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<document ID-DOI="http://dx.doi.org/10.3897/jhr.94.94263" ID-Pensoft-Pub="1314-2607-94-13" ID-Pensoft-UUID="ADCFB8021287566FB2D7E8A8711D5CAE" ID-ZooBank="6CB807239A47403FABEC9AF8AE7F417F" ModsDocID="1314-2607-94-13" checkinTime="1671601504499" checkinUser="pensoft" docAuthor="Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars &amp; Rasplus, Jean-Yves" docDate="2022" docId="59D3A255387452889FD2A8D87D2CFA7E" docLanguage="en" docName="JourHymenoptRes 94: 13-88" docOrigin="Journal of Hymenoptera Research 94" docPubDate="2022-12-20" docSource="http://dx.doi.org/10.3897/jhr.94.94263" docTitle="Eopelma Gibson incertae sedis new placement" docType="treatment" docVersion="1" id="ADCFB8021287566FB2D7E8A8711D5CAE" lastPageNumber="13" masterDocId="ADCFB8021287566FB2D7E8A8711D5CAE" masterDocTitle="From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)" masterLastPageNumber="88" masterPageNumber="13" pageNumber="13" updateTime="1671601504499" updateUser="pensoft">
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<mods:title>From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)</mods:title>
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<mods:namePart>Burks, Roger</mods:namePart>
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<mods:namePart>Fusu, Lucian</mods:namePart>
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<mods:affiliation>Faculty of Biology, Alexandru Ioan Cuza University, Iasi, Romania</mods:affiliation>
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<mods:namePart>Heraty, John M.</mods:namePart>
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<mods:affiliation>Insects Division, Natural History Museum, London, UK</mods:affiliation>
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<mods:namePart>Peters, Ralph S.</mods:namePart>
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<mods:namePart>Tselikh, Ekaterina V.</mods:namePart>
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<mods:namePart>Woolley, James B.</mods:namePart>
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<mods:namePart>van Noort, Simon</mods:namePart>
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<mods:affiliation>Research and Exhibitions Department, South African Museum, Iziko Museums of South Africa, PO Box 61, Cape Town 8000 South Africa &amp; Department of Biological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701, South Africa</mods:affiliation>
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<mods:namePart>Baur, Hannes</mods:namePart>
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<mods:affiliation>Department of Invertebrates, Natural History Museum Bern, Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, Bern, Switzerland</mods:affiliation>
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<mods:namePart>Cruaud, Astrid</mods:namePart>
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<mods:namePart>Darling, Christopher</mods:namePart>
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<mods:namePart>Haas, Michael</mods:namePart>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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<mods:namePart>Hanson, Paul</mods:namePart>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany &amp; Institute of Biology, Biological Systematics (190 n) University of Hohenheim, Stuttgart, Germany</mods:affiliation>
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<mods:namePart>Rasplus, Jean-Yves</mods:namePart>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-20</mods:number>
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<mods:start>13</mods:start>
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<treatment LSID="urn:lsid:plazi:treatment:59D3A255387452889FD2A8D87D2CFA7E" httpUri="http://treatment.plazi.org/id/59D3A255387452889FD2A8D87D2CFA7E" lastPageNumber="13" pageId="0" pageNumber="13">
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<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="59D3A255-3874-5288-9FD2-A8D87D2CFA7E" authority="Gibson incertae sedis new placement" authorityName="Gibson incertae sedis new placement" class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Eopelma Gibson incertae sedis new placement</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="13">
<taxonomicName class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Eopelma</taxonomicName>
Gibson, 1989. Type species.
<taxonomicName authorityName="Gibson" authorityYear="1989" class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma mystax" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mystax">Eopelma mystax</taxonomicName>
Gibson, 1989.
</paragraph>
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<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">
Antenna with 8 flagellomeres, with an undivided clava. Eyes ventrally divergent. Clypeus without transverse subapical groove. Labrum hidden behind clypeus, flexible. Mandibles with 3 teeth or with a ventral tooth and dorsal slightly emarginate truncation. Axilla long, separated anteromedially. Axillular sulcus more or less distinct. Frenum absent. Acropleuron enlarged, convex and pad-like; covering most of mesopleural area, separated from mesocoxa by the lower mesepimeron, metapleuron, and a dorsal extension of the metasternal area. All legs with 5 tarsomeres; protibial spur stout and curved; basitarsal comb longitudinal; ventral membranous area anterior to mesocoxal attachment lacking; mesotibial spur stout; mesotarsus with 1 row of pegs anteroventrally (Fig.
<figureCitation captionStart="Figures 6166" captionStartId="F11" captionText="Figures 61 - 66. 61 Austrosystasis atricorpus Girault (Austrosystasinae, incertae sedis): mesosoma lateral view 62 - 64 Ditropinotella sp. (Ditropinotellinae, incertae sedis) 62 antenna 63 mesosoma lateral view 64 metasoma 65 Enoggera reticulata Naumann (Enoggerinae, incertae sedis): mesosoma lateral view 66 Eopelma sp. (incertae sedis): apex of mesotibia and mesotarsus." figureDoi="10.3897/jhr.94.94263.figures61-66" httpUri="https://binary.pensoft.net/fig/781074" pageId="0" pageNumber="13">66</figureCitation>
). Metasoma with syntergum, therefore without epipygium.
</paragraph>
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<subSubSection pageId="0" pageNumber="13" type="discussion">
<paragraph pageId="0" pageNumber="13">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Eopelma</emphasis>
</taxonomicName>
is consistently the sister group of another
<emphasis italics="true" pageId="0" pageNumber="13">incertae sedis</emphasis>
taxon,
<taxonomicName authorityName="Boucek" authorityYear="1988" class="Insecta" family="Pteromalidae" genus="Storeya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Storeya" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Storeya</emphasis>
</taxonomicName>
<normalizedToken originalValue="Bouček">Boucek</normalizedToken>
(
<taxonomicName family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Storeyinae">Storeyinae</taxonomicName>
), in next-generation molecular analysis (Cruaud et al., submitted). These two groups do not strongly resemble each other in body shape.
<taxonomicName authorityName="Boucek" authorityYear="1988" class="Insecta" family="Pteromalidae" genus="Storeya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Storeya" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Storeya</emphasis>
</taxonomicName>
does not have an expanded acropleuron, and has not previously been indicated as a relative of any eupelmid. They do share an antennal flagellomere count, a long radicle, general coloration, and an undivided clava, but the list of shared features possessed by these two genera is much shorter than the list of differences between them. A new subfamily is not described for
<taxonomicName class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Eopelma</emphasis>
</taxonomicName>
because it is a single genus, the position of which in
<taxonomicName kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="superFamily" superFamily="Chalcidoidea">Chalcidoidea</taxonomicName>
is still in question.
</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName class="Insecta" family="Eupelmidae" genus="Eopelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eopelma" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Eopelma</emphasis>
</taxonomicName>
vaguely resembles
<taxonomicName authorityName="Girault" authorityYear="1913" class="Insecta" family="Neanastatidae" genus="Neanastatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanastatus" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Neanastatus</emphasis>
</taxonomicName>
Girault in body shape and coloration.
<taxonomicName authorityName="Girault" authorityYear="1913" class="Insecta" family="Neanastatidae" genus="Neanastatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neanastatus" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Neanastatus</emphasis>
</taxonomicName>
differs in having a much smaller axilla, which may not be clearly visible.
</paragraph>
</subSubSection>
</treatment>
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