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<document id="3AD036086EBEE21A0A7CE511D0FE3F24" ID-DOI="10.5281/zenodo.194942" ID-GBIF-Dataset="ebd5aa3c-6d5b-4720-a5a8-f6ebaf3dff66" ID-ISSN="1175-5326" ID-Zenodo-Dep="194942" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219771126" checkinUser="plazi" docAuthor="De, Francisco J." docDate="2010" docId="03F787AFFFAD891ED582FBDA7CDAC709" docLanguage="en" docName="zt02440p017.pdf" docOrigin="Zootaxa 2440" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Clupeidae Cuvier 1816, gen. nov." docType="treatment" docVersion="8" lastPageNumber="4" masterDocId="FFCEFFD7FFAF891DD515FFA57E0CC52E" masterDocTitle="Morphological and systematic reassessment of † Knightia brasiliensis Woodward, 1939 (Teleostei: Clupeiformes) from the Pliocene of Parnaíba Basin, northeastern Brazil" masterLastPageNumber="17" masterPageNumber="1" pageNumber="3" updateTime="1698237521598" updateUser="plazi">
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<mods:title id="10879FF23021C0B5E9F03ED89D5595F8">Morphological and systematic reassessment of † Knightia brasiliensis Woodward, 1939 (Teleostei: Clupeiformes) from the Pliocene of Parnaíba Basin, northeastern Brazil</mods:title>
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<mods:namePart id="87C7464216462F98B3CDEBA8EE7B6275">De, Francisco J.</mods:namePart>
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<treatment id="03F787AFFFAD891ED582FBDA7CDAC709" ID-DOI="http://doi.org/10.5281/zenodo.6200375" ID-GBIF-Taxon="119385075" ID-Zenodo-Dep="6200375" LSID="urn:lsid:plazi:treatment:03F787AFFFAD891ED582FBDA7CDAC709" httpUri="http://treatment.plazi.org/id/03F787AFFFAD891ED582FBDA7CDAC709" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C3446532FFAD891FD582FBDA7FD5C1F2" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE136B9FFAD891FD582FBDA7FB6C1B7" blockId="2.[151,442,1151,1177]" box="[151,442,1151,1177]" pageId="2" pageNumber="3">
<heading id="D0A981D5FFAD891FD582FBDA7FB6C1B7" bold="true" box="[151,442,1151,1177]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="4C5E4D3AFFAD891FD582FBDA7FB6C1B7" ID-CoL="8CJ" ID-ENA="55118" authority="Cuvier, 1816" authorityName="Cuvier" authorityYear="1816" box="[151,442,1151,1177]" class="Actinopterygii" family="Clupeidae" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="family">
<emphasis id="B92AEAABFFAD891FD582FBDA7FB6C1B7" bold="true" box="[151,442,1151,1177]" pageId="2" pageNumber="3">Clupeidae Cuvier, 1816</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE136B9FFAD891FD582FB677FD5C1F2" blockId="2.[151,473,1218,1244]" box="[151,473,1218,1244]" pageId="2" pageNumber="3">
<emphasis id="B92AEAABFFAD891FD582FB677F69C1F2" bold="true" box="[151,357,1218,1244]" pageId="2" pageNumber="3">
<taxonomicName id="4C5E4D3AFFAD891FD5B0FB677F69C1F2" box="[165,357,1218,1244]" class="Actinopterygii" family="Clupeidae" genus="Paleopiquitinga" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAABFFAD891FD5B0FB677F69C1F2" bold="true" box="[165,357,1218,1244]" italics="true" pageId="2" pageNumber="3">Paleopiquitinga</emphasis>
</taxonomicName>
</emphasis>
,
<taxonomicNameLabel id="A21957D0FFAD891FD461FB617FD5C1F2" box="[372,473,1220,1244]" pageId="2" pageNumber="3" rank="genus">
<emphasis id="B92AEAABFFAD891FD461FB617FD5C1F2" bold="true" box="[372,473,1220,1244]" pageId="2" pageNumber="3">gen. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3446532FFAD891FD582FAA27CE9C031" box="[151,741,1287,1311]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE136B9FFAD891FD582FAA27CE9C031" blockId="2.[151,741,1287,1311]" box="[151,741,1287,1311]" pageId="2" pageNumber="3">
<emphasis id="B92AEAABFFAD891FD582FAA27F21C031" bold="true" box="[151,301,1287,1311]" pageId="2" pageNumber="3">Type-species.</emphasis>
<taxonomicName id="4C5E4D3AFFAD891FD454FAAD7CEDC031" authority="Woodward, 1939" authorityName="Woodward" authorityYear="1939" box="[321,737,1288,1311]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B92AEAABFFAD891FD454FAAD7C2CC031" box="[321,544,1288,1311]" italics="true" pageId="2" pageNumber="3">Knightia brasiliensis</emphasis>
<bibRefCitation id="EFCF4B48FFAD891FD733FAAD7CEDC031" author="Woodward" box="[550,737,1288,1311]" pageId="2" pageNumber="17" refString="Woodward, A. S. (1939) Tertiary fossil fishes from Maranhao, Brazil. Annals and Magazine of Natural History, 11 (3), 450 - 453." type="journal article" year="1939">Woodward, 1939</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3446532FFAD891FD582FAE87C1BC09C" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BE136B9FFAD891FD582FAE87C1BC09C" blockId="2.[151,1437,1357,2018]" pageId="2" pageNumber="3">
<emphasis id="B92AEAABFFAD891FD582FAE87F2AC049" bold="true" box="[151,294,1357,1383]" pageId="2" pageNumber="3">Etymology.</emphasis>
The generic epithet is derived from the Greek word,
<emphasis id="B92AEAABFFAD891FD6BAFAE87A0BC048" box="[943,1031,1357,1382]" italics="true" pageId="2" pageNumber="3">palaios</emphasis>
, ancient; plus
<taxonomicName id="4C5E4D3AFFAD891FD1ACFAE87B39C048" box="[1209,1333,1357,1382]" class="Actinopterygii" family="Clupeidae" genus="Lile" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="piquitinga">
<emphasis id="B92AEAABFFAD891FD1ACFAE87B39C048" box="[1209,1333,1357,1382]" italics="true" pageId="2" pageNumber="3">piquitinga</emphasis>
</taxonomicName>
, a Tupi- Guarani name for a living herring [
<taxonomicName id="4C5E4D3AFFAD891FD724FAD67A3DC0A2" authority="Schreiner &amp; Ribeiro, 1903" authorityName="Schreiner &amp; Ribeiro" authorityYear="1903" box="[561,1073,1394,1420]" class="Actinopterygii" family="Clupeidae" genus="Lile" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="piquitinga">
<emphasis id="B92AEAABFFAD891FD724FAD67CD1C0A2" box="[561,733,1395,1420]" italics="true" pageId="2" pageNumber="3">Lile piquitinga</emphasis>
(
<bibRefCitation id="EFCF4B48FFAD891FD7F8FAD77A24C0A2" author="Schreiner" box="[749,1064,1394,1420]" pageId="2" pageNumber="17" refString="Schreiner, C. &amp; Ribeiro, A. de M., (1903) A colleccao de peixes do Museu Nacional do Rio de Janeiro. Arquivos do Museu Nacional, 1 - 41." type="book chapter" year="1903">Schreiner &amp; Ribeiro, 1903</bibRefCitation>
)
</taxonomicName>
] originally described in
<bibRefCitation id="EFCF4B48FFAD891FD058FAD77F51C09C" author="Pies" pageId="2" pageNumber="17" refString="Pies, W. &amp; Marcgraf, G., (1648) Historia Naturalis Brasiliae. Auspicio et Beneficio Illustriss. I. Mauritii Com. Nassau … non tantum Plantae et Animalia, sed et Indigenarum morbid, ingenia et mores describuntur et iconibus supra quingentas illustrantur. Amsterdam: Franciscus Hackium, Leiden and Ludovic Elzevier. 293 + (4) p." type="journal article" year="1648">Pies &amp; Marcgraf (1648)</bibRefCitation>
; gender female.
</paragraph>
</subSubSection>
<subSubSection id="C3446532FFAD891FD5D3FA187B86C2CC" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BE136B9FFAD891FD5D3FA187B86C2CC" blockId="2.[151,1437,1357,2018]" pageId="2" pageNumber="3">
<emphasis id="B92AEAABFFAD891FD5D3FA187F4AC0F9" bold="true" box="[198,326,1469,1495]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Slender and small-sized (rarely exceeding
<quantity id="4CA69B5CFFAD891FD67EFA187DC9C0F9" box="[875,965,1469,1495]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="mm" value="80.0">80 mm</quantity>
TL) clupeoid fish recognized by the following combination of features: head length contained 3 ½ in SL and equals to maximum body depth; anterior and posterior fontanels absent in the midline of cranial roof; longitudinal ridges on frontals; preepioccipital and large temporal fossae present; supratemporal commissure piercing parietal and supraoccipital bones; pterotic contributes to the temporal fossa; quadrate-mandible joint placed below the anterior half of orbit; jaws toothless; short lower jaw with deep coronoid process; two supramaxillae; seven branchiostegal rays; third postcleithrum rod-shaped; eight slender supraneural bones; 38 vertebrae, 15 caudal; large notochordal canal; short-based dorsal and anal fins; dorsal fin originating at the middle point of the body, anterior to pelvic fin; seven-rayed pelvic fin originating slightly behind vertical line anterior to the origin of dorsal fin; intermuscular epineural and epipleural bones extending over caudal region; predorsal scutes absent; at least 11 prepelvic abdominal scutes; eight postpelvic scutes; triangular pelvic scute; anteriormost preural vertebrae bearing long and thin neural spines; elongate leaf-like process of neural arch of first preural centrum; one epural; parhypurapophysis present; neural arch of the first preural centrum long and leaf-like; first ural centrum fused to first ural centrum; six hypurals; three uroneurals, with the first forming a strong pleurostyle; mid-sized cycloid scales with vertically arranged circuli; perforated scales of lateral line absent.
</paragraph>
</subSubSection>
<subSubSection id="C3446532FFAC891ED5D3FF327CDAC709" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BE136B9FFAC891ED5D3FF327CDAC709" blockId="3.[151,1437,151,551]" pageId="3" pageNumber="4">
<emphasis id="B92AEAABFFAC891ED5D3FF327F4CC59F" bold="true" box="[198,320,151,177]" pageId="3" pageNumber="4">Remarks.</emphasis>
<taxonomicName id="4C5E4D3AFFAC891ED440FF3D7C28C59F" box="[341,548,152,177]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="eoceana">
<emphasis id="B92AEAABFFAC891ED440FF3D7C28C59F" box="[341,548,152,177]" italics="true" pageId="3" pageNumber="4">Knightia eoceana</emphasis>
</taxonomicName>
from the Lower Eocene of Wyoming was described by
<bibRefCitation id="EFCF4B48FFAC891ED1BAFF3D7B5FC59C" author="Jordan" box="[1199,1363,152,178]" pageId="3" pageNumber="16" refString="Jordan, D. S. (1907) The fossil fishes of California; with suplementary notes on other species of extinct fishes. Bulletin of the Department of Geology, University of California, 5, 95 - 145." type="journal article" year="1907">
<collectingCountry id="F3497629FFAC891ED1BAFF3D7AF1C59C" box="[1199,1277,152,178]" name="Jordan" pageId="3" pageNumber="4">Jordan</collectingCountry>
(1907)
</bibRefCitation>
as the type-species of †
<taxonomicName id="4C5E4D3AFFAC891ED448FF187FCFC5F8" box="[349,451,189,214]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED448FF187FCFC5F8" box="[349,451,189,214]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFCF4B48FFAC891ED4C7FF187C71C5F9" author="Grande" box="[466,637,189,215]" pageId="3" pageNumber="16" refString="Grande, L. (1982) A revision of the fossil genus † Knightia, with a description of a new genus from the Green River Formation (Teleostei, Clupeidae). American Museum Novitates, 2731, 1 - 22." type="journal article" year="1982">Grande (1982)</bibRefCitation>
furnished a revised diagnosis for †
<taxonomicName id="4C5E4D3AFFAC891ED10CFF187A8CC5F8" box="[1049,1152,189,214]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED10CFF187A8CC5F8" box="[1049,1152,189,214]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
based on a combination of the following features: i) one supramaxillary, ii) complete series of 1214 dorsal scutes (each one with unsculptured wings and median keel), and (iii), relatively few vertebrae (36 to 40) and pleural ribs (20-22 pairs). He listed seven valid species: †
<taxonomicName id="4C5E4D3AFFAC891ED76CFE887D59C468" box="[633,853,301,326]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="eoceana">
<emphasis id="B92AEAABFFAC891ED76CFE887D59C468" box="[633,853,301,326]" italics="true" pageId="3" pageNumber="4">Knightia eoceana</emphasis>
</taxonomicName>
, †
<taxonomicName id="4C5E4D3AFFAC891ED663FE887A10C468" box="[886,1052,301,326]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="alta">
<emphasis id="B92AEAABFFAC891ED663FE887A10C468" box="[886,1052,301,326]" italics="true" pageId="3" pageNumber="4">Knightia alta</emphasis>
</taxonomicName>
, †
<taxonomicName id="4C5E4D3AFFAC891ED129FE887AA5C468" box="[1084,1193,301,326]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED129FE887AA5C468" box="[1084,1193,301,326]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
<emphasis id="B92AEAABFFAC891ED1AAFE887B0EC468" box="[1215,1282,301,326]" italics="true" pageId="3" pageNumber="4">copei</emphasis>
”, †
<taxonomicName id="4C5E4D3AFFAC891ED025FE887EE2C445" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="vetusta">
<emphasis id="B92AEAABFFAC891ED025FE887EE2C445" italics="true" pageId="3" pageNumber="4">Knightia vetusta</emphasis>
</taxonomicName>
, †
<taxonomicName id="4C5E4D3AFFAC891ED41BFEF77F75C445" box="[270,377,338,363]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED41BFEF77F75C445" box="[270,377,338,363]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
n.sp (Tertiary of eastern
<collectingCountry id="F3497629FFAC891ED7A7FEF77D0CC442" box="[690,768,338,364]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
), †
<taxonomicName id="4C5E4D3AFFAC891ED63BFEF77D96C445" box="[814,922,338,363]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED63BFEF77D96C445" box="[814,922,338,363]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
?
<emphasis id="B92AEAABFFAC891ED6ADFEF77A13C445" box="[952,1055,338,363]" italics="true" pageId="3" pageNumber="4">yuyanga</emphasis>
, and †“
<emphasis id="B92AEAABFFAC891ED197FEF77B98C445" box="[1154,1428,338,363]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C5E4D3AFFAC891ED197FEF77AFFC445" box="[1154,1267,338,363]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">Knightia</taxonomicName>
<taxonomicName id="4C5E4D3AFFAC891ED013FEF77B98C445" box="[1286,1428,338,363]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="brasiliensis">brasiliensis</taxonomicName>
</emphasis>
. Afterwards,
<bibRefCitation id="EFCF4B48FFAC891ED43AFEDD7FE8C4BC" author="Grande" box="[303,484,376,402]" pageId="3" pageNumber="16" refString="Grande, L. (1985) Recent and fossil clupeomorph fishes with materials for revision of the subgroups of clupeoids. Bulletin of the American Museum of Natural History, 181, 231 - 372." type="journal article" year="1985">Grande (1985)</bibRefCitation>
, concerning † “
<taxonomicName id="4C5E4D3AFFAC891ED7B0FEDD7D03C4BF" box="[677,783,376,401]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED7B0FEDD7D03C4BF" box="[677,783,376,401]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
<taxonomicName id="4C5E4D3AFFAC891ED633FEDD7DBEC4BF" box="[806,946,376,401]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B92AEAABFFAC891ED633FEDD7DBEC4BF" box="[806,946,376,401]" italics="true" pageId="3" pageNumber="4">brasiliensis</emphasis>
</taxonomicName>
, pointed out that “for reasons discussed above, this species must be removed from †
<taxonomicName id="4C5E4D3AFFAC891ED789FE387D0FC498" box="[668,771,413,438]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED789FE387D0FC498" box="[668,771,413,438]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
(no new genus is proposed here without further study)”. †
<taxonomicName id="4C5E4D3AFFAC891ED5B0FE677FFDC4F5" box="[165,497,450,475]" class="Actinopterygii" family="Clupeidae" genus="Paleopiquitinga" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B92AEAABFFAC891ED5B0FE677FFDC4F5" box="[165,497,450,475]" italics="true" pageId="3" pageNumber="4">Paleopiquitinga brasiliensis</emphasis>
</taxonomicName>
possesses two supramaxillaries and lacks dorsal scutes. Only the lesser number of vertebrae and pleural ribs are shared by †
<taxonomicName id="4C5E4D3AFFAC891ED7DCFE4D7D98C72F" box="[713,916,488,513]" class="Actinopterygii" family="Clupeidae" genus="Paleopiquitinga" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAABFFAC891ED7DCFE4D7D98C72F" box="[713,916,488,513]" italics="true" pageId="3" pageNumber="4">Paleopiquitinga</emphasis>
</taxonomicName>
and †
<taxonomicName id="4C5E4D3AFFAC891ED6F6FE4D7A5DC72F" box="[995,1105,488,513]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAABFFAC891ED6F6FE4D7A5DC72F" box="[995,1105,488,513]" italics="true" pageId="3" pageNumber="4">Knightia</emphasis>
</taxonomicName>
; which are also shared by †
<emphasis id="B92AEAABFFAC891ED5B0FDA87F92C708" box="[165,414,525,550]" italics="true" pageId="3" pageNumber="4">Gosiutichthys parvus</emphasis>
(see
<bibRefCitation id="EFCF4B48FFAC891ED4CEFDA87C72C709" author="Grande" box="[475,638,525,551]" pageId="3" pageNumber="16" refString="Grande, L. (1982) A revision of the fossil genus † Knightia, with a description of a new genus from the Green River Formation (Teleostei, Clupeidae). American Museum Novitates, 2731, 1 - 22." type="journal article" year="1982">Grande, 1982</bibRefCitation>
;
<bibRefCitation id="EFCF4B48FFAC891ED79FFDA87CC5C709" author="Grande" box="[650,713,525,551]" pageId="3" pageNumber="16" refString="Grande, L. (1985) Recent and fossil clupeomorph fishes with materials for revision of the subgroups of clupeoids. Bulletin of the American Museum of Natural History, 181, 231 - 372." type="journal article" year="1985">1985</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>