1367,1391]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
; and the distinctly longer stylocerite, which is easily reaching or sometimes reaching far beyond half-length of the second article of the antennular peduncle vs. not exceeding beyond proximal third of this article in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD9CFA50C6B6FA5F" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[615,748,1438,1462]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD9CFA50C6B6FA5F" box="[615,748,1438,1462]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="E6EF2A56FFC8FFDAFCDCFA51C725FA5E" box="[807,895,1438,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1842,1864]" captionTargetBox="[193,1407,204,1798]" captionTargetId="figure@2.[174,1418,195,1825]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—frontal region, dorsal view; B—same, lateral view; C—telson, dorsal view; D—ventromesial carina on first antennular article, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, distal portion of propodus and dactylus, mesial view; I—fifth pereiopod, lateral view; J—same, distal portion of propodus and dactylus, mesial view; K—second pleopod, appendix masculina and appendix interna, mesial view; L—uropod, dorsal view." httpUri="https://zenodo.org/record/1290345/files/figure.png" pageId="4" pageNumber="578">Fig. 1A</figureCitation>
|
||
and
|
||
<bibRefCitation id="1A454B22FFC8FFDAFC46FA51C035FA5E" box="[957,1135,1438,1463]" pageId="4" pageNumber="578" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho 1989</bibRefCitation>
|
||
: fig. 1a). The presence of widely and irregularly spaced, low tubercles along the ventral surface of the major cheliped palm and pollex in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFA7FFA0DC4AAFA17" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFA7FFA0DC4AAFA17" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFF0DFA2AC515FA17" box="[246,335,1509,1534]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFF0DFA2AC515FA17" bold="true" box="[246,335,1509,1534]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="E6EF2A56FFC8FFDAFEA5FA2AC58DFA17" box="[350,471,1509,1534]" captionStart="FIGURE 2" captionStartId="5.[151,250,1910,1932]" captionTargetBox="[184,1388,351,1872]" captionTargetId="figure@5.[181,1413,336,1897]" captionTargetPageId="5" captionText="FIGURE 2. Alpheus naranjo sp. nov., paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—major cheliped, chela, lateral view; B—same, mesial view; C—same, coxa to carpus, lateral view; D—same, ischium, merus and carpus, mesial view, conventional setae omitted; E—same, dactylus, lateral view; F—minor cheliped, lateral view; G—same, chela, mesial view; H—same, ischium, merus and carpus, mesial view, conventional setae omitted." httpUri="https://zenodo.org/record/1290347/files/figure.png" pageId="4" pageNumber="578">Fig. 2A, B</figureCitation>
|
||
), is another feature not seen in the other three species (cf.
|
||
<bibRefCitation id="1A454B22FFC8FFDAFBA4FA2AC131FA17" box="[1119,1387,1509,1534]" pageId="4" pageNumber="578" refId="ref5168" refString="Crosnier, A. & Forest, J. (1966) Crustaces Decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19: Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article">Crosnier & Forest 1966</bibRefCitation>
|
||
: fig. 3c, d;
|
||
<bibRefCitation id="1A454B22FFC8FFDAFF27F9C6C5ECF9CB" box="[220,438,1545,1570]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig. 16e, f;
|
||
<bibRefCitation id="1A454B22FFC8FFDAFDC0F9C6C6B3F9CB" box="[571,745,1545,1570]" pageId="4" pageNumber="578" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho 1989</bibRefCitation>
|
||
: fig. 1e, f). Finally, the body colour of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB63F9C5C14AF9C8" ID-CoL="85LLW" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1176,1296,1546,1569]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB63F9C5C14AF9C8" box="[1176,1296,1546,1569]" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFAEDF9C6C135F9CB" box="[1302,1391,1545,1570]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFAEDF9C6C135F9CB" bold="true" box="[1302,1391,1545,1570]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is a rich orange-red (
|
||
<figureCitation id="E6EF2A56FFC8FFDAFEA8F9E2C597F9AF" box="[339,461,1581,1606]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3A, B</figureCitation>
|
||
), differing from the more deep red-brown with tiny yellowish spots of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB16F9E1C104F9AC" ID-CoL="85LFP" authorityName="Crosnier & Forest" authorityYear="1965" box="[1261,1374,1581,1605]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB16F9E1C104F9AC" box="[1261,1374,1581,1605]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E6EF2A56FFC8FFDAFA95F9E2C4E2F980" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3C</figureCitation>
|
||
) and the more yellow tones of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDEFF99EC6CCF980" ID-CoL="85LGL" authorityName="Carvacho" authorityYear="1989" box="[532,662,1617,1641]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDEFF99EC6CCF980" box="[532,662,1617,1641]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E6EF2A56FFC8FFDAFD5EF99EC6A4F980" box="[677,766,1617,1642]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[195,1400,193,1849]" captionTargetId="figure@6.[195,1400,193,1849]" captionTargetPageId="6" captionText="FIGURE 3. Three species from the Alpheus blachei species complex, dorsal views of living individuals. A—Alpheus naranjo sp. nov., holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—Alpheus blachei Crosnier & Forest, 1965, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—Alpheus confusus Carvacho, 1989, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann." httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="4" pageNumber="578">Fig. 3D</figureCitation>
|
||
), both illustrated here for the first time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36CE6558FFC8FFD9FF3CF9BBC07BFDD4" lastPageId="7" lastPageNumber="581" pageId="4" pageNumber="578" type="description">
|
||
<paragraph id="7E6B36D3FFC8FFDBFF3CF9BBC16CFEF2" blockId="4.[151,1437,151,2035]" lastBlockId="5.[151,1436,151,284]" lastPageId="5" lastPageNumber="579" pageId="4" pageNumber="578">
|
||
The current paucity of the material of the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD56F9BAC77BF964" authorityName="Crosnier & Forest" authorityYear="1965" box="[685,801,1653,1677]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD56F9BAC77BF964" box="[685,801,1653,1677]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
complex, especially from the eastern Pacific, currently precludes a more rigorous molecular analysis of this group. Based on morphological grounds,
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB3AF956C160F959" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1217,1338,1689,1712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB3AF956C160F959" box="[1217,1338,1689,1712]" italics="true" pageId="4" pageNumber="578">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC8FFDAFAB9F957C1C6F958" box="[1346,1436,1688,1713]" pageId="4" pageNumber="578" rank="species">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFAB9F957C1C6F958" bold="true" box="[1346,1436,1688,1713]" pageId="4" pageNumber="578">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
appears to be closest to either
|
||
<taxonomicName id="B9D44D50FFC8FFDAFE1EF972C60CF93D" authorityName="Crosnier & Forest" authorityYear="1965" box="[485,598,1724,1748]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFE1EF972C60CF93D" box="[485,598,1724,1748]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
(in which case it could be its trans-Atlantic sister) or
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB50F972C171F93D" authorityName="Carvacho" authorityYear="1989" box="[1195,1323,1724,1748]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB50F972C171F93D" box="[1195,1323,1724,1748]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
(in which case it could be its trans-isthmian sister), and slightly more distant to
|
||
<taxonomicName id="B9D44D50FFC8FFDAFC26F92FC0D5F911" authorityName="Kim & Abele" authorityYear="1988" box="[989,1167,1760,1784]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFC26F92FC0D5F911" box="[989,1167,1760,1784]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
. The fairly inadequate description of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFEC6F8CBC59BF8F5" authorityName="Carvacho" authorityYear="1989" box="[317,449,1796,1820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFEC6F8CBC59BF8F5" box="[317,449,1796,1820]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="1A454B22FFC8FFDAFE0BF8CCC6F5F8F5" box="[496,687,1795,1820]" pageId="4" pageNumber="578" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho (1989)</bibRefCitation>
|
||
does not make it easy to compare some of the characters of this taxon to those of other species in the
|
||
<taxonomicName id="B9D44D50FFC8FFDAFDC8F8E7C6F9F8D6" authorityName="Crosnier & Forest" authorityYear="1965" box="[563,675,1831,1855]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFDC8F8E7C6F9F8D6" box="[563,675,1831,1855]" italics="true" pageId="4" pageNumber="578">A. blachei</emphasis>
|
||
</taxonomicName>
|
||
complex. For instance,
|
||
<bibRefCitation id="1A454B22FFC8FFDAFC55F8E8C031F8A9" box="[942,1131,1831,1856]" pageId="4" pageNumber="578" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho (1989)</bibRefCitation>
|
||
provided a very superficial drawing of the frontal region and failed to illustrate or to describe any of the walking legs of
|
||
<taxonomicName id="B9D44D50FFC8FFDAFB4EF884C16DF88A" authorityName="Carvacho" authorityYear="1989" box="[1205,1335,1867,1891]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFB4EF884C16DF88A" box="[1205,1335,1867,1891]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
. He also did not compare
|
||
<taxonomicName id="B9D44D50FFC8FFDAFEA3F8A0C580F86E" authorityName="Carvacho" authorityYear="1989" box="[344,474,1903,1927]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFEA3F8A0C580F86E" box="[344,474,1903,1927]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
with the sympatric
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD45F8A0C731F86E" authorityName="Kim & Abele" authorityYear="1988" box="[702,875,1903,1927]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD45F8A0C731F86E" box="[702,875,1903,1927]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
, possibly because it was described only one year earlier (
|
||
<bibRefCitation id="1A454B22FFC8FFDAFF16F85DC590F842" box="[237,458,1938,1963]" pageId="4" pageNumber="578" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
). Based on the present evidence,
|
||
<taxonomicName id="B9D44D50FFC8FFDAFCC4F85CC7B1F842" authorityName="Kim & Abele" authorityYear="1988" box="[831,1003,1939,1963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFCC4F85CC7B1F842" box="[831,1003,1939,1963]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFC8FFDAFBD9F85CC0F9F842" authorityName="Carvacho" authorityYear="1989" box="[1058,1187,1939,1963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFBD9F85CC0F9F842" box="[1058,1187,1939,1963]" italics="true" pageId="4" pageNumber="578">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
represent two distinct species that can be separated from each other by the shape of the scaphocerite (with a very small distolateral tooth hardly exceeding beyond the blade in
|
||
<taxonomicName id="B9D44D50FFC8FFDAFD8EF815C770F81B" authorityName="Kim & Abele" authorityYear="1988" box="[629,810,2010,2034]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="578" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC8FFDAFD8EF815C770F81B" box="[629,810,2010,2034]" italics="true" pageId="4" pageNumber="578">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
vs. with a fairly strong distolateral tooth strongly overreaching the blade in
|
||
<taxonomicName id="B9D44D50FFC9FFDBFE39FF57C61FFF59" authorityName="Carvacho" authorityYear="1989" box="[450,581,152,176]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFE39FF57C61FFF59" box="[450,581,152,176]" italics="true" pageId="5" pageNumber="579">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
) and orbital hoods (distinctly swollen and rounded anteriorly, with minute teeth, in
|
||
<taxonomicName id="B9D44D50FFC9FFDBFF06FF73C5F6FF3D" authorityName="Kim & Abele" authorityYear="1988" box="[253,428,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFF06FF73C5F6FF3D" box="[253,428,188,212]" italics="true" pageId="5" pageNumber="579">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
vs. slightly flared and sinuous anteriorly, with larger teeth, in
|
||
<taxonomicName id="B9D44D50FFC9FFDBFB7FFF73C15DFF3D" authorityName="Carvacho" authorityYear="1989" box="[1156,1287,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFB7FFF73C15DFF3D" box="[1156,1287,188,212]" italics="true" pageId="5" pageNumber="579">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
) (cf.
|
||
<bibRefCitation id="1A454B22FFC9FFDBFAB1FF74C545FF11" pageId="5" pageNumber="579" refId="ref5456" refString="Kim, W. & Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article">Kim & Abele 1988</bibRefCitation>
|
||
: fig. 16a;
|
||
<bibRefCitation id="1A454B22FFC9FFDBFE6FFF10C61EFF11" box="[404,580,223,248]" pageId="5" pageNumber="579" refId="ref4999" refString="Carvacho, A. (1989) Les crevettes carides du Golfe de Californie. IV. Alpheus confusus n. sp. Crustaceana, 56, 58 - 62. https: // doi. org / 10.1163 / 156854089 X 00798" type="journal article">Carvacho 1989</bibRefCitation>
|
||
: fig. 1a), and by the proportions of the third pereiopod dactylus (short and stout in
|
||
<taxonomicName id="B9D44D50FFC9FFDBFF0AFECCC5C7FEF2" authorityName="Kim & Abele" authorityYear="1988" box="[241,413,259,283]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="felgenhaueri">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFF0AFECCC5C7FEF2" box="[241,413,259,283]" italics="true" pageId="5" pageNumber="579">A. felgenhaueri</emphasis>
|
||
</taxonomicName>
|
||
vs. more slender in
|
||
<taxonomicName id="B9D44D50FFC9FFDBFD7AFECCC6A1FEF2" authorityName="Carvacho" authorityYear="1989" box="[641,763,259,283]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFD7AFECCC6A1FEF2" box="[641,763,259,283]" italics="true" pageId="5" pageNumber="579">A confusus</emphasis>
|
||
</taxonomicName>
|
||
(A. Anker, pers. obs., OUMNH.ZC.2014-02-028).
|
||
</paragraph>
|
||
<caption id="2AAB665BFFC9FFDBFF6CF8B9C128F801" httpUri="https://zenodo.org/record/1290347/files/figure.png" pageId="5" pageNumber="579" startId="5.[151,250,1910,1932]" targetBox="[184,1388,351,1872]" targetPageId="5">
|
||
<paragraph id="7E6B36D3FFC9FFDBFF6CF8B9C128F801" blockId="5.[151,1436,1910,2024]" pageId="5" pageNumber="579">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFF6CF8B9C557F862" bold="true" box="[151,269,1910,1932]" pageId="5" pageNumber="579">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="B9D44D50FFC9FFDBFEE2F8B9C59BF865" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[281,449,1910,1932]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="579" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFEE2F8B9C59BF865" box="[281,449,1910,1932]" italics="true" pageId="5" pageNumber="579">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFC9FFDBFE33F8B8C643F865" box="[456,537,1911,1932]" pageId="5" pageNumber="579" rank="species">
|
||
<emphasis id="4CA0EAC1FFC9FFDBFE33F8B8C643F865" bold="true" box="[456,537,1911,1932]" pageId="5" pageNumber="579">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male (cl 8.4 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44448): A—major cheliped, chela, lateral view; B—same, mesial view; C—same, coxa to carpus, lateral view; D—same, ischium, merus and carpus, mesial view, conventional setae omitted; E—same, dactylus, lateral view; F—minor cheliped, lateral view; G—same, chela, mesial view; H—same, ischium, merus and carpus, mesial view, conventional setae omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2AAB665BFFCAFFD8FF6CF88AC665F81D" httpUri="https://zenodo.org/record/1290349/files/figure.png" pageId="6" pageNumber="580" startId="6.[151,250,1861,1883]" targetBox="[195,1400,193,1849]" targetPageId="6">
|
||
<paragraph id="7E6B36D3FFCAFFD8FF6CF88AC665F81D" blockId="6.[151,1436,1861,2036]" pageId="6" pageNumber="580">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FF6CF88AC549F8B3" bold="true" box="[151,275,1861,1883]" pageId="6" pageNumber="580">FIGURE 3.</emphasis>
|
||
Three species from the
|
||
<taxonomicName id="B9D44D50FFCAFFD8FDF2F88AC6F6F8B2" authorityName="Crosnier & Forest" authorityYear="1965" box="[521,684,1861,1883]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="580" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FDF2F88AC6F6F8B2" box="[521,684,1861,1883]" italics="true" pageId="6" pageNumber="580">Alpheus blachei</emphasis>
|
||
</taxonomicName>
|
||
species complex, dorsal views of living individuals. A—
|
||
<taxonomicName id="B9D44D50FFCAFFD8FB0FF88AC1C6F8B2" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1268,1436,1861,1883]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="580" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FB0FF88AC1C6F8B2" box="[1268,1436,1861,1883]" italics="true" pageId="6" pageNumber="580">Alpheus naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCAFFD8FF6CF8ABC4BDF890" box="[151,231,1892,1913]" pageId="6" pageNumber="580" rank="species">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FF6CF8ABC4BDF890" bold="true" box="[151,231,1892,1913]" pageId="6" pageNumber="580">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, male (9.0 mm) from Bocas del Toro, Caribbean coast of Panama (FLMNH UF 44471); B—same, ovigerous female (cl 9.7 mm) from the same locality (OUMNH.ZC. 2018.01.03); C—
|
||
<taxonomicName id="B9D44D50FFCAFFD8FC6BF84DC174F871" authority="Crosnier & Forest, 1965" authorityName="Crosnier & Forest" authorityYear="1965" box="[912,1326,1922,1944]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="580" phylum="Arthropoda" rank="species" species="blachei">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FC6BF84DC068F871" box="[912,1074,1922,1944]" italics="true" pageId="6" pageNumber="580">Alpheus blachei</emphasis>
|
||
Crosnier & Forest, 1965
|
||
</taxonomicName>
|
||
, ovigerous female (cl 8.3 mm) from São Tomé (OUMNH.ZC.2016-01-036); D—
|
||
<taxonomicName id="B9D44D50FFCAFFD8FC98F86EC0E5F85E" authority="Carvacho, 1989" authorityName="Carvacho" authorityYear="1989" box="[867,1215,1953,1975]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="580" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="4CA0EAC1FFCAFFD8FC98F86EC04FF85E" box="[867,1045,1953,1975]" italics="true" pageId="6" pageNumber="580">Alpheus confusus</emphasis>
|
||
Carvacho, 1989
|
||
</taxonomicName>
|
||
, ovigerous female (cl 5.4 mm) from Coiba, Pacific coast of Panama (OUMNH.ZC.2014-02-028). Photographic credits: (A, B) by Robert Lasley; (C) by the author; (D) by Ingo S. Wehrtmann.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E6B36D3FFCBFFD9FF3CFF58C733FE61" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="581">
|
||
The only two western Atlantic snapping shrimps that have some morphological similarities with
|
||
<taxonomicName id="B9D44D50FFCBFFD9FB06FF57C12EFF46" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1277,1396,152,175]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FB06FF57C12EFF46" box="[1277,1396,152,175]" italics="true" pageId="7" pageNumber="581">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCBFFD9FA81FF58C492FF3C" pageId="7" pageNumber="581" rank="species">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FA81FF58C492FF3C" bold="true" pageId="7" pageNumber="581">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, such as the armed orbital hoods, the general shape of the major and minor cheliped, and the dark-coloured spiniform seta of the uropod, are
|
||
<taxonomicName id="B9D44D50FFCBFFD9FDEAFF2FC76FFF11" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[529,821,223,248]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="formosus">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FDEAFF2FC6C2FF11" box="[529,664,224,248]" italics="true" pageId="7" pageNumber="581">A. formosus</emphasis>
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD64FF10C76FFF11" box="[671,821,223,248]" pageId="7" pageNumber="581" refId="ref5400" refString="Gibbes, L. R. (1850) On the carcinological collections of the cabinets of natural history in the United States: with an enumeration of the species contained therein and descriptions of new species. Proceedings of third meeting of American Association for Advancement of Science, 3, 165 - 201." type="journal article">Gibbes, 1850</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFCBFFD9FC95FF2FC1C3FF11" authority="Anker, Hurt & Knowlton, 2008" authorityName="Anker, Hurt & Knowlton" authorityYear="2008" box="[878,1433,223,248]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="paraformosus">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FC95FF2FC070FF11" box="[878,1066,224,248]" italics="true" pageId="7" pageNumber="581">A. paraformosus</emphasis>
|
||
<bibRefCitation id="1A454B22FFCBFFD9FBCAFF10C1C3FF11" box="[1073,1433,223,248]" pageId="7" pageNumber="581" refId="ref4658" refString="Anker, A., Hurt, C. & Knowlton, N. (2008) Revision of the Alpheus formosus Gibbes, 1850 complex, with redescription of A. formosus and description of a new species from the tropical western Atlantic (Crustacea: Decapoda: Alpheidae). Zootaxa, 1707, 1 - 22." type="journal article">Anker, Hurt & Knowlton, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
. Both are readily distinguishable from the new species by their dorsally abruptly flattened post-rostral areas, their orbital teeth distinctly removed from the anterior orbital margin, their much shorter stylocerites, as well as by their very different and characteristic colour patterns, dominated by one median longitudinal yellow or cream band, two lateral brown bands and bluish walking legs (
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD69FEA0C707FE61" box="[658,861,367,392]" pageId="7" pageNumber="581" refId="ref4658" refString="Anker, A., Hurt, C. & Knowlton, N. (2008) Revision of the Alpheus formosus Gibbes, 1850 complex, with redescription of A. formosus and description of a new species from the tropical western Atlantic (Crustacea: Decapoda: Alpheidae). Zootaxa, 1707, 1 - 22." type="journal article">
|
||
Anker
|
||
<emphasis id="4CA0EAC1FFCBFFD9FD1AFEBEC743FE61" box="[737,793,368,392]" italics="true" pageId="7" pageNumber="581">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E6B36D3FFCBFFD9FF3CFE5AC07BFDD4" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="581">
|
||
In the Indo-West Pacific,
|
||
<taxonomicName id="B9D44D50FFCBFFD9FE12FE5BC769FE45" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[489,819,404,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="splendidus">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FE12FE5BC6DBFE45" box="[489,641,404,428]" italics="true" pageId="7" pageNumber="581">A. splendidus</emphasis>
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD72FE5BC769FE45" box="[649,819,404,429]" pageId="7" pageNumber="581" refId="ref5042" refString="Coutiere, H. (1897) Note sur quelques Alpheides nouveaux ou peu connus rapportes de Djibouti (Afrique orientale). Bulletin du Museum d'Histoire Naturelle, 3, 233 - 236. https: // doi. org / 10.5962 / bhl. part. 19259" type="journal article">Coutière, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9D44D50FFCBFFD9FCC4FE5BC003FE45" authority="De Man, 1908" authorityName="De Man" authorityYear="1908" box="[831,1113,404,428]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="facetus">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FCC4FE5BC7F4FE45" box="[831,942,404,428]" italics="true" pageId="7" pageNumber="581">A. facetus</emphasis>
|
||
<bibRefCitation id="1A454B22FFCBFFD9FC4DFE5AC003FE45" box="[950,1113,404,428]" pageId="7" pageNumber="581" refId="ref5308" refString="De Man, J. G. (1908) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ". Notes from the Leyden Museum, 30, 98 - 112." type="journal article">De Man, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9D44D50FFCBFFD9FB69FE5BC495FE39" authority="Coutiere, 1908" authorityName="Coutiere" authorityYear="1908" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="coetivensis">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FB69FE5BC177FE45" box="[1170,1325,404,428]" italics="true" pageId="7" pageNumber="581">A. coetivensis</emphasis>
|
||
<bibRefCitation id="1A454B22FFCBFFD9FACFFE5BC495FE39" pageId="7" pageNumber="581" refId="ref5091" refString="Coutiere, H. (1908) Sur quelques nouvelles especes d'Alpheidae. Bulletin de la Societe Philomathique de Paris, Serie 9, 10, 191 - 216." type="journal article">Coutière, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="B9D44D50FFCBFFD9FF00FE77C600FE39" authority="(Banner, 1953)" authorityName="Banner" authorityYear="1953" baseAuthorityName="Banner" baseAuthorityYear="1953" box="[251,602,439,464]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="edmondsoni">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FF00FE77C5F9FE39" box="[251,419,440,464]" italics="true" pageId="7" pageNumber="581">A. edmondsoni</emphasis>
|
||
(
|
||
<bibRefCitation id="1A454B22FFCBFFD9FE4EFE77C60BFE39" box="[437,593,439,464]" pageId="7" pageNumber="581" refId="ref4858" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article">Banner, 1953</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
] share some general characteristics with
|
||
<taxonomicName id="B9D44D50FFCBFFD9FBCFFE77C0F4FE26" authority="Anker, 2018" authorityName="Anker" authorityYear="2018" box="[1076,1198,440,463]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="581" phylum="Arthropoda" rank="species" species="naranjo" status="sp. nov.">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FBCFFE77C0F4FE26" box="[1076,1198,440,463]" italics="true" pageId="7" pageNumber="581">A. naranjo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="579357BAFFCBFFD9FB43FE78C14EFE39" box="[1208,1300,439,464]" pageId="7" pageNumber="581" rank="species">
|
||
<emphasis id="4CA0EAC1FFCBFFD9FB43FE78C14EFE39" bold="true" box="[1208,1300,439,464]" pageId="7" pageNumber="581">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but can be easily distinguished from the new species by a combination of morphological features, and most easily, by their strikingly different colour patterns (
|
||
<bibRefCitation id="1A454B22FFCBFFD9FDDDFE30C690FDF1" box="[550,714,511,536]" pageId="7" pageNumber="581" refId="ref5091" refString="Coutiere, H. (1908) Sur quelques nouvelles especes d'Alpheidae. Bulletin de la Societe Philomathique de Paris, Serie 9, 10, 191 - 216." type="journal article">Coutière 1908</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD2DFDCFC729FDF1" box="[726,883,512,536]" pageId="7" pageNumber="581" refId="ref5308" refString="De Man, J. G. (1908) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ". Notes from the Leyden Museum, 30, 98 - 112." type="journal article">De Man 1908</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1A454B22FFCBFFD9FC84FDCFC7E3FDF1" box="[895,953,512,536]" pageId="7" pageNumber="581" refId="ref5344" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465." type="journal article">1911</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCBFFD9FC3FFDCFC00DFDF1" box="[964,1111,511,536]" pageId="7" pageNumber="581" refId="ref4858" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article">Banner 1953</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCBFFD9FB98FE30C178FDF1" box="[1123,1314,511,536]" pageId="7" pageNumber="581" refId="ref4951" refString="Bhuti, G. S., Shakuntala, S. & Sankolli, K. N. (1975) On a new record of Alpheus splendidus Coutiere from India and its colouration. The Karnatak University Journal, Science, 20, 292 - 297." type="journal article">
|
||
Bhuti
|
||
<emphasis id="4CA0EAC1FFCBFFD9FB53FDCEC082FDF1" box="[1192,1240,512,536]" italics="true" pageId="7" pageNumber="581">et al</emphasis>
|
||
. 1975
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCBFFD9FAD5FDCFC570FDD5" pageId="7" pageNumber="581" refId="ref4886" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article">Banner & Banner 1982</bibRefCitation>
|
||
; for colour photographs see
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD93FDEBC6AAFDD5" box="[616,752,548,573]" pageId="7" pageNumber="581" refId="ref4776" refString="Anker, A. (2010) On two snapping shrimps, Alpheus baccheti n. sp. and A. coetivensis Coutiere from the Tuamotu Islands. Zootaxa, 2492, 49 - 62." type="journal article">Anker 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A454B22FFCBFFD9FD07FDEBC04FFDD4" box="[764,1045,548,573]" pageId="7" pageNumber="581" refId="ref4812" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34 (Supplement), 343 - 454." type="journal article">Anker & De Grave 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |