734 lines
84 KiB
XML
734 lines
84 KiB
XML
<document ID-CLB-Dataset="83510" ID-DOI="http://dx.doi.org/10.3897/vz.72.e84516" ID-GBIF-Dataset="dc582891-1697-4a2b-878c-f5b3c0327037" ID-Pensoft-Pub="2625-8498-72-433" ID-Pensoft-UUID="257AA52675F45F79A9960A25A9C39668" ID-ZooBank="3001932D04B64A2AAB254FFF3FF96E30" ModsDocID="2625-8498-72-433" checkinTime="1656404867800" checkinUser="pensoft" docAuthor="Yeung, Ho Yuen, Lau, Michael W. N. & Yang, Jian-Huan" docDate="2022" docId="5DB6C563C9C851E4A1FDD68E5FBF1FC1" docLanguage="en" docName="VertZool 72: 433-444" docOrigin="Vertebrate Zoology 72" docPubDate="2022-06-27" docSource="http://dx.doi.org/10.3897/vz.72.e84516" docTitle="Calamaria arcana Yeung & Lau & Yang 2022, sp. nov." docType="treatment" docUuid="CFCB3C7F-842E-4D6D-A11F-C77189B11224" docUuidSource="ZooBank" docVersion="5" id="257AA52675F45F79A9960A25A9C39668" lastPageNumber="433" masterDocId="257AA52675F45F79A9960A25A9C39668" masterDocTitle="A new species of Calamaria (Squamata: Colubridae) from Guangdong Province, southern China" masterLastPageNumber="444" masterPageNumber="433" pageNumber="433" updateTime="1732858539943" updateUser="ExternalLinkService">
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<mods:titleInfo id="97E64D781DCBF8FC41FD4B3CFCAB41BE">
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<mods:title id="E1D57E6E99AFE06946C66952EA149CFE">A new species of Calamaria (Squamata: Colubridae) from Guangdong Province, southern China</mods:title>
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<mods:namePart id="008933DB432822D5C0D52B7E3225C8F4">Yeung, Ho Yuen</mods:namePart>
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<mods:affiliation id="2B2DE31191573AC45D6B5FBC745F82EA">Kadoorie Conservation China, Kadoorie Farm and Botanic Garden, Lam Kam Road, Tai Po, Hong Kong SAR, PR China</mods:affiliation>
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<mods:roleTerm id="61BB603FE9305D5D402F240AC5567716">Author</mods:roleTerm>
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<mods:namePart id="EDDF8152769EE23AACC7B45A145E8FC8">Lau, Michael W. N.</mods:namePart>
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<mods:affiliation id="67FA08179C26FE355FE363AC8756B3D9">Hong Kong Wetlands Conservation Association, Hong Kong SAR, PR China</mods:affiliation>
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<mods:namePart id="9C21D1D819E9D503F3FB0114310E4415">Yang, Jian-Huan</mods:namePart>
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<mods:nameIdentifier id="4BFCEB0564D0F400465D8CA7264E61B5" type="ORCID">https://orcid.org/0000-0002-3706-5380</mods:nameIdentifier>
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<mods:affiliation id="88DAAB7E65D889A658ED994509A2922E">Kadoorie Conservation China, Kadoorie Farm and Botanic Garden, Lam Kam Road, Tai Po, Hong Kong SAR, PR China</mods:affiliation>
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<mods:nameIdentifier id="3D17C19697169E8ADB0EDDCDF2D69AA7" type="email">jhyang@kfbg.org</mods:nameIdentifier>
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<mods:title id="A9EE5132F1B640D726B8D60226FB5D5F">Vertebrate Zoology</mods:title>
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<mods:date id="D251745E0E32C528C0C94D576F464F13">2022</mods:date>
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<mods:number id="41AA32C358CE72110F7A72C90AFB83DC">72</mods:number>
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<treatment id="5DB6C563C9C851E4A1FDD68E5FBF1FC1" ID-GBIF-Taxon="196365423" LSID="urn:lsid:zoobank.org:act:CFCB3C7F-842E-4D6D-A11F-C77189B11224" httpUri="http://treatment.plazi.org/id/5DB6C563C9C851E4A1FDD68E5FBF1FC1" lastPageNumber="433" pageId="0" pageNumber="433" scope_class="Squamata" scope_family="Colubridae">
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<subSubSection id="2729168ED71AC0DDDB61DF44ED0379FA" pageId="0" pageNumber="433" type="nomenclature">
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<paragraph id="AB31D6B427B1E9A90510B28D45CC0690" pageId="0" pageNumber="433">
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<taxonomicName id="E6990A5FD89B353700A68C07F27864A3" LSID="https://zoobank.org/CFCB3C7F-842E-4D6D-A11F-C77189B11224" authority="Yeung & Lau & Yang, 2022" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana" status="sp. nov.">Calamaria arcana</taxonomicName>
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<taxonomicNameLabel id="808099B396E1FD91BBCC90D2D5442625" pageId="0" pageNumber="433">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5107A70DFB4DEAA1F38C6D5E7DD4853C" pageId="0" pageNumber="433" type="description">
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<paragraph id="5782C0526C75BFB83011A05D54349C1A" pageId="0" pageNumber="433">
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<figureCitation id="CC37EB4EA1313CD5C0D33FD4225DB3B1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Calamaria arcana sp. nov. (KFBG 14611) in life: A dorsolateral view B ventral view C lateral view of left side of head. Photographs by Michael W. N. Lau." figureDoi="10.3897/vz.72.e84516.figure2" httpUri="https://binary.pensoft.net/fig/706621" pageId="0" pageNumber="433">Figs 2</figureCitation>
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<figureCitation id="88B741433C62A620DC4C1908E1AA7350" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of the holotype of Calamaria arcana sp. nov. (KFBG 14611): A right side B left side C dorsal view D ventral view." figureDoi="10.3897/vz.72.e84516.figure3" httpUri="https://binary.pensoft.net/fig/706622" pageId="0" pageNumber="433">, 3</figureCitation>
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<figureCitation id="29D2B4AFB805D7E0FEB9E61A7036A27D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Calamaria arcana sp. nov. (KFBG 14611): A dorsal view B ventral view." figureDoi="10.3897/vz.72.e84516.figure4" httpUri="https://binary.pensoft.net/fig/706623" pageId="0" pageNumber="433">, 4</figureCitation>
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<figureCitation id="733D298FA12D3154E3747CCF8BAA693E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">, 5G-I</figureCitation>
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||
<figureCitation id="B64D3FE86FA4F026CDBB427A38F280C7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">, 6G-I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FCAF6F33BF80917CDC8BC7AE202F8815" pageId="0" pageNumber="433" type="reference_group">
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<paragraph id="FE06E4CB796BBC2B7602071CB9026651" pageId="0" pageNumber="433">
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||
<taxonomicName id="7EF708D4FAD2BEEF7879926201185209" authority="Yeung & Lau & Yang, 2022" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Calamaria pavimentata" order="Squamata" pageId="0" pageNumber="433" rank="species" species="pavimentata" status="sp. nov.">Calamaria pavimentata</taxonomicName>
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||
-
|
||
<bibRefCitation id="74EEA90F5643E1EBFD71A5B94C7BA9BC" author="Johns, GC" journalOrPublisher="Molecular Biology and Evolution," pageId="0" pageNumber="433" refId="B9" refString="Kadoorie Farm and Botanic Garden (2003) Report of a Rapid Biodiversity Assessment at Nanling National Nature Reserve, Northwest Guangdong, China, June-July 2000, South China Forest Biodiversity Survey Report Series: No. 29.KFBG, Hong Kong SAR, 33 pp." year="1998">Kadoorie Farm and Botanic Garden (2003)</bibRefCitation>
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||
;
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||
<bibRefCitation id="61D1BC38A28C619359AD2E1C852DCC86" author="Li, ZC" journalOrPublisher="Guangdong Science and Technology Press, Guangzhou" pageId="0" pageNumber="433" refId="B17" refString="Li, ZC, Xiao, Z, Liu, SR, 2011. Amphibians and reptiles of Guangdong. Guangdong Science and Technology Press, Guangzhou" title="Amphibians and reptiles of Guangdong." year="2011">Li et al. (2011)</bibRefCitation>
|
||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="85E3E44BA408B8922A329FC1EC6AA467" pageId="0" pageNumber="433" type="suggested common names">
|
||
<paragraph id="8DC4101C4F85232B834E4FEC9B5D92AE" pageId="0" pageNumber="433">Suggested common names.</paragraph>
|
||
<paragraph id="36EB42FBC0CC645CD3AD08AD64B2BE39" pageId="0" pageNumber="433">
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||
Lingnan Reed Snake (English),
|
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<normalizedToken id="64098A946A32AAECF47863428699B412" originalValue="岭南两头蛇">岭南两头蛇</normalizedToken>
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(Chinese)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="6E287C5D61C74A08499433F2F2ED3B67" pageId="0" pageNumber="433" type="holotype">
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<paragraph id="7B4BB176D640A45EF21C7404875E392A" pageId="0" pageNumber="433">Holotype.</paragraph>
|
||
<paragraph id="5D0C26CC15E469D28D3886A873D4E0B9" pageId="0" pageNumber="433">
|
||
KFBG 14611 (Figs
|
||
<figureCitation id="01886816E97DCB60EE720440A8BD03A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Calamaria arcana sp. nov. (KFBG 14611) in life: A dorsolateral view B ventral view C lateral view of left side of head. Photographs by Michael W. N. Lau." figureDoi="10.3897/vz.72.e84516.figure2" httpUri="https://binary.pensoft.net/fig/706621" pageId="0" pageNumber="433">2</figureCitation>
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||
,
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||
<figureCitation id="78ECD4F9F3800EDB079F88B79860AA6D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head of the holotype of Calamaria arcana sp. nov. (KFBG 14611): A right side B left side C dorsal view D ventral view." figureDoi="10.3897/vz.72.e84516.figure3" httpUri="https://binary.pensoft.net/fig/706622" pageId="0" pageNumber="433">3</figureCitation>
|
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,
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||
<figureCitation id="9934397A875B1F886805181E3F53137D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Calamaria arcana sp. nov. (KFBG 14611): A dorsal view B ventral view." figureDoi="10.3897/vz.72.e84516.figure4" httpUri="https://binary.pensoft.net/fig/706623" pageId="0" pageNumber="433">4</figureCitation>
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||
,
|
||
<figureCitation id="CB06312DD63FE7580FAB1D265963B9A8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">5G-I</figureCitation>
|
||
,
|
||
<figureCitation id="E10F24816330AD61F3CA33F9A9868652" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6G-I</figureCitation>
|
||
), male, Mt. Dadongshan, Nanling National Nature Reserve, Lianzhou County, Qingyuan City, Guangdong Province, China (中国广东南岭国家级自然保护区大东山管理站). The specimen was found at between the hydroelectric station near Dadongshan Management Station (
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<geoCoordinate id="27C61CE289A1CC34B68DE6FBD1EFA452" degrees="24" direction="north" minutes="55" orientation="latitude" precision="1" seconds="58.5" value="24.932917">
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24°55
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<normalizedToken id="3BD89AB6FEE62C9E918A8340B96B58DC" originalValue="’58.5”">'58.5"</normalizedToken>
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N
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</geoCoordinate>
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<geoCoordinate id="316DA3F16598857422E32D08398462E1" degrees="112" direction="east" minutes="42" orientation="longitude" precision="1" seconds="51.8" value="112.714386">
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112°42
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<normalizedToken id="1D19CE264C9A184DC5CB459AD291904D" originalValue="’51.8”">'51.8"</normalizedToken>
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E
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</geoCoordinate>
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) and Shizi Ping (石仔坪), 800-900 m above sea level, collected by Michael W.N. Lau on 7 July 2000 at 9:20 am.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FA74A30D6D052F4562253026EDF638E9" pageId="0" pageNumber="433" type="etymology">
|
||
<paragraph id="876B953032D73084AA2CBF01A437BD99" pageId="0" pageNumber="433">Etymology.</paragraph>
|
||
<paragraph id="3F276E8A2448EA33584BF5508B91C5E5" pageId="0" pageNumber="433">
|
||
The species epithet "
|
||
<taxonomicName id="508F3FC5AB7DA67888578B7B16BFF7B1" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">Calamaria arcana</taxonomicName>
|
||
" is the nominative form the Latin adjective
|
||
<normalizedToken id="3F8B37334B6C9261FB449AB46A569D49" originalValue="“arcanus”">"arcanus"</normalizedToken>
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||
meaning
|
||
<normalizedToken id="BBC332DD798432D09B42D2EA6E5B9334" originalValue="“hidden”">"hidden"</normalizedToken>
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,
|
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<normalizedToken id="590E1CBE7604461186D66BFEC6473838" originalValue="“secret”">"secret"</normalizedToken>
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||
and is given in feminine form to match the female genitive declension of the genus name
|
||
<taxonomicName id="C6EC972DAB24DD706DCADA0D381DA602" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria" order="Squamata" pageId="0" pageNumber="433" rank="genus">
|
||
<emphasis id="466FEB3F64E3FEF7224BA3605DCA257B" italics="true" pageId="0" pageNumber="433">Calamaria</emphasis>
|
||
</taxonomicName>
|
||
. It refers to the fact that this species is difficult to find with only a single specimen discovered during our long term survey in southern China. For common name we suggested as "Lingnan reed snake" in English and "Ling Nan Liang Tou She (
|
||
<normalizedToken id="30575972FF9A4DB2F69C869B181071ED" originalValue="岭南两头蛇)”">岭南两头蛇)"</normalizedToken>
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||
in Chinese, referring to the Lingnan (岭南, means 'South of the Nanling
|
||
<normalizedToken id="9B6D1FA70627659E63A2FAAB658E2C57" originalValue="Mountains’">Mountains'</normalizedToken>
|
||
) region where the new species is found. Lingnan is a geographic area referring to the lands in the south of the Nanling Mountains, covering the modern Chinese provinces of Guangdong, Guangxi and Hainan, as well as northern Vietnam.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7B3CE1338DEB2E72D8E143D08650CC9D" pageId="0" pageNumber="433" type="diagnosis">
|
||
<paragraph id="1B1363FDD9FCF2FDD22193989D49849A" pageId="0" pageNumber="433">Diagnosis.</paragraph>
|
||
<paragraph id="FD64EC1E9FE2CE648CDF19993B269A21" pageId="0" pageNumber="433">
|
||
<taxonomicName id="CDFD1D6409D2AB35AD7A70DC4C7FEECE" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="143E32A77C0604A8F441EDF234213D94" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB72EE4B3498CB0917D8B4E39F252E31" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
is assigned to the genus
|
||
<taxonomicName id="C394BD0693F40537BAEFA711D129E2B6" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria" order="Squamata" pageId="0" pageNumber="433" rank="genus">
|
||
<emphasis id="138AC93C23A1E3E02FE1615DF5D70AC6" italics="true" pageId="0" pageNumber="433">Calamaria</emphasis>
|
||
</taxonomicName>
|
||
by its molecular phylogenetic position and the following morphological characters: dorsal scales in 13 rows throughout body, possessing elongate, cylindrical bodies, along with internasals and prefrontals fused, four supralabials and five infralabials (
|
||
<bibRefCitation id="6D3BB8759F65FA19F128321CC2AA3D14" author="Inger, RF" journalOrPublisher="Fieldiana Zoology" pageId="0" pageNumber="433" pagination="1 - 304" refId="B6" refString="Inger, RF, Marx, H, 1965. The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria. Fieldiana Zoology 49: 1 - 304" title="The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria." volume="49" year="1965">Inger and Marx 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="00991BE9CAFD2C7A3FD11F7BB9507910" DOI="https://doi.org/10.3105/1881-1019-27.2.71" author="Ziegler, T" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="433" pagination="71 - 80" refId="B36" refString="Ziegler, T, Nguyen, VS, Nguyen, QT, 2008. A new reed snake of the genus Calamaria Boie (Squamata: Colubridae) from Vietnam. Current Herpetology 27 (2): 71 - 80, DOI: https://doi.org/10.3105/1881-1019-27.2.71" title="A new reed snake of the genus Calamaria Boie (Squamata: Colubridae) from Vietnam." url="https://doi.org/10.3105/1881-1019-27.2.71" volume="27" year="2008">Ziegler et al. 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6306B5C1DDA596E3868EC91AFEC4ADEF" pageId="0" pageNumber="433">
|
||
<taxonomicName id="E72C85EBDE13B22F8575729E3BF78D1E" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="C261724A7A1BD15E063FA3EFD806FC3A" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5C19B6CD7878BBDECEFD6FC397DD7C80" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
could be distinguished from all other species of
|
||
<taxonomicName id="6FADB5D68604EBE7580ADFDBDAEF285B" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria" order="Squamata" pageId="0" pageNumber="433" rank="genus">
|
||
<emphasis id="52FC5732A047F5247AE7CB40FC6F9185" italics="true" pageId="0" pageNumber="433">Calamaria</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following characters: (1) ten modified maxillary teeth; (2) four supralabials, second and third supralabials entering orbit; (3) preocular present; (4) mental not touching anterior chin shields; (5) six scales and shields surrounding the paraparietal; (6) 170 ventral scales; (7) 22 paired subcaudals; (8) tail not gradually tapering, abruptly tapering at the tip; (9) dorsal scales reduced to five rows above last subcaudal at tail; (10) dorsum of body and tail brownish; (11) dark collar on nuchal region absent; (12) two outermost dorsal scale rows light yellow with upper margins partly dark pigmented; (13) ventral scales immaculate, without dark outermost corners and pigmentation anteriorly; and (14) absence of distinct dark longitudinal line or scattered spots on the underside of tail.
|
||
</paragraph>
|
||
<caption id="12A36BA50E4C4CBDFC3FEAE3F4DA7858" doi="10.3897/vz.72.e84516.figure2" httpUri="https://binary.pensoft.net/fig/706621" pageId="0" pageNumber="433" start="Figure 2" startId="F2">
|
||
<paragraph id="5DB4B33A52CBED8654E7749B0970EE79" pageId="0" pageNumber="433">
|
||
<emphasis id="2CC6B6C585C0B909898C9A6E1D0DEC48" bold="true" pageId="0" pageNumber="433">Figure 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="2BDCA238E43F1EF275D9341530009792" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="CBFDFAE807ADE3D44806957C9ACE743B" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="EABE28460FC785C396B368CFFF4A97FA" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
(KFBG 14611) in life:
|
||
<emphasis id="202D37888E1742FC86D9F11B113E4C1F" bold="true" pageId="0" pageNumber="433">A</emphasis>
|
||
dorsolateral view
|
||
<emphasis id="DA26DF0235395DC67E1EF0F9CC6E29BA" bold="true" pageId="0" pageNumber="433">B</emphasis>
|
||
ventral view
|
||
<emphasis id="8488179627B428CC7FAD0BD9C0F4861D" bold="true" pageId="0" pageNumber="433">C</emphasis>
|
||
lateral view of left side of head. Photographs by Michael W.N. Lau.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2314FDD9D61A35DE7192D8CB9055AE94" doi="10.3897/vz.72.e84516.figure3" httpUri="https://binary.pensoft.net/fig/706622" pageId="0" pageNumber="433" start="Figure 3" startId="F3">
|
||
<paragraph id="2ABCFE5A2175F3D682AD9B210135C9A1" pageId="0" pageNumber="433">
|
||
<emphasis id="F215F0636FF689BEF2C1A91485C3732C" bold="true" pageId="0" pageNumber="433">Figure 3.</emphasis>
|
||
Head of the holotype of
|
||
<taxonomicName id="3245CD002CF6D1B401659F25A8D51AE0" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="19C046891253BBE696406D6E09158103" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="24EFBF90C9E1389E63867E520D378C90" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
(KFBG 14611):
|
||
<emphasis id="9A3FA718EA0E6D08878A37DA3D84E2E7" bold="true" pageId="0" pageNumber="433">A</emphasis>
|
||
right side
|
||
<emphasis id="8E7B64C3AFC0B1AE75E68C790555885C" bold="true" pageId="0" pageNumber="433">B</emphasis>
|
||
left side
|
||
<emphasis id="6F3A6D6B0F661E26C0857F5890312F92" bold="true" pageId="0" pageNumber="433">C</emphasis>
|
||
dorsal view
|
||
<emphasis id="9B6FA7B12FC9F1DFA98AC7E6E8D17567" bold="true" pageId="0" pageNumber="433">D</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="608607BC6B37C81A19520458FE4F82A5" doi="10.3897/vz.72.e84516.figure4" httpUri="https://binary.pensoft.net/fig/706623" pageId="0" pageNumber="433" start="Figure 4" startId="F4">
|
||
<paragraph id="4DBB7319122F3D608848C036B87B1A67" pageId="0" pageNumber="433">
|
||
<emphasis id="BDBC646342C879EB8F0C439D11ABD32B" bold="true" pageId="0" pageNumber="433">Figure 4.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="BE94ACA2F7A4FD11198033DE6497E099" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="ED44291C94CF91C7AD8330F7D9D8EA99" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9D762D83B9F3B94EBBD4CB2726463562" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
(KFBG 14611):
|
||
<emphasis id="6F07570EF9B9DB163D0378D72B2BA939" bold="true" pageId="0" pageNumber="433">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="92FA16784AA0F33E7C1BCF78E290645B" bold="true" pageId="0" pageNumber="433">B</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5B62A3D274D70A483C03E9643FD94C7D" pageId="0" pageNumber="433" type="description">
|
||
<paragraph id="0F6AA9658E6318E530FE416A8EA8F9F0" pageId="0" pageNumber="433">Description of holotype.</paragraph>
|
||
<paragraph id="504CA10355E745F74ECA6A330DC0938D" pageId="0" pageNumber="433">Male. Rostral wider than high (height 1.8 mm, width 2.2 mm), portion visible from above about equal to prefrontal suture; prefrontal shorter than frontal, not entering orbit, and touching first two supralabials; frontal hexagonal, about 2.2 times maximum width of supraocular; paraparietal surrounded by six shields and scales; a single preocular present, higher than postocular; postocular one, higher than wide, not as high as eye diameter; eye diameter 1.4 mm, larger than eye-mouth distance 0.8 mm; distance from anterior corner of eye to nostril 1.9 mm and to the tip of snout 3.1 mm; pupil rounded; supralabials 4/4 (left/right, hereafter), second and third entering orbit, fourth longest, third about 0.9 times of second in length, first shorter than third and as wide as second; mental semicircular to triangular, not touching anterior chin shields; infralabials 5/5, first three touching anterior chin shields; first pair of chin shields meeting in midline, second pair touching anteriorly and separated posteriorly by first gular scales; three gulars in midline between second chin shields and first ventrals; dorsal scales in 13 rows throughout body, reducing to eight rows above 3rd subcaudal, to seven rows above 8th subcaudal, to six rows above 10th subcaudal, and to five rows above the last subcaudal on tail; ventral scales 170, subcaudals 22, divided, followed by a shield covering tail tip; anal scale single.</paragraph>
|
||
<paragraph id="20C4504F41A32B419DFF4A78C4D34494" pageId="0" pageNumber="433">Head length 7.1 mm; head width 5.1 mm; SVL 281.5 mm; TaL 21.7 mm; TL 303.2 mm; body thickness about 5.8 to 6.4 mm; body thickness index at mid body 0.021; tail root 4.5 mm thick; tail not as thick as body, slightly tapering then abruptly tapering at tip; TaL/TL ratio 7.2%; maxillary teeth modified, 10/10.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFDD0CBBD00192E5BD3601FA4F577EC1" pageId="0" pageNumber="433" type="coloration of the holotype in life">
|
||
<paragraph id="093CF8632A5D821987A5676ECDA58122" pageId="0" pageNumber="433">Coloration of the holotype in life.</paragraph>
|
||
<paragraph id="7632B907D5FCCFA7183223C94CE96A65" pageId="0" pageNumber="433">Dorsal surface of body grey-brown above, somewhat iridescent; dark collar at nuchal region absent; indistinct light ring behind head present, nearly invisible; a narrow interrupted light ring at root of tail present, followed by few light spots on middle and tip of tail. Underside of head orangish-beige with black flecks on the infralabials; ventral scales of body and tail immaculate orangish-red, without dark pigmentation on outermost corners and anterior margins; underside of tail without dark longitudinal line or blotches.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2C3942C0024FF72A8536CBB2BDB673A5" pageId="0" pageNumber="433" type="coloration in preservation">
|
||
<paragraph id="BEB7303CFEF454203E5DFBED0040CEFB" pageId="0" pageNumber="433">Coloration in preservation.</paragraph>
|
||
<paragraph id="6B1CCA93871E7234EB7E6A13A4880B4B" pageId="0" pageNumber="433">Color preserved in ethanol fade to caramel-brown above. Dorsal head and upper parts of supralabials brown, lower parts of supralabials light yellow; underside of head light yellow with brownish spots on the infralabials; presence of an indistinct narrow yellow ring about 5 scales wide on neck, nearly invisible; dorsum of body caramel-brown, with light flecks on each dorsal scale; two outermost dorsal scale rows light yellow with upper margins partly dark pigmented; a narrow interrupted light ring at root of tail, light spots on middle and tip of tail still visible; ventral scales of body and tail uniformly light yellowish-beige; underside of tail without dark median line or blotches.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FB8530EDB47AA6608370EE6BC07CD1D" pageId="0" pageNumber="433" type="comparisons">
|
||
<paragraph id="8EE459691566D6EE3DB00C18819E2276" pageId="0" pageNumber="433">Comparisons.</paragraph>
|
||
<paragraph id="6950E969BF018204A8E9B356ABD9B5FF" pageId="0" pageNumber="433">
|
||
We compared
|
||
<taxonomicName id="14A65F1046C23EA510A82FFCC21301A0" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="9BA222D173CA95DFD15320DBF4CDABF5" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="292FE610475AC5C713F6005C66A72E9F" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
with 18 congeners of the genus
|
||
<taxonomicName id="962F8B3AC282ABD75C5127D11A2E2A7E" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria" order="Squamata" pageId="0" pageNumber="433" rank="genus">
|
||
<emphasis id="582A709BBE3A59E8A86A3555876BEF68" italics="true" pageId="0" pageNumber="433">Calamaria</emphasis>
|
||
</taxonomicName>
|
||
from China and mainland Southeast Asia listed above. Unrelated taxa from the Greater Sunda Islands are omitted from comparisons for the sake of simplicity.
|
||
</paragraph>
|
||
<paragraph id="20417BAFCF7EFC3A16402DB31C8D8486" pageId="0" pageNumber="433">
|
||
<taxonomicName id="BCA5CF7358E5EF4217E6A87899E6E1DD" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="306F5762CE7B76A672AAC6C8D09BC385" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="833768D06A799D2D0B1D96680626BD97" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
can be readily distinguished from
|
||
<taxonomicName id="5823F760EC102EE8A47192B0B5EC3DD7" class="Reptilia" kingdom="Animalia" lsidName="C. albiventer" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="albiventer">
|
||
<emphasis id="F4AD0CA83246E05A3D91BBDB5BD7E2CA" italics="true" pageId="0" pageNumber="433">C. albiventer</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="72F7F5085A3C6BEB0A640EC6EB7CF9AD" class="Reptilia" kingdom="Animalia" lsidName="C. lumbricoidea" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="lumbricoidea">
|
||
<emphasis id="0AAF8836A9BC453DE0AAE908B9EA8312" italics="true" pageId="0" pageNumber="433">C. lumbricoidea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A020635FD3A8CBE390AD89963EB2ACAB" class="Reptilia" kingdom="Animalia" lsidName="C. prakkei" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="prakkei">
|
||
<emphasis id="C0CD38E5ADAC424212DC2785FB0F4D43" italics="true" pageId="0" pageNumber="433">C. prakkei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FF63BDA55FF05E6D164A21F3E3FBE0D8" class="Reptilia" kingdom="Animalia" lsidName="C. schlegeli" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="schlegeli">
|
||
<emphasis id="43BE63B73136E01D8C99373D314586EC" italics="true" pageId="0" pageNumber="433">C. schlegeli</emphasis>
|
||
</taxonomicName>
|
||
by having lower number of supralabials (four vs. five), and the pattern of supralabials shields touching the orbit (2nd and 3rd supralabials touching orbit vs. 3rd and 4th supralabials touching orbit). Additionally, these four species only occur south of the Isthmus of Kra in Peninsular Malaysia where is extremely far from the type locality of
|
||
<taxonomicName id="A4B166426FF32259DBA5686BABA6B0BC" class="Reptilia" kingdom="Animalia" lsidName="C. arcana" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="arcana">
|
||
<emphasis id="14A809B89A9B4B011A9AD974E4A7FFE2" italics="true" pageId="0" pageNumber="433">C. arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7F8A6D8AEA84AC350546ABA0B72AA81" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="B841ADDB0C82A0054A4C857D313165DB" pageId="0" pageNumber="433">
|
||
<taxonomicName id="DDD2AB29F9616EA1B7F849031BFB9397" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="1E5B2B33837BF1327C901DBA102AA23D" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8A484063F6F34EAC68FBD9A9B66F817C" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
can be further easily distinguished with
|
||
<taxonomicName id="540CCC71B2309219F45817D89E2C1B57" class="Reptilia" kingdom="Animalia" lsidName="C. lovii" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="lovii">
|
||
<emphasis id="65C5E73DA5D9E3C7D4C7A1DA9D2BCF6B" italics="true" pageId="0" pageNumber="433">C. lovii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91C0D68220C1CAAB778BACCB0BA4CB3B" class="Reptilia" kingdom="Animalia" lsidName="C. nebulosa" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="01E66DB751043B2FFD55D76E0646BE12" italics="true" pageId="0" pageNumber="433">C. nebulosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8C718F8679F9F956143EFDE7FAAB2FA3" class="Reptilia" kingdom="Animalia" lsidName="C. thanhi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="thanhi">
|
||
<emphasis id="4465CB2A50A11E68C939CC4063033CC0" italics="true" pageId="0" pageNumber="433">C. thanhi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="86BE1B143D9CC51023C2B86097CB4E6D" class="Reptilia" kingdom="Animalia" lsidName="C. yunnanensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="yunnanensis">
|
||
<emphasis id="400BF894533E6CABB5D18AFF5BF70C51" italics="true" pageId="0" pageNumber="433">C. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a preocular scale (vs. absence of preocular scale).
|
||
</paragraph>
|
||
<paragraph id="8BD73FF4DE5BAEBD897FA23F2D59614A" pageId="0" pageNumber="433">
|
||
<taxonomicName id="80022F0239D0EF85FD68D9772D372F78" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="48AA358D0F77EE3A9DFFB5669EE95E87" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B17729F00DC855C3AEF53B0F6E58DF36" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="0EABF6BD54F3122BAA7C2F5DC14F9852" class="Reptilia" kingdom="Animalia" lsidName="C. abramovi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="abramovi">
|
||
<emphasis id="E03587BC1CD9D1DC3522404911AA2450" italics="true" pageId="0" pageNumber="433">C. abramovi</emphasis>
|
||
</taxonomicName>
|
||
by having eye diameter larger than distance from eye to mouth edge (vs. reverse condition in
|
||
<taxonomicName id="9BC63DACC0F17AF9C5F290ECE70356F2" class="Reptilia" kingdom="Animalia" lsidName="C. abramovi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="abramovi">
|
||
<emphasis id="E89C7F63723F2145FCE150D1CF288D6E" italics="true" pageId="0" pageNumber="433">C. abramovi</emphasis>
|
||
</taxonomicName>
|
||
), posterior chin shields meeting in midline (vs. separated in
|
||
<taxonomicName id="0DFF7E8AEE3E66941200F8CD87A2259C" class="Reptilia" kingdom="Animalia" lsidName="C. abramovi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="abramovi">
|
||
<emphasis id="A58159DBFF1B3C72652DE6C7B6D3E52A" italics="true" pageId="0" pageNumber="433">C. abramovi</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="0AB673C0BBDDD9FBB53B4D2BE69020D3" class="Reptilia" kingdom="Animalia" lsidName="C. abramovi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="abramovi">
|
||
<emphasis id="7B78D6FA2E1D327DA7F3AAC4A8E214F3" italics="true" pageId="0" pageNumber="433">C. abramovi</emphasis>
|
||
</taxonomicName>
|
||
), and a distinctly different coloration (body black and with yellow-orange spots on venter in
|
||
<taxonomicName id="AEC1FAE60B01943518D08DB81026C57B" class="Reptilia" kingdom="Animalia" lsidName="C. abramovi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="abramovi">
|
||
<emphasis id="AE7D14967D2F3045C747C37972E416CC" italics="true" pageId="0" pageNumber="433">C. abramovi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E08E19B082346D3111407DD1B137FD0" pageId="0" pageNumber="433">
|
||
<taxonomicName id="AEF11DB81AAB7B50356402963FF748F8" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="53C4F81434F67EF3697F5A2484408996" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0A894A35BE0E18301D94998DEDC3FE0A" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="A62943A55CFD1C2497A37C1878E3414C" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="4DD0CFDFCDA5CA93C1ED084BDAB6DE8A" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
by having dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="8BA6E57B8242E51F1EA3FF5DB2D9E70F" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="F69803B9A80C31DF83F9E14373273F7A" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
), light color spots at root of tail present (vs. absent in
|
||
<taxonomicName id="C3D92523CFB05DE8B269120DBB0FD240" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="BC7DE4941C06BB00B982522C3B61D164" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
. Fig.
|
||
<figureCitation id="488662FFD87D8605B702563A2DCB2FA5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6</figureCitation>
|
||
), ventral scales immaculate, without dark outermost corners (vs. presence of dark outermost corners on ventral scales in
|
||
<taxonomicName id="38BA940C45C788DD14890B38381EB163" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="784CCBDDA8C6B14E70EE0B5B0B2090A4" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="5827A38ADC062B27E27D7BC86A7ECA27" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6</figureCitation>
|
||
), two outermost dorsal scale rows light yellow with upper margins partly dark pigmented (vs. all dorsal scale rows in same coloration in
|
||
<taxonomicName id="B99120D8F1A755B2C90630BF80D7C1F4" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="6CF591DDD685C3019B72385D56BC35AD" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="EE74198AF4F59BD0CD750439CF8AC012" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B0759428141AE5DE79A69DC33BFF961D" pageId="0" pageNumber="433">
|
||
<taxonomicName id="3AD3EC614D80D269416451376163ADA5" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="3DC8D75015EACD6710711FF9CCBB1780" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1C4243F97D8F869F0AA505B99922BD08" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="1F12B8647DA16C710D149A1BBE40564E" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="FDACD9E0F67B25C589368F49356134A5" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
by having rostral shield width larger than length (vs. reverse condition in
|
||
<taxonomicName id="06B9F5E29F10CCAA318EC3EC4ADA92DE" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="50AB09A0A4F06EC034FE757E70B0E450" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
), eye diameter larger than distance from eye to mouth edge (vs. equal to or shorter in
|
||
<taxonomicName id="9DEC74FEB2F1F6217F52C56FE69DE900" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="F2CC7801769A11420AC7C9102591CC4A" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
), mental not touching anterior chin shields (vs. touching in
|
||
<taxonomicName id="DA03CD0937B829A7114EF036AC173284" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="D6C26BEF75B01C072ECB05365158F0EB" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales reduced to five rows at tail (reduced to four rows in
|
||
<taxonomicName id="902A29522363C233E5BE2DA280443FE9" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="6ED59EBC394ED1BD12EAEE14D9770485" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
), and six shields and scales surrounding paraparietal (vs. five in
|
||
<taxonomicName id="37784A358423593E67E6BF1BD3CFA0E4" class="Reptilia" kingdom="Animalia" lsidName="C. buchi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="buchi">
|
||
<emphasis id="EDB7B4E702E46B25AB2784ABD58AE9A6" italics="true" pageId="0" pageNumber="433">C. buchi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC6187F9D89B92396720DBD1A65D7E57" pageId="0" pageNumber="433">
|
||
<taxonomicName id="F13DEC72ABDDD40F3FD78CD5FA8C4179" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="C0077732BE06C42B5ECCFF3ED161D5DF" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FF13E1FCE5A26AA5C0F4A40D78E96A79" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="8CD8B9CE10336942DA5AE8C9D5F4C9BC" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="EA438485606B4E32253506998F9FAD28" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
by having lower number of supralabials (4 vs. 5 in
|
||
<taxonomicName id="04E0B6DE7BA3BC51574EE45360707976" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="C19CC91F70F7D722A674B72C822CFDD8" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
), eye diameter larger than distance from eye to mouth edge (vs. reverse condition in
|
||
<taxonomicName id="6248FDE614BAAD92D0DF7F68F6F040DF" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="6E7C1F1D8FE9E8B49BA5C020BF4CCB62" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
), mental not touching anterior chin shields (vs. touching in
|
||
<taxonomicName id="D5B302EBCD914CE38049B2B28F6FB28A" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="20966EF3EE22516CE8F95B79F5D8AB21" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
), tail not flatted (vs. slight flatted laterally in
|
||
<taxonomicName id="F73900570FA7768500EEA0822CDE8EEA" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="29705BFF74E5191E63BEF62244F47CBA" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="1600378A660702FDBF890C85C1EAD054" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="9ABF8CD73CC15E93BC536B58F7923FDC" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
), and lower number of ventral scales in males (170 vs. 209 in
|
||
<taxonomicName id="5CD6E26A84748FF68E6B947340C7E001" class="Reptilia" kingdom="Animalia" lsidName="C. concolor" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="concolor">
|
||
<emphasis id="19C4849CC179AE2AED2AE123CD07F301" italics="true" pageId="0" pageNumber="433">C. concolor</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="67A18D5052F15179AEBCC5C54E89FB30" pageId="0" pageNumber="433">
|
||
<taxonomicName id="94102A95F509FA16D9E1C33C6132A38A" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="4BCD55FD2B966224DEB429C657FC7411" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="06CFD5EF6A1FE0231E8D4C675C2A968D" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="8DD17C4528EC32FAF5C6B46B653C1851" class="Reptilia" kingdom="Animalia" lsidName="C. dominici" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="dominici">
|
||
<emphasis id="94BDCA807752A34211F2CD473751FAA4" italics="true" pageId="0" pageNumber="433">C. dominici</emphasis>
|
||
</taxonomicName>
|
||
by having dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="4A9A27F93CD6EF40FFF523B116F4B766" class="Reptilia" kingdom="Animalia" lsidName="C. dominici" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="dominici">
|
||
<emphasis id="0CAC5D06490DC8DD9B082E4FBC0666A9" italics="true" pageId="0" pageNumber="433">C. dominici</emphasis>
|
||
</taxonomicName>
|
||
), absence of distinct yellow blotches on dorsum (vs. present in
|
||
<taxonomicName id="DB729C8BD2AC14C3C1E0376E1FBCC887" class="Reptilia" kingdom="Animalia" lsidName="C. dominici" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="dominici">
|
||
<emphasis id="749161E7F16F73CC4F3C1CB1BF82C99A" italics="true" pageId="0" pageNumber="433">C. dominici</emphasis>
|
||
</taxonomicName>
|
||
), venter immaculate without any dark stripes or spots (vs. ventral side dark with yellow blotches and bands).
|
||
</paragraph>
|
||
<paragraph id="0E7A189082C9A1E4B6742BEBE5A8C64F" pageId="0" pageNumber="433">
|
||
<taxonomicName id="AA067BEF7382AA404F755E90A6C66E28" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="1EF36192A12544780A8905BE4B5F3D8B" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CD2B9BC7792237977FE1B7B25CABBF16" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="C3DBE7543832B2326203E6534CB400F2" class="Reptilia" kingdom="Animalia" lsidName="C. gialaiensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="gialaiensis">
|
||
<emphasis id="0AF48D30E543FF152A47EE7A82BB1ACB" italics="true" pageId="0" pageNumber="433">C. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
by having mental not touching anterior chin shields (vs. touching in
|
||
<taxonomicName id="6D7AD283E91B247A6F3BDF23836A7674" class="Reptilia" kingdom="Animalia" lsidName="C. gialaiensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="gialaiensis">
|
||
<emphasis id="C71E4A94DD3EBFFB922112D09BD684DE" italics="true" pageId="0" pageNumber="433">C. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
), lower number of ventral scales (170 in males vs. 191 in unknown gender), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="4F365CA5CD6A9AA661320D5AC2CDDCB0" class="Reptilia" kingdom="Animalia" lsidName="C. gialaiensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="gialaiensis">
|
||
<emphasis id="836C2091C53B0B0CD6EA97ABE9F11522" italics="true" pageId="0" pageNumber="433">C. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
), absence of a dark longitudinal line under of tail (vs. present in
|
||
<taxonomicName id="9184F753261A8A6C26861D7288C31138" class="Reptilia" kingdom="Animalia" lsidName="C. gialaiensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="gialaiensis">
|
||
<emphasis id="1CD5FC620C5FCD4D7B5BC765F9A6D5B6" italics="true" pageId="0" pageNumber="433">C. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
), and absence of dark collar in nuchal region (vs. an indistinct dark collar present in nuchal region in
|
||
<taxonomicName id="B51F0FF9A5AD425FFBE5C7929A66399A" class="Reptilia" kingdom="Animalia" lsidName="C. gialaiensis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="gialaiensis">
|
||
<emphasis id="C2B460517A104C4F5AFD320343D7B3DA" italics="true" pageId="0" pageNumber="433">C. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="462A090C3DE228CF551E15AD1240CAB3" pageId="0" pageNumber="433">
|
||
<taxonomicName id="FC9D81D98D4FADCB5C12114429CA12AA" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="B3A24D61D356DED1BC4C144B2FD1E461" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7522FE474B2157B1D63FE240CCE7D314" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="906BAEA5140111328ECF06B06F722FFA" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="DF05AF06AE18BBCA4991A32BF28F5592" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
by having mental not touching anterior chin shields (vs. touching in
|
||
<taxonomicName id="9E9C50EE6C3DAF416F0AABA1DECE14FC" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="EFFADEF9671BF4A819FEF2FD41152320" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="8597FB97BEB7D4A8A0367477A05C7CFD" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="B3D70116F18E0AEE3B7FBE44AC6EF0BF" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
), lower number of ventral scales (170 vs. 190 in
|
||
<taxonomicName id="E868F65C4773CB4B0D89A76F9526E5C0" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="4DAA11724155798ED747770A79CEC986" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
), absence of dark collar in nuchal region (vs. presence of dark collar in nuchal region in
|
||
<taxonomicName id="0DAC28DC6EEAD488072646E0BE6E606E" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="22144BAE54658FFF880E91CC3894BA7A" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
), and venter immaculate (vs. venter cream with dark transverse bands in
|
||
<taxonomicName id="58A706AA79B9B09AF2D66C6AFDF7ED1C" class="Reptilia" kingdom="Animalia" lsidName="C. sangi" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="sangi">
|
||
<emphasis id="EF1F2EC07F343765D58CB46F2B5487D3" italics="true" pageId="0" pageNumber="433">C. sangi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D91CD1CB361FF0B1F4F3462CFDE52B3A" pageId="0" pageNumber="433">
|
||
<taxonomicName id="D7421C1A318A4DD0E42C24F20D91F850" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="5764EC6455457EF7E1C4252AC34EBF48" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E5C76DBE016B77B48B314EE92614EB5E" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="B889B2266374FADCE75627FFA2F71437" class="Reptilia" kingdom="Animalia" lsidName="C. strigiventris" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="strigiventris">
|
||
<emphasis id="8B2B1DBAEF8A5FDA38FE4D065294B30B" italics="true" pageId="0" pageNumber="433">C. strigiventris</emphasis>
|
||
</taxonomicName>
|
||
by having eye diameter larger than distance from eye to mouth edge (vs. reverse condition in
|
||
<taxonomicName id="0D222C53F1875E33D8CC6B01F681021A" class="Reptilia" kingdom="Animalia" lsidName="C. strigiventris" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="strigiventris">
|
||
<emphasis id="40CC5BEFCD60E3D796EA318D1063F39B" italics="true" pageId="0" pageNumber="433">C. strigiventris</emphasis>
|
||
</taxonomicName>
|
||
), lower number of subcaudals in males (22 vs. 29-33 in males), higher number of ventrals in males (170 vs. 130-157), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="E91F22F2C0333E03BBE250913D7B54A0" class="Reptilia" kingdom="Animalia" lsidName="C. strigiventris" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="strigiventris">
|
||
<emphasis id="E7C932D534EEC13E313DD7767B74A1F4" italics="true" pageId="0" pageNumber="433">C. strigiventris</emphasis>
|
||
</taxonomicName>
|
||
), venter immaculate (vs. presence of three interrupted longitudinal black stripes in
|
||
<taxonomicName id="3B89DBD141BB046164E08032E78360C5" class="Reptilia" kingdom="Animalia" lsidName="C. strigiventris" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="strigiventris">
|
||
<emphasis id="EFBE26C84D36384EC8C2E52745851306" italics="true" pageId="0" pageNumber="433">C. strigiventris</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C89F66B1234F89AB97BEB7F02B30487E" pageId="0" pageNumber="433">
|
||
<taxonomicName id="C87404E5016CB303EC794BB3D559818B" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="4B18618655303B9D004F89D78F7BA499" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FCD4EF043FA26120E93038941D50A645" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="E55A700B8E0E439FBD32532FDCFFCAA2" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="B5061FE48EB628E4F0C4C4B1652027C5" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
by having rostral shield width larger than length (vs. reverse condition in
|
||
<taxonomicName id="AC4467CAFC5E4BFEFEC6900190DA4884" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="1AD7466D5376AAE200D99A52970F3E75" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
), tail slowly tapering anteriorly and abruptly tapering at tip (vs. tail tapering gradually to a point in
|
||
<taxonomicName id="5B469ADCBC5355BDC9DC7E0016BF1405" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="61F3870F5BDAF91A39A5526BEAFDDEE9" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="7C2C3FB88D1D97C8BD52AA9EF1667575" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6</figureCitation>
|
||
), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="4DDF3CA9503FA1BB5025C85A58F40FF3" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="BFA01B07B7C969C3585757DBCF09D6C4" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
), absence of dark collar at nuchal region (vs. present in
|
||
<taxonomicName id="A59C4CDB2E8312E5D70CEE9D319B0A12" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="D66E04DEC6B6866D087903E5340B7161" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="D6CC28D4D7D3ABEBAE9BFC41A35B2BFF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">5</figureCitation>
|
||
), absence of distinct narrow and dark longitudinal lines on dorsum (vs. present in
|
||
<taxonomicName id="5D4CE815C9E3983827E817746B941C38" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="7E884DD0C6A67160FE89842F98CC06C2" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="2FD90E6AAF244BD853D219BCA13E9CE9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">5</figureCitation>
|
||
), absence of dark outermost corners on ventral scales (vs. ventral scales with dark outermost corners in
|
||
<taxonomicName id="02CF553A036757C3CA403EBB1A017F9D" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="CC39A8DED422D916E46F0AB1D5C39109" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="F268AC80E2FF371E6CB3D9AC137E7B5E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="44024C6D81328010BF9BCAD49326A79A" pageId="0" pageNumber="433">
|
||
<taxonomicName id="B1506493A26F638819DA3F7A74C488BF" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="FE13DA6F42E82EC35C2D463E96FA74A8" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CD5EE3F5E7FC77FA420B8BCF5B330BC" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
can be distinguished from the sister taxon of
|
||
<taxonomicName id="26ED96DA8C7239D25A0A2DFC8EE1FB41" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="D4BE885C8F4A901F6137E86634E2FA0F" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
by having tail abruptly tapering at the tip of end (vs. tail not tapering in
|
||
<taxonomicName id="5C589D12A5B0E147A87DB34AD8CCB914" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="BAD313E15D6DC5B0CE8CE4A4C66E0A9A" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), higher number of maxillary teeth (10 vs. 8-9 in
|
||
<taxonomicName id="4760E3480E33103A94E310A95D5EA9FB" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="2ABE6968BEE76FBB8C2021EEDDF6AAAA" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), higher number of ventral scales in males (170 vs. 148-166 in
|
||
<taxonomicName id="82A7ED0815C681C0C0027E1EEBE8E213" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="2DEDAF408DA14603325CFA274D6E24E0" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), higher numbers of subcaudals in males (22 vs. 15-19 in
|
||
<taxonomicName id="8087E937377D2D60DD4A1D87EECF80AC" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="2B655114874C8B2EFEC937C854B564FA" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), dorsal scales reduced to five rows at tail (vs. reduced to four rows in
|
||
<taxonomicName id="4CAA7CA585DF10FBB4069C54305DF650" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="B822923D4649446E100B3B57FF24C795" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), absence of black line or spots on venter of tail (vs. venter of tail with broad and distinct median black stripe), absence of dark outermost corners on ventral scales (vs. present in
|
||
<taxonomicName id="22A83D9D740E93AD459F9634CC8573BD" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="9C6CB113F78FEA1A26D5C1D40E11F2EA" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="8E321B434D9E11596BEACC2DF0603899" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the tail shape and coloration among Calamaria arcana sp. nov., C. andersoni, C. septentrionalis and C. pavimentata. A-C dorsal, lateral and ventral views of the tail of C. pavimentata (KFBG 14507). D-F dorsal, lateral and ventral views of the tail of C. septentrionalis (KFBG 14506). G-I dorsal, lateral and ventral views of the tail of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal, lateral and ventral views of the tail of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433">6</figureCitation>
|
||
), two outermost dorsal scale rows light yellow with upper margins partly dark pigmented (vs. all dorsal scale rows in same coloration in
|
||
<taxonomicName id="30FB35D86513B7C66D6985609AD13BB6" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="599F60797BF201AC98CE38C9CFAF01EE" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="24CC15352166A12FA9D9A85A0286C31A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparative characters of the head shape and body coloration of Calamaria arcana sp. nov., C. andersoni, C. septentrionalis, C. pavimentata. A-C dorsal view of head and body, and dorsolateral view of body of C. pavimentata (KFBG 14507). D-F dorsal view of head and body, and dorsolateral view of body of C. septentrionalis (KFBG 14506). G-I dorsal view of head and body, and dorsolateral view of body of holotype of C. arcana sp. nov. (KFBG 14611). J-L dorsal view of head and body, and dorsolateral view of body of C. andersoni (SYS r 001699)." figureDoi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="236DE559A3D502E5BA92632556FB50A9" pageId="0" pageNumber="433" type="distribution">
|
||
<paragraph id="04EEEC187D7248852014E4AAB39DD275" pageId="0" pageNumber="433">Natural history, distribution and conservation status.</paragraph>
|
||
<paragraph id="675325E1AA1591E8F314178376A2BBEE" pageId="0" pageNumber="433">The holotype was found active on a cloudy morning along a trail in tall shrubland and secondary forest.</paragraph>
|
||
<paragraph id="A92715FED25BE04AFFD0A58662BBDCB5" pageId="0" pageNumber="433">
|
||
<taxonomicName id="FB7A00D4069AD6BDCD4EAAE7AB3183CB" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="039DB7A5A3C503FAD6F8B6701845D30A" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="36F9B47D4B5192C0D5CBDC03FD084316" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
is currently only known from a single specimen from its type locality, Mt. Dadongshan, Nanling National Nature Reserve, Guangdong Province, China (Fig.
|
||
<figureCitation id="1CD4A2F4E1DC04D7973BE9B840970221" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Map of Southern China showing the type locality of Calamaria arcana sp. nov.; Mt. Dadongshan, Lianzhou County, Guangdong Province, China (star)." figureDoi="10.3897/vz.72.e84516.figure7" httpUri="https://binary.pensoft.net/fig/706626" pageId="0" pageNumber="433">7</figureCitation>
|
||
). It can be expected that the new species should be distributed across adjacent mountain areas, but in view of the uncertainty of its extent of occurrence, and little information on its ecology and threats, we recommend the new species be listed as Data Deficient (DD) according the International Union for Conservation of Nature (IUCN) Red List categories and Criteria (
|
||
<bibRefCitation id="B875C5026F9E2437AAB4ED5B89577BF2" author="Inger, RF" journalOrPublisher="Fieldiana Zoology" pageId="0" pageNumber="433" publicationUrl="https://www.iucnredlist.org" refId="B7" refString="2022. IUCN International Union for Conservation of Nature. https://www.iucnredlist.org [ [Accessed 16.01.2022] ]" title="IUCN International Union for Conservation of Nature" url="https://www.iucnredlist.org" year="2022">IUCN 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="7F0B38C4E6655074F30B867193F3EBB2" doi="10.3897/vz.72.e84516.figure5" httpUri="https://binary.pensoft.net/fig/706624" pageId="0" pageNumber="433" start="Figure 5" startId="F5">
|
||
<paragraph id="FE8645D6ED177F4C4EFD01FF3AF2DA55" pageId="0" pageNumber="433">
|
||
<emphasis id="8D97E275747434640E8CE30F64DE82DF" bold="true" pageId="0" pageNumber="433">Figure 5.</emphasis>
|
||
Comparative characters of the head shape and body coloration of
|
||
<taxonomicName id="3A854D5FBCC07DB1513E517D6CA2918D" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="1F37399925E770A00CB169C1DB96A508" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F833D35CCE4C39CDF40E118631EB1A1" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
,
|
||
<taxonomicName id="E83050C44DE1B903F2CE8FDD2BF6641E" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="83DEC60B2F07636E57F62158EB9EBAA1" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5E926D54A460BF238FB5DA8DE4800962" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="9B302A939DD58136ED7ED84D799F5C5C" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F8FFBBA5586EAE013BC9805F1D2905A8" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="AD73DC4EAF2F6FEB2F54B09C1321D9D5" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="BC8C08C24057ED7BE65024BC23317B6C" bold="true" pageId="0" pageNumber="433">A-C</emphasis>
|
||
dorsal view of head and body, and dorsolateral view of body of
|
||
<taxonomicName id="F8D65E8EA582E6418FBB24E6829290DC" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="79573196A9B2496FE94AB71F1BF03E0B" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
(KFBG 14507).
|
||
<emphasis id="2BB4E4163ED0289127250716B466BCDC" bold="true" pageId="0" pageNumber="433">D-F</emphasis>
|
||
dorsal view of head and body, and dorsolateral view of body of
|
||
<taxonomicName id="D76B2B089A8747AD790C24A60698989D" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="1572C89ACF7131689B8ED0207E2BC978" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
(KFBG 14506).
|
||
<emphasis id="B12F17BADDFF39FD84BBA88BD30FDCE8" bold="true" pageId="0" pageNumber="433">G-I</emphasis>
|
||
dorsal view of head and body, and dorsolateral view of body of holotype of
|
||
<taxonomicName id="44915D914A6C30FAB5EABA3FEF253561" class="Reptilia" kingdom="Animalia" lsidName="C. arcana" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="arcana">
|
||
<emphasis id="B62FD95FED10F16334DB5FC00F037717" italics="true" pageId="0" pageNumber="433">C. arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5C2110A4D473A6DC98AF48289F003C84" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
(KFBG 14611).
|
||
<emphasis id="E1D24AF34F25E546E53D7EA697B8D884" bold="true" pageId="0" pageNumber="433">J-L</emphasis>
|
||
dorsal view of head and body, and dorsolateral view of body of
|
||
<taxonomicName id="59FE6AD10A4A93E76BCDC90ACF5BF8B1" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="2D104566742D341E4E552B1B4411E6C2" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
(SYS r001699).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3BF181BC6446C6DB656455A1C49328C6" doi="10.3897/vz.72.e84516.figure6" httpUri="https://binary.pensoft.net/fig/706625" pageId="0" pageNumber="433" start="Figure 6" startId="F6">
|
||
<paragraph id="24B17F5C1655386C472AF06CBFE1DFD5" pageId="0" pageNumber="433">
|
||
<emphasis id="FE8EF94001B0A94EA615C3602CAC0022" bold="true" pageId="0" pageNumber="433">Figure 6.</emphasis>
|
||
Comparison of the tail shape and coloration among
|
||
<taxonomicName id="01F2F5258BD7A08235DEE7A6A242902F" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="6ADD03DA0E9FEC64B3195657434CC877" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AC1E45D63132900C762DFB493F58F04F" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
,
|
||
<taxonomicName id="E1DFB15FFC3692FE2432E23898C3A691" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="C6218D42E4F4C232297398CF190F4ED6" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3609EE35E8CF5F49090F7EC9FCDCEE1" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="68BB49047CD3738C42F118065BFE65D4" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="243A01A66B17BE4F0358CEB66636FFC9" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="9646635BC4E4E30147DCFF7384A70A7A" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E42A40A50D4E2885F9B14A3F33939EA3" bold="true" pageId="0" pageNumber="433">A-C</emphasis>
|
||
dorsal, lateral and ventral views of the tail of
|
||
<taxonomicName id="D15317D319CD6BA552D78D1C775D5C61" class="Reptilia" kingdom="Animalia" lsidName="C. pavimentata" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="pavimentata">
|
||
<emphasis id="D34D550CAB91F2ED8636F83FDDFFD634" italics="true" pageId="0" pageNumber="433">C. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
(KFBG 14507).
|
||
<emphasis id="2CB6FA98A5BCB8C3202EB046D5AEF6D3" bold="true" pageId="0" pageNumber="433">D-F</emphasis>
|
||
dorsal, lateral and ventral views of the tail of
|
||
<taxonomicName id="8E3BE09BA765627A90007C93DA01657D" class="Reptilia" kingdom="Animalia" lsidName="C. septentrionalis" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="BD83DFBC60304D4D4071C54B1FC99F48" italics="true" pageId="0" pageNumber="433">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
(KFBG 14506).
|
||
<emphasis id="81F3D60F615839C504373FA62279FFA0" bold="true" pageId="0" pageNumber="433">G-I</emphasis>
|
||
dorsal, lateral and ventral views of the tail of holotype of
|
||
<taxonomicName id="18254BD933A6BF60280E8F0788ECBD22" class="Reptilia" kingdom="Animalia" lsidName="C. arcana" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="arcana">
|
||
<emphasis id="DA9FE7B16A5923C4CAA06D2469AD386C" italics="true" pageId="0" pageNumber="433">C. arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1C9622977F3A421BA7712F249481189" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
(KFBG 14611).
|
||
<emphasis id="9DFFDF41C3D0DA80D972F1AC891D4093" bold="true" pageId="0" pageNumber="433">J-L</emphasis>
|
||
dorsal, lateral and ventral views of the tail of
|
||
<taxonomicName id="A9C79ABFE1CFAB875F87167E21BF5C7E" class="Reptilia" kingdom="Animalia" lsidName="C. andersoni" order="Squamata" pageId="0" pageNumber="433" phylum="Chordata" rank="species" species="andersoni">
|
||
<emphasis id="36A2FD8C8DF90670774543F548050FE5" italics="true" pageId="0" pageNumber="433">C. andersoni</emphasis>
|
||
</taxonomicName>
|
||
(SYS r001699).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C4D7745762986F97F36337C59C9DB19D" doi="10.3897/vz.72.e84516.figure7" httpUri="https://binary.pensoft.net/fig/706626" pageId="0" pageNumber="433" start="Figure 7" startId="F7">
|
||
<paragraph id="D4A5EAB61FDE65C5A31E2BBBC7CA5D3E" pageId="0" pageNumber="433">
|
||
<emphasis id="57A081CD3BD38C086AD575895C0E76E9" bold="true" pageId="0" pageNumber="433">Figure 7.</emphasis>
|
||
Map of Southern China showing the type locality of
|
||
<taxonomicName id="FF8DF0E3299E39BE8BC028B2EECF5874" authorityName="Yeung & Lau & Yang" authorityYear="2022" family="Colubridae" genus="Calamaria" kingdom="Animalia" lsidName="Calamaria arcana" order="Squamata" pageId="0" pageNumber="433" rank="species" species="arcana">
|
||
<emphasis id="DDB009CECC493FE94D2A730C430C0B61" italics="true" pageId="0" pageNumber="433">Calamaria arcana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7E93B7A4EC79893ED8686943CAD8C308" bold="true" pageId="0" pageNumber="433">sp. nov.</emphasis>
|
||
; Mt. Dadongshan, Lianzhou County, Guangdong Province, China (star).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |