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<document id="ABE76A921ACFBB48186E1C5AC820120F" ID-DOI="10.1111/j.1096-3642.2011.00738.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442323" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452876177" checkinUser="valdenar" docAuthor="Karaman, Ivo, Hammouti, Nasera, Pavićević, Dragan, Kiefer, Andreas, Horvatović, Mladen &amp; Seitz, Alfred" docDate="2011" docId="03F75E0EFFD3FFA8FCF4FAE1FC7BF8B6" docLanguage="en" docName="j.1096-3642.2011.00738.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00738.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Troglophilus neglectus VLASINENSIS" docType="treatment" docVersion="3" lastPageNumber="1045" masterDocId="FFCE2676FFDBFFA2FF98FFDBFFD3FFD9" masterDocTitle="The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans" masterLastPageNumber="1063" masterPageNumber="1035" pageNumber="1043" updateTime="1699275884152" updateUser="plazi">
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<mods:title id="8FA59CE565E8C1F078176ECAC400902D">The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans</mods:title>
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<mods:namePart id="47C386E45F83906267570F788EF7CFE4">Karaman, Ivo</mods:namePart>
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<mods:namePart id="A0A58745037760ABBD0ED11E0300BB67">Hammouti, Nasera</mods:namePart>
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<mods:namePart id="E2251EF8384B3A17E736D9E304F8CA9A">Pavićević, Dragan</mods:namePart>
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<mods:namePart id="1C90F6FAEEEBA8E8F9C3015BC851F618">Kiefer, Andreas</mods:namePart>
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<mods:namePart id="1DE199714EBAAE047604C1416A49B77B">Horvatović, Mladen</mods:namePart>
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<mods:namePart id="3008BFC752A0CB9A6B030B2F0E1AF6AB">Seitz, Alfred</mods:namePart>
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<mods:title id="2027B5C25720AECE6EEC835AFA296422">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="8D81C9C1F21620DB57BB2B796D1FD315">2011</mods:date>
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<mods:number id="2D15330E33D72185C2ABCE62DD6F69C2">163</mods:number>
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<mods:identifier id="0DD42A0E2941FB7009BA6E4B8311A2C3" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F75E0EFFD3FFA8FCF4FAE1FC7BF8B6" LSID="urn:lsid:plazi:treatment:03F75E0EFFD3FFA8FCF4FAE1FC7BF8B6" httpUri="http://treatment.plazi.org/id/03F75E0EFFD3FFA8FCF4FAE1FC7BF8B6" lastPageId="10" lastPageNumber="1045" pageId="8" pageNumber="1043">
<subSubSection id="C344BC93FFD3FFAAFCF4FAE1FB4EFAB6" pageId="8" pageNumber="1043" type="nomenclature">
<paragraph id="8BE1EF18FFD3FFAAFCF4FAE1FB4EFAB6" blockId="8.[876,1352,1337,1391]" pageId="8" pageNumber="1043">
<heading id="D0A95874FFD3FFAAFCF4FAE1FB4EFAB6" centered="true" fontSize="8" level="2" pageId="8" pageNumber="1043" reason="8">
<taxonomicName id="4C5E949BFFD3FFAAFCF4FAE1FA92FA94" ID-CoL="5993L" authority="VLASINENSIS" authorityName="VLASINENSIS" box="[876,1345,1338,1359]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="1043" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B92A330AFFD3FFAAFCF4FAE1FA92FA94" box="[876,1345,1338,1359]" italics="true" pageId="8" pageNumber="1043">TROGLOPHILUS NEGLECTUS VLASINENSIS</emphasis>
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:
<bibRefCitation id="EFCF92E9FFD3FFAAFB8EFA82FB4EFAB6" author="Harz K" box="[1046,1181,1369,1391]" pageId="8" pageNumber="1043" refId="ref10657" refString="Harz K. 1969. Die otrhopteren europas. The Hague: Dr. W. Junk B. V, vol. I." type="journal volume" year="1969">HARZ, 1969</bibRefCitation>
</heading>
</paragraph>
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<subSubSection id="C344BC93FFD3FFA8FCBEFAA4FC7BF8B6" lastPageId="10" lastPageNumber="1045" pageId="8" pageNumber="1043" type="materials_examined">
<paragraph id="8BE1EF18FFD3FFAAFCBEFAA4FB65FA4C" blockId="8.[806,1206,1407,1429]" box="[806,1206,1407,1429]" pageId="8" pageNumber="1043">
<emphasis id="B92A330AFFD3FFAAFCBEFAA4FBD5FA4D" box="[806,1030,1407,1428]" italics="true" pageId="8" pageNumber="1043">Material examined:</emphasis>
See Appendix.
</paragraph>
<paragraph id="8BE1EF18FFD3FFAAFCBEFA17FBE8F8B6" blockId="8.[806,1421,1483,1903]" pageId="8" pageNumber="1043">
<emphasis id="B92A330AFFD3FFAAFCBEFA17FC4EFA38" box="[806,925,1484,1505]" italics="true" pageId="8" pageNumber="1043">Diagnosis:</emphasis>
Medium to large sized species. Male body length ranges between 13.5
<quantity id="4CA642FDFFD3FFAAFB52FA32FAC8FA26" box="[1226,1307,1513,1535]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="1043" unit="mm" value="20.0">20 mm</quantity>
; females,
<quantity id="4CA642FDFFD3FFAAFCBEF9D3FC73F9C7" box="[806,928,1544,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.4" metricValueMin="1.4" pageId="8" pageNumber="1043" unit="mm" value="19.0" valueMax="24.0" valueMin="14.0">1424 mm</quantity>
. Body colour dark brown, light brown to light green marbled. Males tenth tergite with narrow, partially square, medial lobe which is divided by a central deep depression in two triangular-shaped lobes (
<figureCitation id="1365F39DFFD3FFAAFCEFF958FC3CF940" box="[887,1007,1667,1689]" captionStart="Figure 6" captionStartId="9.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-230@9.[323,1283,196,1286]" captionTargetPageId="9" captionText="Figure 6. AF, Troglophilus neglectus, tenth tergite, dorsal view: A, male 16.5 mm (Bihac´, Bosnia); B, male 16.5 mm (Kranj, Slovenia); C, male 15.5 mm (Banjani, Macedonia); D, male 17 mm (Smarna Gora, Slovenia); E, female 17 mm (Blace, Macedonia); F, female 15.5 mm (Kopajkošare, Serbia)." figureDoi="http://doi.org/10.5281/zenodo.5442335" httpUri="https://zenodo.org/record/5442335/files/figure.png" pageId="8" pageNumber="1043">Fig. 6AD</figureCitation>
). Female tenth abdominal tergite with two paramedial, sharp spine-like protrusions (
<figureCitation id="1365F39DFFD3FFAAFCB6F91BFC70F90F" box="[814,931,1728,1750]" captionStart="Figure 6" captionStartId="9.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-230@9.[323,1283,196,1286]" captionTargetPageId="9" captionText="Figure 6. AF, Troglophilus neglectus, tenth tergite, dorsal view: A, male 16.5 mm (Bihac´, Bosnia); B, male 16.5 mm (Kranj, Slovenia); C, male 15.5 mm (Banjani, Macedonia); D, male 17 mm (Smarna Gora, Slovenia); E, female 17 mm (Blace, Macedonia); F, female 15.5 mm (Kopajkošare, Serbia)." figureDoi="http://doi.org/10.5281/zenodo.5442335" httpUri="https://zenodo.org/record/5442335/files/figure.png" pageId="8" pageNumber="1043">Fig. 6EF</figureCitation>
). The titillator distally protruded with abruptly narrowed obtuse tip (
<figureCitation id="1365F39DFFD3FFAAFB30F905FB2EF92C" box="[1192,1277,1758,1781]" captionStart="Figure 7" captionStartId="10.[144,223,1316,1335]" captionTargetBox="[303,1263,196,1286]" captionTargetId="figure-218@10.[303,1263,196,1286]" captionTargetPageId="10" captionText="Figure 7. AB, Troglophilus neglectus, male 17 mm, phallus with titillator (arrow) (Smarna gora, Slovenia): A, dorsal view; B, lateral view. CD, Troglophilus ovuliformis, male 20 mm, phallus with titillator (arrow) (G. Seoca, Montenegro): C, dorsal view; D, lateral view. E, T. ovuliformis, male 20 mm, titillator (arrow), dorsal view (Island Pag, Croatia). F, T. ovuliformis, male 20 mm, titillator (arrow), dorsal view (D. Seoca, Montenegro)." figureDoi="http://doi.org/10.5281/zenodo.5442337" httpUri="https://zenodo.org/record/5442337/files/figure.png" pageId="8" pageNumber="1043">Fig. 7A</figureCitation>
); in lateral view, distally voluminous (
<figureCitation id="1365F39DFFD3FFAAFBE8F926FB15F8CB" box="[1136,1222,1789,1811]" captionStart="Figure 7" captionStartId="10.[144,223,1316,1335]" captionTargetBox="[303,1263,196,1286]" captionTargetId="figure-218@10.[303,1263,196,1286]" captionTargetPageId="10" captionText="Figure 7. AB, Troglophilus neglectus, male 17 mm, phallus with titillator (arrow) (Smarna gora, Slovenia): A, dorsal view; B, lateral view. CD, Troglophilus ovuliformis, male 20 mm, phallus with titillator (arrow) (G. Seoca, Montenegro): C, dorsal view; D, lateral view. E, T. ovuliformis, male 20 mm, titillator (arrow), dorsal view (Island Pag, Croatia). F, T. ovuliformis, male 20 mm, titillator (arrow), dorsal view (D. Seoca, Montenegro)." figureDoi="http://doi.org/10.5281/zenodo.5442337" httpUri="https://zenodo.org/record/5442337/files/figure.png" pageId="8" pageNumber="1043">Fig. 7B</figureCitation>
). The ovipositor short and wide, length ranges between
<quantity id="4CA642FDFFD3FFAAFB69F8C7FA88F8E8" box="[1265,1371,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="8" pageNumber="1043" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
; the widest point is in its proximal third (
<figureCitation id="1365F39DFFD3FFAAFB54F8E1FAF5F889" box="[1228,1318,1850,1872]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. AC, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). DI ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="8" pageNumber="1043">Fig. 12I</figureCitation>
), ventral margin almost straight.
</paragraph>
<caption id="DF21BF90FFD2FFABFF3CFAFFFCE5FAA8" ID-DOI="http://doi.org/10.5281/zenodo.5442335" ID-Zenodo-Dep="5442335" httpUri="https://zenodo.org/record/5442335/files/figure.png" pageId="9" pageNumber="1044" startId="9.[164,243,1316,1335]" targetBox="[323,1283,196,1286]" targetPageId="9">
<paragraph id="8BE1EF18FFD2FFABFF3CFAFFFCE5FAA8" blockId="9.[164,1441,1316,1394]" pageId="9" pageNumber="1044">
<emphasis id="B92A330AFFD2FFABFF3CFAFFFEDFFAEE" bold="true" box="[164,268,1316,1335]" pageId="9" pageNumber="1044">Figure 6.</emphasis>
AF,
<taxonomicName id="4C5E949BFFD2FFABFECBFAFFFDECFAEE" box="[339,575,1316,1335]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="1044" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B92A330AFFD2FFABFECBFAFFFDECFAEE" box="[339,575,1316,1335]" italics="true" pageId="9" pageNumber="1044">Troglophilus neglectus</emphasis>
</taxonomicName>
, tenth tergite, dorsal view: A, male 16.5 mm (Bihac´, Bosnia); B, male 16.5 mm (Kranj, Slovenia); C, male 15.5 mm (Banjani, Macedonia); D, male 17 mm (Smarna Gora, Slovenia); E, female 17 mm (Blace, Macedonia); F, female 15.5 mm (Kopajkošare, Serbia).
</paragraph>
</caption>
<paragraph id="8BE1EF18FFD2FFABFF3CFA77FF2CF90F" blockId="9.[164,779,1452,1903]" pageId="9" pageNumber="1044">
<emphasis id="B92A330AFFD2FFABFF3CFA77FEE6FA18" box="[164,309,1452,1473]" italics="true" pageId="9" pageNumber="1044">Distribution:</emphasis>
This species is widely distributed over a large part of the Balkans (
<figureCitation id="1365F39DFFD2FFABFE76FA10FDFDFA38" box="[494,558,1483,1505]" captionStart="Figure 8" captionStartId="11.[164,243,1076,1095]" captionTargetBox="[164,1445,196,1047]" captionTargetId="figure-307@11.[164,1445,196,1047]" captionTargetPageId="11" captionText="Figure 8. The geographical distribution of Troglophilus neglectus (arrows indicate supposed directions of spreading). (ITA-Italy; A-Austria; SLO-Slovenia; CRO-Croatia; BIH-Bosnia and Herzegovina; SRB-Serbia; MNE-Montenegro; BUL- Bulgaria; ALB-Albania; MK-Macedonia; GRE-Greece)." figureDoi="http://doi.org/10.5281/zenodo.5442339" httpUri="https://zenodo.org/record/5442339/files/figure.png" pageId="9" pageNumber="1044">Fig. 8</figureCitation>
) and neighbouring parts of
<collectingCountry id="F349AF88FFD2FFABFE97FA31FEB6F9D9" box="[271,357,1514,1536]" name="Austria" pageId="9" pageNumber="1044">Austria</collectingCountry>
and
<collectingCountry id="F349AF88FFD2FFABFE34FA31FE30F9D9" box="[428,483,1514,1536]" name="Italy" pageId="9" pageNumber="1044">Italy</collectingCountry>
. We can roughly distinguish two relatively compact units of its distribution range. The first one extends from northern parts of
<collectingCountry id="F349AF88FFD2FFABFF3CF99EFECCF982" box="[164,287,1605,1627]" name="Macedonia" pageId="9" pageNumber="1044">Macedonia</collectingCountry>
and southern parts of
<collectingCountry id="F349AF88FFD2FFABFDDFF99EFD42F982" box="[583,657,1605,1627]" name="Serbia" pageId="9" pageNumber="1044">Serbia</collectingCountry>
to northeast
<collectingCountry id="F349AF88FFD2FFABFF7DF9BFFEFCF9A3" box="[229,303,1636,1658]" name="Serbia" pageId="9" pageNumber="1044">Serbia</collectingCountry>
and western-most parts of
<collectingCountry id="F349AF88FFD2FFABFD38F9BFFCD4F9A3" box="[672,775,1636,1658]" name="Bulgaria" pageId="9" pageNumber="1044">Bulgaria</collectingCountry>
. The southern distribution borders are still not clear; it is possible that the species is present in
<collectingCountry id="F349AF88FFD2FFABFD29F97AFCD9F96E" box="[689,778,1697,1719]" name="Albania" pageId="9" pageNumber="1044">Albania</collectingCountry>
as well.
</paragraph>
<paragraph id="8BE1EF18FFD2FFABFF24F905FC0DF8CB" blockId="9.[164,779,1452,1903]" lastBlockId="9.[826,1441,1452,1810]" pageId="9" pageNumber="1044">
The second unit comprises western-most parts of the Balkans and neighbouring regions of
<collectingCountry id="F349AF88FFD2FFABFD2CF926FCD9F8CA" box="[692,778,1789,1811]" name="Austria" pageId="9" pageNumber="1044">Austria</collectingCountry>
and
<collectingCountry id="F349AF88FFD2FFABFF41F8C7FEC3F8EB" box="[217,272,1820,1842]" name="Italy" pageId="9" pageNumber="1044">Italy</collectingCountry>
. The species is present in western
<collectingCountry id="F349AF88FFD2FFABFD2EF8C7FCD4F8EB" box="[694,775,1820,1842]" name="Bosnia and Herzegovina" pageId="9" pageNumber="1044">Bosnia</collectingCountry>
, in
<collectingCountry id="F349AF88FFD2FFABFF5DF8E1FEC8F889" box="[197,283,1850,1872]" name="Croatia" pageId="9" pageNumber="1044">Croatia</collectingCountry>
from northern Dalmatia to the north and west, and widely in
<collectingCountry id="F349AF88FFD2FFABFE0AF882FE24F8B6" box="[402,503,1881,1903]" name="Slovenia" pageId="9" pageNumber="1044">Slovenia</collectingCountry>
. The eastern-most area of this unit of the species distribution range is in Slavonia and western
<collectingCountry id="F349AF88FFD2FFABFBD5FA10FB4DFA38" box="[1101,1182,1483,1505]" name="Bosnia and Herzegovina" pageId="9" pageNumber="1044">Bosnia</collectingCountry>
. However, the distribution pattern in
<collectingCountry id="F349AF88FFD2FFABFB95FA31FB88F9D9" box="[1037,1115,1514,1536]" name="Bosnia and Herzegovina" pageId="9" pageNumber="1044">Bosnia</collectingCountry>
is still unclear. The species is not present in western parts of
<collectingCountry id="F349AF88FFD2FFABFB7DF9D3FAE2F9C7" box="[1253,1329,1544,1566]" name="Serbia" pageId="9" pageNumber="1044">Serbia</collectingCountry>
, in
<collectingCountry id="F349AF88FFD2FFABFAFEF9D2FC41F9E5" name="Serbia" pageId="9" pageNumber="1044">Montenegro</collectingCountry>
, in
<collectingCountry id="F349AF88FFD2FFABFC51F9FCFB88F9E4" box="[969,1115,1575,1597]" name="Bosnia and Herzegovina" pageId="9" pageNumber="1044">Herzegovina</collectingCountry>
, and in south and central Dalmatia. It seems that it is absent from east
<collectingCountry id="F349AF88FFD2FFABFCA2F9BFFC5BF9A3" box="[826,904,1636,1658]" name="Bosnia and Herzegovina" pageId="9" pageNumber="1044">Bosnia</collectingCountry>
too. Southern populations appear only at higher altitudes. There are no data on the sympatric appearance of this species with other
<taxonomicName id="4C5E949BFFD2FFABFA8AF97AFC59F90F" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="1044" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92A330AFFD2FFABFA8AF97AFA72F96F" box="[1298,1441,1697,1718]" italics="true" pageId="9" pageNumber="1044">Troglophilus</emphasis>
species
</taxonomicName>
in the eastern unit of its distribution range. In the western unit, sympatry with
<taxonomicName id="4C5E949BFFD2FFABFA9FF904FAADF92A" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[1287,1406,1758,1780]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="1044" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="B92A330AFFD2FFABFA9FF904FAADF92A" box="[1287,1406,1758,1780]" italics="true" pageId="9" pageNumber="1044">T. cavicola</emphasis>
</taxonomicName>
is very common.
</paragraph>
<paragraph id="8BE1EF18FFD2FFA8FCA2F8E1FE78F9C7" blockId="9.[826,1441,1850,1903]" lastBlockId="10.[144,759,1483,1903]" lastPageId="10" lastPageNumber="1045" pageId="9" pageNumber="1044">
<emphasis id="B92A330AFFD2FFABFCA2F8E1FC75F896" box="[826,934,1850,1871]" italics="true" pageId="9" pageNumber="1044">Remarks:</emphasis>
The citation of this species from the Imanthia Cave near Nousa in
<collectingCountry id="F349AF88FFD2FFABFB3CF882FB21F8B7" box="[1188,1266,1881,1902]" name="Greece" pageId="9" pageNumber="1044">Greece</collectingCountry>
most probably refers to the species
<taxonomicName id="4C5E949BFFD1FFA8FE13FA17FE30FA38" authority="Karaman &amp; Hammouti &amp; Pavićević &amp; Kiefer &amp; Horvatović &amp; Seitz, 2011" authorityName="Karaman &amp; Hammouti &amp; Pavićević &amp; Kiefer &amp; Horvatović &amp; Seitz" authorityYear="2011" box="[395,483,1484,1505]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="zorae" status="sp. nov.">
<emphasis id="B92A330AFFD1FFA8FE13FA17FE30FA38" box="[395,483,1484,1505]" italics="true" pageId="10" pageNumber="1045">T. zorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198E71FFD1FFA8FE76FA10FD94FA39" box="[494,583,1483,1504]" pageId="10" pageNumber="1045" rank="species">sp. nov.</taxonomicNameLabel>
This was concluded according to the illustration of a male tenth tergite in
<bibRefCitation id="EFCF92E9FFD1FFA8FE9EF9D2FE75F9C7" author="Maran J" box="[262,422,1544,1566]" pageId="10" pageNumber="1045" pagination="387 - 393" refId="ref11056" refString="Maran J. 1958. Beitrag zur Kenntnis der geographischen Variabilitat von Troglophilus neglectus Krauss. Orthoptera Raphidophoridae. Acta Entomologica Musei Nationalis Pragae 32: 387 - 393." type="journal article" year="1958">Maran (1958)</bibRefCitation>
.
</paragraph>
<caption id="DF21BF90FFD1FFA8FF08FAFFFBD1FA56" ID-DOI="http://doi.org/10.5281/zenodo.5442337" ID-Zenodo-Dep="5442337" httpUri="https://zenodo.org/record/5442337/files/figure.png" pageId="10" pageNumber="1045" startId="10.[144,223,1316,1335]" targetBox="[303,1263,196,1286]" targetPageId="10">
<paragraph id="8BE1EF18FFD1FFA8FF08FAFFFBD1FA56" blockId="10.[144,1422,1316,1423]" pageId="10" pageNumber="1045">
<emphasis id="B92A330AFFD1FFA8FF08FAFFFF2BFAEE" bold="true" box="[144,248,1316,1335]" pageId="10" pageNumber="1045">Figure 7.</emphasis>
AB,
<taxonomicName id="4C5E949BFFD1FFA8FEDAFAFFFDFDFAEE" box="[322,558,1316,1335]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B92A330AFFD1FFA8FEDAFAFFFDFDFAEE" box="[322,558,1316,1335]" italics="true" pageId="10" pageNumber="1045">Troglophilus neglectus</emphasis>
</taxonomicName>
, male 17 mm, phallus with titillator (arrow) (Smarna gora, Slovenia): A, dorsal view; B, lateral view. CD,
<taxonomicName id="4C5E949BFFD1FFA8FE29FA9AFD61FA8D" authorityName="KARNY" authorityYear="1907" box="[433,690,1345,1364]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="ovuliformis">
<emphasis id="B92A330AFFD1FFA8FE29FA9AFD61FA8D" box="[433,690,1345,1364]" italics="true" pageId="10" pageNumber="1045">Troglophilus ovuliformis</emphasis>
</taxonomicName>
, male 20 mm, phallus with titillator (arrow) (G. Seoca, Montenegro): C, dorsal view; D, lateral view. E,
<taxonomicName id="4C5E949BFFD1FFA8FD95FA84FD4FFAA8" authorityName="KARNY" authorityYear="1907" box="[525,668,1374,1393]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="ovuliformis">
<emphasis id="B92A330AFFD1FFA8FD95FA84FD4FFAA8" box="[525,668,1374,1393]" italics="true" pageId="10" pageNumber="1045">T. ovuliformis</emphasis>
</taxonomicName>
, male 20 mm, titillator (arrow), dorsal view (Island Pag, Croatia). F,
<taxonomicName id="4C5E949BFFD1FFA8FF08FAA7FECCFA56" authorityName="KARNY" authorityYear="1907" box="[144,287,1404,1423]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="ovuliformis">
<emphasis id="B92A330AFFD1FFA8FF08FAA7FECCFA56" box="[144,287,1404,1423]" italics="true" pageId="10" pageNumber="1045">T. ovuliformis</emphasis>
</taxonomicName>
, male 20 mm, titillator (arrow), dorsal view (D. Seoca, Montenegro).
</paragraph>
</caption>
<paragraph id="8BE1EF18FFD1FFA8FF30F9FCFE48F92D" blockId="10.[144,759,1483,1903]" pageId="10" pageNumber="1045">
Morphological characteristics of
<taxonomicName id="4C5E949BFFD1FFA8FD85F9FCFF1BF982" authorityName="Maran" authorityYear="1958" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="vlasinensis">
<emphasis id="B92A330AFFD1FFA8FD85F9FCFF1BF982" italics="true" pageId="10" pageNumber="1045">T. neglectus vlasinensis</emphasis>
</taxonomicName>
from south-east
<collectingCountry id="F349AF88FFD1FFA8FE09F99EFE0EF982" box="[401,477,1605,1627]" name="Serbia" pageId="10" pageNumber="1045">Serbia</collectingCountry>
, used for differentiation in
<bibRefCitation id="EFCF92E9FFD1FFA8FF2AF9BFFE87F9A3" author="Maran J" box="[178,340,1636,1658]" pageId="10" pageNumber="1045" pagination="387 - 393" refId="ref11056" refString="Maran J. 1958. Beitrag zur Kenntnis der geographischen Variabilitat von Troglophilus neglectus Krauss. Orthoptera Raphidophoridae. Acta Entomologica Musei Nationalis Pragae 32: 387 - 393." type="journal article" year="1958">Maran (1958)</bibRefCitation>
, are in the boundaries of the variability of
<taxonomicName id="4C5E949BFFD1FFA8FE9DF958FE5AF941" box="[261,393,1667,1688]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B92A330AFFD1FFA8FE9DF958FE5AF941" box="[261,393,1667,1688]" italics="true" pageId="10" pageNumber="1045">T. neglectus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5E949BFFD1FFA8FE03F958FF60F96E" authority="neglectus" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="neglectus">
<emphasis id="B92A330AFFD1FFA8FE03F958FF60F96E" italics="true" pageId="10" pageNumber="1045">Troglophilus neglectus neglectus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E949BFFD1FFA8FE9DF979FDC9F96F" authorityName="Maran" authorityYear="1958" box="[261,538,1697,1719]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="vlasinensis">
<emphasis id="B92A330AFFD1FFA8FE9DF979FDC9F96F" box="[261,538,1697,1719]" italics="true" pageId="10" pageNumber="1045">T. neglectus vlasinensis</emphasis>
</taxonomicName>
share the same haplotype,
<collectionCode id="ED4F77DDFFD1FFA8FE8AF91BFEF1F90C" box="[274,290,1728,1749]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="1045">T</collectionCode>
neg HT1, which also suggests synonymization of these taxa.
</paragraph>
<paragraph id="8BE1EF18FFD1FFA8FF30F926FC7BF8B6" blockId="10.[144,759,1483,1903]" lastBlockId="10.[806,1421,1483,1903]" pageId="10" pageNumber="1045">
After examination of a large number of specimens, we can say with certainty that the populations of
<taxonomicName id="4C5E949BFFD1FFA8FF08F8E0FEC0F896" box="[144,275,1850,1872]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="1045" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="B92A330AFFD1FFA8FF08F8E0FEC0F896" box="[144,275,1850,1872]" italics="true" pageId="10" pageNumber="1045">T. neglectus</emphasis>
</taxonomicName>
in eastern
<collectingCountry id="F349AF88FFD1FFA8FE33F8E1FE26F889" box="[427,501,1850,1872]" name="Serbia" pageId="10" pageNumber="1045">Serbia</collectingCountry>
are parthenogenetic. We suppose that it is the same in the neighbouring parts of
<collectingCountry id="F349AF88FFD1FFA8FC08FA10FC27FA38" box="[912,1012,1483,1505]" name="Bulgaria" pageId="10" pageNumber="1045">Bulgaria</collectingCountry>
where this species is present. As females dominate in the material from the south populations, this led us to the supposition that parthenogenetic populations also prevail in these areas (the only males were collected from a cave north of Skopje). The assumption that the marginal populations are often parthenogenetic imposes a conclusion that these areas were secondarily inhabited from the west. However, in keeping with the recent distribution, we conclude quite the opposite. The primary distribution range of this species must be related to the southern parts of
<collectingCountry id="F349AF88FFD1FFA8FBBBF8C7FBBEF8EB" box="[1059,1133,1820,1842]" name="Serbia" pageId="10" pageNumber="1045">Serbia</collectingCountry>
and neighbouring areas. The explanation for such an opinion is given in the Discussion.
</paragraph>
</subSubSection>
</treatment>
</document>