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<document id="72FA4B04DBE7703C4E3A19C1AE6223F9" ID="10.11646/zootaxa.4244.1.1SLASH10518" ID-CLB-Dataset="36256" ID-DOI="10.11646/zootaxa.4244.1.1" ID-GBIF-Dataset="05f1a37b-29f3-4bca-a735-178a3373bb8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="400400" ID-ZooBank="781D71C8-4632-4D1B-8D82-F77CA1146029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489738266586" checkinUser="plazi" docAuthor="Hogans, W. E." docDate="2017" docId="03F777535B62FFE9D6A0FE8F7325F344" docLanguage="en" docName="zootaxa.4244.1.1SLASH10518.pdf" docOrigin="Zootaxa 4244 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pennella hawaiiensis Kazachenko &amp; Kurochkin 1974" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FFCE0F2B5B77FFFFD637FFC57039F11B" masterDocTitle="Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean" masterLastPageNumber="38" masterPageNumber="1" pageNumber="22" updateTime="1698438091026" updateUser="ExternalLinkService">
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<mods:title id="6E580FD7BD4A9E83E19DAC60430C95A0">Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean</mods:title>
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<mods:date id="A220DEFE5D55B11A07B98B8B3745CCBC">2017</mods:date>
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<treatment id="03F777535B62FFE9D6A0FE8F7325F344" ID-DOI="http://doi.org/10.5281/zenodo.6052553" ID-GBIF-Taxon="127847784" ID-Zenodo-Dep="6052553" LSID="urn:lsid:plazi:treatment:03F777535B62FFE9D6A0FE8F7325F344" httpUri="http://treatment.plazi.org/id/03F777535B62FFE9D6A0FE8F7325F344" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C34495CE5B62FFEAD6A0FE8F70C1F09D" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BE1C6455B62FFEAD6A0FE8F730BF07F" blockId="21.[151,818,330,390]" box="[151,818,330,356]" pageId="21" pageNumber="22">
<heading id="D0A971295B62FFEAD6A0FE8F730BF07F" bold="true" box="[151,818,330,356]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C5EBDC65B62FFEAD6A0FE8F730BF07F" authority="Kazachenko &amp; Kurochkin, 1974" authorityName="Kazachenko &amp; Kurochkin" authorityYear="1974" box="[151,818,330,356]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="B92A1A575B62FFEAD6A0FE8F730BF07F" bold="true" box="[151,818,330,356]" pageId="21" pageNumber="22">
<emphasis id="B92A1A575B62FFEAD6A0FE8F71AEF07F" bold="true" box="[151,407,330,356]" italics="true" pageId="21" pageNumber="22">Pennella hawaiiensis</emphasis>
<bibRefCitation id="EFCFBBB45B62FFEAD7A9FE8F730BF07F" author="Kazachenko" box="[414,818,330,356]" pageId="21" pageNumber="22" refString="Kazachenko, V. N. &amp; Kurochkin, Y. V. (1974) On a new species of parasitic copepod, Pennella hawaiiensis sp. nov. from the boarfish, Pentoceros richardsoni. Proceedings of the Pacific Sciences Research Institute of Oceanography, 88, 42 - 53." type="journal article" year="1974">Kazachenko &amp; Kurochkin, 1974</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE1C6455B62FFEAD6A0FEA870C1F09D" blockId="21.[151,818,330,390]" box="[151,248,365,390]" pageId="21" pageNumber="22">
(
<figureCitation id="1365DAC05B62FFEAD6A8FEA870D6F09D" box="[159,239,365,390]" captionStart="FIGURE 11" captionStartId="21.[151,250,1922,1944]" captionTargetBox="[215,1372,441,1894]" captionTargetId="figure@21.[196,1390,441,1903]" captionTargetPageId="21" captionText="FIGURE 11. Pennella hawaiiensis, a. habitus, old female; b. habitus, younger female; c. cephalothorax of young specimen, ventral; d. first antenna; e. second antenna; f. first swimming leg; g. second swimming leg; h. detail of abdominal plume (from Kazachenko &amp; Kurochkin 1974). Scale bars: c = 0.1; d-f = 0.02 mm; g = 0.2 mm." httpUri="https://zenodo.org/record/400410/files/figure.png" pageId="21" pageNumber="22">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF2196CD5B62FFEAD6A0F84773FCF6CE" httpUri="https://zenodo.org/record/400410/files/figure.png" pageId="21" pageNumber="22" targetBox="[215,1372,441,1894]" targetPageId="21">
<paragraph id="8BE1C6455B62FFEAD6A0F84773FCF6CE" blockId="21.[151,1436,1922,2005]" pageId="21" pageNumber="22">
<emphasis id="B92A1A575B62FFEAD6A0F8477126F683" bold="true" box="[151,287,1922,1944]" pageId="21" pageNumber="22">FIGURE 11.</emphasis>
<taxonomicName id="4C5EBDC65B62FFEAD71FF847723BF683" box="[296,514,1922,1944]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="B92A1A575B62FFEAD71FF847723BF683" box="[296,514,1922,1944]" italics="true" pageId="21" pageNumber="22">Pennella hawaiiensis</emphasis>
</taxonomicName>
, a. habitus, old female; b. habitus, younger female; c. cephalothorax of young specimen, ventral; d. first antenna; e. second antenna; f. first swimming leg; g. second swimming leg; h. detail of abdominal plume (from Kazachenko &amp; Kurochkin 1974). Scale bars: c =0.1; d-f = 0.02 mm; g =0.2 mm.
</paragraph>
</caption>
<subSubSection id="C34495CE5B61FFE9D6A0FF52715DF1B4" box="[151,356,151,176]" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE1C6455B61FFE9D6A0FF52715DF1B4" blockId="22.[151,1437,151,608]" box="[151,356,151,176]" pageId="22" pageNumber="23">
<emphasis id="B92A1A575B61FFE9D6A0FF527120F1AB" bold="true" box="[151,281,151,176]" pageId="22" pageNumber="23">Synonyms.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C34495CE5B61FFE9D6F0FF7973CDF05B" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BE1C6455B61FFE9D6F0FF7973EDF1CF" blockId="22.[151,1437,151,608]" box="[199,980,188,213]" pageId="22" pageNumber="23">
<materialsCitation id="3B36CC185B61FFE9D6F0FF7973EDF1CF" ID-GBIF-Occurrence="1440667655" box="[199,980,188,213]" location="Pentoceros" municipality="south Pacific Ocean." pageId="22" pageNumber="23" specimenCount="1" typeStatus="holotype">
<emphasis id="B92A1A575B61FFE9D6F0FF7971EBF1CE" bold="true" box="[199,466,188,213]" pageId="22" pageNumber="23">
<typeStatus id="54E578E75B61FFE9D6F0FF79713BF1CE" box="[199,258,188,213]" pageId="22" pageNumber="23">Type</typeStatus>
host and locality.
</emphasis>
<emphasis id="B92A1A575B61FFE9D7EEFF7872E6F1CE" box="[473,735,188,213]" italics="true" pageId="22" pageNumber="23">
<location id="8E81909E5B61FFE9D7EEFF78726FF1CF" LSID="urn:lsid:plazi:treatment:03F777535B62FFE9D6A0FE8F7325F344:8E81909E5B61FFE9D7EEFF78726FF1CF" box="[473,598,189,212]" municipality="south Pacific Ocean." name="Pentoceros" pageId="22" pageNumber="23">Pentoceros</location>
<taxonomicName id="4C5EBDC65B61FFE9D46AFF7972E6F1CE" box="[605,735,188,213]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="richardsoni">richardsoni</taxonomicName>
</emphasis>
,
<collectingMunicipality id="6B855C3F5B61FFE9D4DBFF7973EDF1CF" box="[748,980,188,213]" pageId="22" pageNumber="23">south Pacific Ocean.</collectingMunicipality>
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</paragraph>
<paragraph id="8BE1C6455B61FFE9D6F0FF1A73CDF05B" blockId="22.[151,1437,151,608]" pageId="22" pageNumber="23">
<emphasis id="B92A1A575B61FFE9D6F0FF1A7158F1E3" bold="true" box="[199,353,223,248]" pageId="22" pageNumber="23">Morphology.</emphasis>
Size:
<quantity id="4CA66BA05B61FFE9D79FFF25721FF1EC" box="[424,550,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="22" pageNumber="23" unit="mm" value="85.0" valueMax="90.0" valueMin="80.0">8090 mm</quantity>
. Papillae: partial coverage, generally spherical, variable in size and shape, not found in organized groups. Holdfasts: three, two laterals short; dorsal horn shorter. First antenna with four segments, second with two segments. Plumes: dendritic, complex branching.
</paragraph>
</subSubSection>
<subSubSection id="C34495CE5B61FFE9D6F0FE897325F344" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BE1C6455B61FFE9D6F0FE897325F344" blockId="22.[151,1437,151,608]" pageId="22" pageNumber="23">
<emphasis id="B92A1A575B61FFE9D6F0FE897102F07E" bold="true" box="[199,315,332,357]" pageId="22" pageNumber="23">Remarks.</emphasis>
A valid species. Well described, figured and discussed in the original report. An intermediate size
<taxonomicName id="4C5EBDC65B61FFE9D6A0FEAA70C2F093" box="[151,251,367,392]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B92A1A575B61FFE9D6A0FEAA70C2F093" box="[151,251,367,392]" italics="true" pageId="22" pageNumber="23">Pennella</emphasis>
</taxonomicName>
species parasitic on a unique host, the pelagic armourhead (boarfish),
<taxonomicName id="4C5EBDC65B61FFE9D219FEB474F6F093" box="[1070,1231,367,392]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="B92A1A575B61FFE9D219FEB474F6F093" box="[1070,1231,367,392]" italics="true" pageId="22" pageNumber="23">P. richardsoni</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFCFBBB45B61FFE9D2E8FEAA7151F0B7" author="Kazachenko" pageId="22" pageNumber="23" refString="Kazachenko, V. N. &amp; Kurochkin, Y. V. (1974) On a new species of parasitic copepod, Pennella hawaiiensis sp. nov. from the boarfish, Pentoceros richardsoni. Proceedings of the Pacific Sciences Research Institute of Oceanography, 88, 42 - 53." type="journal article" year="1974">Kazachenko and Kurochkin (1974)</bibRefCitation>
documented several specimens of the adult female and younger pre-adult individuals in metamorphosis.
<taxonomicName id="4C5EBDC65B61FFE9D765FE72727BF0CB" box="[338,578,439,464]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="B92A1A575B61FFE9D765FE72727BF0CB" box="[338,578,439,464]" italics="true" pageId="22" pageNumber="23">Pennella hawaiiensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C5EBDC65B61FFE9D4F3FE7C7326F0CB" box="[708,799,439,464]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B92A1A575B61FFE9D4F3FE7C7326F0CB" box="[708,799,439,464]" italics="true" pageId="22" pageNumber="23">P. filosa</emphasis>
</taxonomicName>
but is a smaller parasite; it also superficially resembles and is close in size to
<emphasis id="B92A1A575B61FFE9D7A5FE187236F0EF" box="[402,527,477,500]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C5EBDC65B61FFE9D7A5FE187232F0EF" box="[402,523,477,500]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="remorae">P. remorae</taxonomicName>
,
</emphasis>
but has a different cephalothoracic papillae configuration, has one less segment on the first antenna, and parasitizes a much different
<typeStatus id="54E578E75B61FFE9D53EFDC57300F303" box="[777,825,512,536]" pageId="22" pageNumber="23">type</typeStatus>
of host.
<taxonomicName id="4C5EBDC65B61FFE9D591FE3A74A1F303" box="[934,1176,511,536]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="B92A1A575B61FFE9D591FE3A74A1F303" box="[934,1176,511,536]" italics="true" pageId="22" pageNumber="23">Pennella hawaiiensis</emphasis>
</taxonomicName>
is the only species of
<taxonomicName id="4C5EBDC65B61FFE9D6A0FDE170C2F326" box="[151,251,548,573]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B92A1A575B61FFE9D6A0FDE170C2F326" box="[151,251,548,573]" italics="true" pageId="22" pageNumber="23">Pennella</emphasis>
</taxonomicName>
other than
<taxonomicName id="4C5EBDC65B61FFE9D74AFDE071CEF327" box="[381,503,549,572]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="exocoeti">
<emphasis id="B92A1A575B61FFE9D74AFDE071CEF327" box="[381,503,549,572]" italics="true" pageId="22" pageNumber="23">P. exocoeti</emphasis>
</taxonomicName>
(a smaller parasite which does not possess a dorsal holdfast horn and is found on flyingfish) which exhibits a four-segmented first antenna.
</paragraph>
</subSubSection>
</treatment>
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