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<document id="6F0CBACBA5E128601EC8A20D3BB42883" ID-DOI="10.5281/zenodo.176054" ID-GBIF-Dataset="8087be13-7a23-4270-beb1-65f12a342f19" ID-ISSN="1175-5326" ID-Zenodo-Dep="176054" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459826746963" checkinUser="plazi" docAuthor="Gaimari, Stephen D." docDate="2007" docId="3449AB7BC760FF9BFF4CFE57572C58A0" docLanguage="en" docName="zt01443p016.pdf" docOrigin="Zootaxa 1443" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Helgreelia parkeri Gaimari, 2007, sp. nov." docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="C870D303C764FF9EFFDBFFA853615A32" masterDocTitle="Three new Neotropical genera of Odiniidae (Diptera: Acalyptratae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="5" updateTime="1698222377076" updateUser="plazi">
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<mods:title id="692AD0FB6ED3CCC50A2D4078122766E4">Three new Neotropical genera of Odiniidae (Diptera: Acalyptratae)</mods:title>
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<mods:namePart id="46C13D10762CB859A6B1BF2C9B6FAEC1">Gaimari, Stephen D.</mods:namePart>
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<mods:typeOfResource id="D983D90AABE2B72E8204D8E751F7619F">text</mods:typeOfResource>
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<mods:title id="D6F29B7DD94AF7947FD73AB634D66A1F">Zootaxa</mods:title>
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<mods:part id="A43E681731065F5F3EDD6FF945DB4445">
<mods:date id="1F60468A977228F086C72D353292030A">2007</mods:date>
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<mods:classification id="A45068A5F33F03E75B8C660C3E322BA8">journal article</mods:classification>
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<treatment id="3449AB7BC760FF9BFF4CFE57572C58A0" ID-DOI="http://doi.org/10.5281/zenodo.6249775" ID-GBIF-Taxon="119356965" ID-Zenodo-Dep="6249775" LSID="urn:lsid:zoobank.org:act:6F09DBD9-A1DA-43B9-8C45-23B730B6D75C" httpUri="http://treatment.plazi.org/id/3449AB7BC760FF9BFF4CFE57572C58A0" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="F4FA49E6C760FF9AFF4CFE5751D6582B" box="[151,695,511,537]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="BC5F1A6DC760FF9AFF4CFE5751D6582B" blockId="4.[151,695,511,537]" box="[151,695,511,537]" pageId="4" pageNumber="5">
<heading id="E717AD01C760FF9AFF4CFE5751D6582B" bold="true" box="[151,695,511,537]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="8E94C67FC760FF9AFF4CFE5751D6582B" bold="true" box="[151,695,511,537]" pageId="4" pageNumber="5">
<taxonomicName id="7BE061EEC760FF9AFF4CFE575214582B" ID-CoL="3K4YF" box="[151,373,511,537]" class="Insecta" family="Odiniidae" genus="Helgreelia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parkeri" status="sp. nov.">
<emphasis id="8E94C67FC760FF9AFF4CFE575214582B" bold="true" box="[151,373,511,537]" italics="true" pageId="4" pageNumber="5">Helgreelia parkeri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="95A77B04C760FF9AFE5FFE575280582B" box="[388,481,511,537]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="24DB06E8C760FF9AFE2AFE575123582B" box="[497,578,511,537]" captionStart="FIGURE 1" captionStartId="10.[151,255,1209,1233]" captionTargetBox="[150,1436,194,1183]" captionTargetId="figure@10.[150,1436,194,1185]" captionTargetPageId="10" captionText="FIGURE 1. Lateral habitus of holotypes of new species (for B D, post-dissection state); inset is a dorsal view of each abdomen. Scale bar = 1.0 mm. A) Helgreelia albeto, sp. nov., Ψ (inset is a paratype). B) Helgreelia parkeri, sp. nov., ♂. C) Neoschildomyia fusca, sp. nov., ♂. D) Pradomyia hadromera, sp. nov., ♂." httpUri="https://zenodo.org/record/176055/files/figure.png" pageId="4" pageNumber="5">Figs. 1</figureCitation>
B, 4AC)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4FA49E6C760FF9AFF4CFDE2502058BE" pageId="4" pageNumber="5" type="etymology">
<paragraph id="BC5F1A6DC760FF9AFF4CFDE2502058BE" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF4CFDE2527A5851" box="[151,283,586,611]" italics="true" pageId="4" pageNumber="5">Etymology.</emphasis>
Named for Dr. Frank Parker, whose collecting efforts in
<collectingCountry id="C4F75AFDC760FF9AFC66FDE2575C5856" box="[957,1085,586,612]" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
have yielded such interesting materials, including most of the specimens of this species.
</paragraph>
</subSubSection>
<subSubSection id="F4FA49E6C760FF9AFF1DFD3251FE5996" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="BC5F1A6DC760FF9AFF1DFD325200594E" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFD3252245881" box="[198,325,666,691]" italics="true" pageId="4" pageNumber="5">Diagnosis.</emphasis>
Ocellar setae as strong as anteriormost reclinate fronto-orbital seta. Abdominal tergites silvergrey pruinose laterally and in posterior half, brownish-grey pruinose in anterior half; densely setulose, with setulae black; tergites 45 with dark spots at bases of setulae in male, spots lacking in female. In female: shape of sternite 2 trapezoidal; sternite 3 1.2 X wider than long; sternites 4 and 5 each with width and length subequal; sternite 7 with desclorotized area medioposteriorly, appearing as nearly divided; with tiny setulae along posterior margin.
</paragraph>
<paragraph id="BC5F1A6DC760FF9AFF1DFC2251FE5996" blockId="4.[151,1437,586,2012]" box="[198,671,906,932]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFC2252705991" box="[198,273,906,931]" italics="true" pageId="4" pageNumber="5">Adults</emphasis>
, ɗ, Ψ. Body length 4.0
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.
</paragraph>
</subSubSection>
<subSubSection id="F4FA49E6C760FF9BFF1DFC1A52595B18" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="BC5F1A6DC760FF9AFF1DFC1A53895E2E" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFC1A526759F9" box="[198,262,946,971]" italics="true" pageId="4" pageNumber="5">Head</emphasis>
(as in
<figureCitation id="24DB06E8C760FF9AFE81FC1A52CA59FE" box="[346,427,946,972]" captionStart="FIGURE 2" captionStartId="11.[151,255,1921,1945]" captionTargetBox="[187,1360,164,1905]" captionTargetId="figure@11.[187,1398,141,1918]" captionTargetPageId="11" captionText="FIGURE 2. Heads of new species. Scale bar = 1.0 mm. A B) Helgreelia albeto, sp. nov., Ψ (holotype); A) anterodorsal view, B) lateral view. C D) Neoschildomyia fusca, sp. nov., ♂ (holotype); C) anterodorsal view, D) lateral view. E F) Pradomyia hadromera, sp. nov., ♂ (holotype); E) anterodorsal view, F) lateral view." httpUri="https://zenodo.org/record/176056/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
AB). Height 1.92.2 X length; width 1.11.3 X height. Frons 2.42.9 X wider than long. Ocellar setae as strong as anteriormost reclinate fronto-orbital seta. Gena 0.500.53 X as high as head height.
</paragraph>
<paragraph id="BC5F1A6DC760FF9AFF1DFB8251B15E5E" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFB82527A5E71" box="[198,283,1066,1091]" italics="true" pageId="4" pageNumber="5">Thorax</emphasis>
(
<figureCitation id="24DB06E8C760FF9AFEF7FB8252115E76" box="[300,368,1066,1092]" captionStart="FIGURE 1" captionStartId="10.[151,255,1209,1233]" captionTargetBox="[150,1436,194,1183]" captionTargetId="figure@10.[150,1436,194,1185]" captionTargetPageId="10" captionText="FIGURE 1. Lateral habitus of holotypes of new species (for B D, post-dissection state); inset is a dorsal view of each abdomen. Scale bar = 1.0 mm. A) Helgreelia albeto, sp. nov., Ψ (inset is a paratype). B) Helgreelia parkeri, sp. nov., ♂. C) Neoschildomyia fusca, sp. nov., ♂. D) Pradomyia hadromera, sp. nov., ♂." httpUri="https://zenodo.org/record/176055/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
B). Scutum
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long. Scutellum 1.41.5 X wider than long.
<emphasis id="8E94C67FC760FF9AFB79FB8257BC5E71" box="[1186,1245,1066,1091]" italics="true" pageId="4" pageNumber="5">Wing</emphasis>
(as in
<figureCitation id="24DB06E8C760FF9AFAEBFB8256145E76" box="[1328,1397,1066,1092]" captionStart="FIGURE 3" captionStartId="12.[151,255,1357,1381]" captionTargetBox="[399,1188,203,1333]" captionTargetId="figure@12.[399,1189,194,1333]" captionTargetPageId="12" captionText="FIGURE 3. Wings of new species. Scale bar = 1.0 mm. A) Helgreelia albeto, sp. nov., Ψ (paratype). B) Neoschildomyia fusca, sp. nov., ♂ (holotype). C) Pradomyia hadromera, sp. nov., ♂ (holotype)." httpUri="https://zenodo.org/record/176057/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
A). Length
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, 2.83.0 X longer than high.
</paragraph>
<paragraph id="BC5F1A6DC760FF9AFF1DFBD351DB5F9E" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFBD352E15EA6" box="[198,384,1147,1172]" italics="true" pageId="4" pageNumber="5">Abdomen, male</emphasis>
(
<figureCitation id="24DB06E8C760FF9AFE49FBD252B95EA6" box="[402,472,1146,1172]" captionStart="FIGURE 1" captionStartId="10.[151,255,1209,1233]" captionTargetBox="[150,1436,194,1183]" captionTargetId="figure@10.[150,1436,194,1185]" captionTargetPageId="10" captionText="FIGURE 1. Lateral habitus of holotypes of new species (for B D, post-dissection state); inset is a dorsal view of each abdomen. Scale bar = 1.0 mm. A) Helgreelia albeto, sp. nov., Ψ (inset is a paratype). B) Helgreelia parkeri, sp. nov., ♂. C) Neoschildomyia fusca, sp. nov., ♂. D) Pradomyia hadromera, sp. nov., ♂." httpUri="https://zenodo.org/record/176055/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
B, inset). Tergites silver-grey pruinose laterally and in posterior half, brownishgrey pruinose in anterior half, except tergite 3 mostly brownish; densely setulose, setulae black; setulae along posterior edges of syntergite 1+2 through tergite 5 longer than others, more so laterally; tergites 45 with dark spots at bases of setulae. Sternites 25 also with setulae longest along posterior margin. Tergite 6 a thin transverse strip. Sternite 6 asymmetrical; a thin sclerotized strip only present on left side; fused to tergite 7 laterally; bare. Tergite 7 entire, as a thin, symmetrical, transverse, sclerotized strip (fused to sternite 6 on left side); bare; spiracle embedded in sclerite on left side. Tergite 8 separated from tergite 7, or only fused medially; 1/2 width of tergite 8; 2.0 X wider than long; bare.
</paragraph>
<paragraph id="BC5F1A6DC760FF9AFF1DFA13516B5D0E" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DFA1352125FE6" box="[198,371,1467,1492]" italics="true" pageId="4" pageNumber="5">Male genitalia</emphasis>
(
<figureCitation id="24DB06E8C760FF9AFE5EFA1252AC5FE6" box="[389,461,1466,1492]" captionStart="FIGURE 4" captionStartId="13.[151,255,1237,1261]" captionTargetBox="[324,1250,208,1204]" captionTargetId="figure@13.[308,1278,194,1213]" captionTargetPageId="13" captionText="FIGURE 4. Helgreelia parkeri, sp. nov., ♂ (holotype). Male genitalia. Scale bars = 0.01 mm. A) Epandrial complex, lateral view. B) Aedeagal complex, lateral view. C) Aedeagal complex, ventral view. a = aedeagus; aa = aedeagal apodeme; c = cercus; e = epandrium; g = gonopod; gl = articulated lobe of gonopod; h = hypandrium." httpUri="https://zenodo.org/record/176058/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
AC). Epandrium small; subquadrate from lateral view; setulose laterally; lacking discrete surstylar lobes, but slightly bulging in surstylar area, with 34 corresponding enlarged setulae. Hypandrium elongate, more than
<date id="C85E3CADC760FF9AFDFEF9A251315C16" box="[549,592,1546,1572]" pageId="4" pageNumber="5">2 X</date>
longer than maximum width; anterior half dorsoventrally flat, with lateral edges subparallel (slightly tapering anteriorly); posterior half broader than anterior part of hypandrium, deeply U-shaped from ventral view, extending laterally around aedeagal apodeme and aedeagus as saddle. Gonopod a simple lobe on posterior part of each hypandrial arm; setulose, with 2 slightly longer setulae posteriorly; with tiny, articulated dorsal lobe, bearing tiny setulae. Paramere absent. Aedeagus small, subequal in length to epandrium; surrounded by membrane; sclerotized part with distinct left and right halves, upcurved and pointed posteriorly. Aedeagal apodeme rod-like, 1.6 X longer than length of hypandrium; posterior end bilobed. Cercus small, setulose.
</paragraph>
<paragraph id="BC5F1A6DC760FF9AFF1DF8E356E55DEE" blockId="4.[151,1437,586,2012]" pageId="4" pageNumber="5">
<emphasis id="8E94C67FC760FF9AFF1DF8E352F85D56" box="[198,409,1867,1892]" italics="true" pageId="4" pageNumber="5">Abdomen, female.</emphasis>
As in male, except lacking dark spots at bases of setulae on tergites 45; setulae of tergite 5 dense, of normal size. Sternite 2 large,
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wide, 2.0 X wider than long; shape trapezoidal, anterior end wider than posterior end by 3.8 X. Sternite 3 3/5 width of sternite 2, 1.2 X wider than long. Sternite 4 1/2 width of sternite 2, width and length subequal. Sternite 5 1/3 width of sternite 2, width and length subequal.
</paragraph>
<paragraph id="BC5F1A6DC761FF9BFF1DFF3052595B18" blockId="5.[151,1437,152,658]" pageId="5" pageNumber="6">
<emphasis id="8E94C67FC761FF9BFF1DFF3052C65A83" box="[198,423,152,177]" italics="true" pageId="5" pageNumber="6">Female terminalia.</emphasis>
Tergite 7 dark, bare of pruinescence in anterior half, silvery-grey pruinose in posterior half; posterior margin densely setulose. Sternite 6 large, width 0.9 X that of sternite 3, 4.4 X wider than long. Sternite 7 with desclorotized area medioposteriorly, appearing as nearly divided; with tiny setulae along posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="F4FA49E6C761FF9BFF1DFE9052985858" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="BC5F1A6DC761FF9BFF1DFE9052985858" blockId="5.[151,1437,152,658]" pageId="5" pageNumber="6">
<emphasis id="8E94C67FC761FF9BFF1DFE9052065B63" box="[198,359,312,337]" italics="true" pageId="5" pageNumber="6">
<typeStatus id="635BA4CFC761FF9BFF1DFE90539A5B63" box="[198,251,312,337]" pageId="5" pageNumber="6">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="635BA4CFC761FF9BFEA1FE9052865B60" box="[378,487,312,338]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
ɗ (pinned, poor condition (antennal first flagellomeres missing, mid legs missing, several setae missing;
<figureCitation id="24DB06E8C761FF9BFE0AFEC851725B48" box="[465,531,352,378]" captionStart="FIGURE 1" captionStartId="10.[151,255,1209,1233]" captionTargetBox="[150,1436,194,1183]" captionTargetId="figure@10.[150,1436,194,1185]" captionTargetPageId="10" captionText="FIGURE 1. Lateral habitus of holotypes of new species (for B D, post-dissection state); inset is a dorsal view of each abdomen. Scale bar = 1.0 mm. A) Helgreelia albeto, sp. nov., Ψ (inset is a paratype). B) Helgreelia parkeri, sp. nov., ♂. C) Neoschildomyia fusca, sp. nov., ♂. D) Pradomyia hadromera, sp. nov., ♂." httpUri="https://zenodo.org/record/176055/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
B), abdomen dissected, in small vial with glycerin on same pin), deposited in
<collectionCode id="DAF182A8C761FF9BFF4CFE20538D5B90" LSID="urn:lsid:biocol.org:col:34871" box="[151,236,392,418]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
(with permission from
<collectionCode id="DAF182A8C761FF9BFDDEFE20513A5B90" box="[517,603,392,418]" pageId="5" pageNumber="6">EMUS</collectionCode>
), with following labels: “
<collectingCountry id="C4F75AFDC761FF9BFC5DFE2057595B90" box="[902,1080,392,418]" name="Costa Rica" pageId="5" pageNumber="6">
<collectionCode id="DAF182A8C761FF9BFC5DFE2057595B90" box="[902,1080,392,418]" pageId="5" pageNumber="6">COSTA RICA</collectionCode>
</collectingCountry>
, Guan[acaste Province].,
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SE R[io] Naranjo,
<date id="C85E3CADC761FF9BFEAAFE1851475BF8" box="[369,550,432,458]" pageId="5" pageNumber="6" value="1991-11-17" valueMax="1991-11-20" valueMin="1991-11-17">1720.XI.1991</date>
, F.D. Parker” / “SDG dissection 1321” / “
<typeStatus id="635BA4CFC761FF9BFBD1FE1857DD5BF8" box="[1034,1212,432,458]" pageId="5" pageNumber="6" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="7BE061EEC761FF9BFB18FE1852605BC0" authority="Gaimari." authorityName="Gaimari." class="Insecta" family="Odiniidae" genus="Helgreelia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="parkeri">
<emphasis id="8E94C67FC761FF9BFB18FE1856FD5BFB" box="[1219,1436,432,457]" italics="true" pageId="5" pageNumber="6">Helgreelia parkeri</emphasis>
Gaimari.
</taxonomicName>
<typeStatus id="635BA4CFC761FF9BFECCFE7052EC5BC0" box="[279,397,472,498]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="C4F75AFDC761FF9BFE46FE7051305BC0" box="[413,593,472,498]" name="Costa Rica" pageId="5" pageNumber="6">
<collectionCode id="DAF182A8C761FF9BFE46FE7051305BC0" box="[413,593,472,498]" pageId="5" pageNumber="6">COSTA RICA</collectionCode>
</collectingCountry>
: same label as
<typeStatus id="635BA4CFC761FF9BFCC9FE70501D5BC0" box="[786,892,472,498]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
, except
<date id="C85E3CADC761FF9BFC3FFE7057F75BC0" box="[996,1174,472,498]" pageId="5" pageNumber="6" value="1991-11-20" valueMax="1991-11-29" valueMin="1991-11-20">2029.XI.1991</date>
(1ɗ, SDG dissection 1320,
<collectionCode id="DAF182A8C761FF9BFF04FDA852575828" box="[223,310,512,538]" pageId="5" pageNumber="6">EMUS</collectionCode>
); Alajuela Province:
<quantity id="7B18B788C761FF9BFDF1FDA851125828" box="[554,627,512,538]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="km" value="20.0">20 km</quantity>
S Upala, F.D. Parker,
<date id="C85E3CADC761FF9BFCACFDA857775828" box="[887,1046,512,538]" pageId="5" pageNumber="6" value="1991-04-01" valueMax="1991-04-10" valueMin="1991-04-01">110.IV.1991</date>
(1Ψ,
<collectionCode id="DAF182A8C761FF9BFB8DFDA857CC5828" box="[1110,1197,512,538]" pageId="5" pageNumber="6">EMUS</collectionCode>
); San José Province: Est. Bijagual, Res. Biol. Carara,
<quantity id="7B18B788C761FF9BFDC9FD80513F5870" box="[530,606,552,578]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="m" value="500.0">500 m</quantity>
, R. Zuniga,
<date id="C85E3CADC761FF9BFD37FD8050225870" box="[748,835,552,578]" pageId="5" pageNumber="6" value="1991-02">II.1991</date>
, L-N-192250,474760,
<collectionCode id="DAF182A8C761FF9BFB94FD8057C15870" box="[1103,1184,552,578]" pageId="5" pageNumber="6">INBIO</collectionCode>
<collectionCode id="DAF182A8C761FF9BFB7CFD8057B95870" LSID="urn:lsid:biocol.org:col:14323" box="[1191,1240,552,578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="5" pageNumber="6">CRI</collectionCode>
000 313062 (1Ψ, SDG dissection 1323,
<collectionCode id="DAF182A8C761FF9BFE45FDF8528C5858" box="[414,493,592,618]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="6">INBio</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="F4FA49E6C761FF9BFF1DFDD0572C58A0" box="[198,1101,632,658]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="BC5F1A6DC761FF9BFF1DFDD0572C58A0" blockId="5.[151,1437,152,658]" box="[198,1101,632,658]" pageId="5" pageNumber="6">
<emphasis id="8E94C67FC761FF9BFF1DFDD0523458A3" box="[198,341,632,657]" italics="true" pageId="5" pageNumber="6">Distribution</emphasis>
. Known only from
<collectingCountry id="C4F75AFDC761FF9BFDE5FDD051DF58A0" box="[574,702,632,658]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
(Alajuela, Guanacaste, San José).
</paragraph>
</subSubSection>
</treatment>
</document>