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<document id="4E292A7092B38A373469E5D75A583C45" ID-CLB-Dataset="29889" ID-DOI="10.11646/zootaxa.4434.1.10" ID-GBIF-Dataset="06e27461-4a99-402f-ae42-eb81da865394" ID-ISSN="1175-5326" ID-Zenodo-Dep="1290884" ID-ZooBank="B22116EA-DF00-4C6D-93FB-D40427809537" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529082540665" checkinUser="plazi" docAuthor="Gnezdilov, Dimir M." docDate="2018" docId="03DE941A747DFFF7FF14FB48FEFFF87A" docLanguage="en" docName="zootaxa.4434.1.10.pdf" docOrigin="Zootaxa 4434 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fowlerium Gnezdilov 2018, gen. n." docType="treatment" docUuid="FBE917F4-60E8-4AF1-A318-FCCE46DEDE78" docUuidSource="ZooBank" docVersion="8" lastPageNumber="163" masterDocId="FFE7EC627478FFF2FF83FFB5FF80FFBD" masterDocTitle="To the revision of the genus Thionia Stål (Hemiptera, Fulgoroidea, Issidae), with description of new genera and new subtribe" masterLastPageNumber="170" masterPageNumber="158" pageNumber="163" updateTime="1698549558969" updateUser="ExternalLinkService">
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<mods:title id="CEB9265B40E9B1B9FA29511D442307D9">To the revision of the genus Thionia Stål (Hemiptera, Fulgoroidea, Issidae), with description of new genera and new subtribe</mods:title>
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<mods:namePart id="7CCBFA894035F755BD811DD10B36488A">Gnezdilov, Dimir M.</mods:namePart>
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<mods:date id="AE4918FC46EF29DFB35526B6127DD657">2018</mods:date>
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<treatment id="03DE941A747DFFF7FF14FB48FEFFF87A" ID-DOI="http://doi.org/10.5281/zenodo.5981752" ID-GBIF-Taxon="145966279" ID-Zenodo-Dep="5981752" LSID="urn:lsid:zoobank.org:act:FBE917F4-60E8-4AF1-A318-FCCE46DEDE78" httpUri="http://treatment.plazi.org/id/03DE941A747DFFF7FF14FB48FEFFF87A" lastPageNumber="163" pageId="5" pageNumber="163">
<subSubSection id="C36D7687747DFFF7FF14FB48FEF6FAAA" box="[151,374,1277,1303]" pageId="5" pageNumber="163" type="nomenclature">
<paragraph id="8BC8250C747DFFF7FF14FB48FEF6FAAA" blockId="5.[151,374,1277,1303]" box="[151,374,1277,1303]" pageId="5" pageNumber="163">
<heading id="D0809260747DFFF7FF14FB48FEF6FAAA" bold="true" box="[151,374,1277,1303]" fontSize="11" level="1" pageId="5" pageNumber="163" reason="1">
<emphasis id="B903F91E747DFFF7FF14FB48FEF6FAAA" bold="true" box="[151,374,1277,1303]" pageId="5" pageNumber="163">
<taxonomicName id="4C775E8F747DFFF7FF14FB48FE9DFAAA" ID-CoL="4KXV" ID-ENA="2837359" authority="Gnezdilov, 2018" authorityName="Gnezdilov" authorityYear="2018" box="[151,285,1277,1303]" class="Insecta" family="Issidae" genus="Fowlerium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B903F91E747DFFF7FF14FB48FE9DFAAA" bold="true" box="[151,285,1277,1303]" italics="true" pageId="5" pageNumber="163">Fowlerium</emphasis>
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<taxonomicNameLabel id="A2304465747DFFF7FEA7FB48FEF6FAAA" box="[292,374,1277,1303]" pageId="5" pageNumber="163" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C36D7687747DFFF7FF14FAF0FDBEFAE7" box="[151,574,1348,1370]" pageId="5" pageNumber="163" type="materials_examined">
<paragraph id="8BC8250C747DFFF7FF14FAF0FDBEFAE7" blockId="5.[151,574,1348,1370]" box="[151,574,1348,1370]" pageId="5" pageNumber="163">
Type species:
<taxonomicName id="4C775E8F747DFFF7FEA5FAF1FDBAFAE7" authority="Fowler, 1905" authorityName="Fowler" authorityYear="1905" box="[294,570,1348,1370]" class="Insecta" family="Issidae" genus="Thionia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="species" species="naso">
<emphasis id="B903F91E747DFFF7FEA5FAF1FE2BFAE7" box="[294,427,1348,1370]" italics="true" pageId="5" pageNumber="163">Thionia naso</emphasis>
<bibRefCitation id="EFE658FD747DFFF7FE32FAF1FDBAFAE7" box="[433,570,1348,1370]" pageId="5" pageNumber="163" refId="ref5620" refString="Fowler, W. W. (1905) Family Issidae. Order Rhynchota. Suborder Hemiptera-Homoptera. Insecta (Continued). Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, 1, 113 - 129." type="journal article">Fowler, 1905</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D7687747DFFF7FF14FA33FCBEF88B" pageId="5" pageNumber="163" type="diagnosis">
<paragraph id="8BC8250C747DFFF7FF14FA33FCBEF88B" blockId="5.[151,1437,1414,1991]" pageId="5" pageNumber="163">
<emphasis id="B903F91E747DFFF7FF14FA33FE91FA22" bold="true" box="[151,273,1414,1439]" pageId="5" pageNumber="163">Diagnosis.</emphasis>
Metope elongate, with distinct median and sublateral carinae joined in one point on its upper margin (
<bibRefCitation id="EFE658FD747DFFF7FF1DFA1FFE54FA7E" box="[158,468,1450,1475]" pageId="5" pageNumber="163" refId="ref5415" refString="Doering, K. C. (1938) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part 2. The University of Kansas Science Bulletin, 25 (20), 447 - 575." type="journal article">Doering 1938, Pl. 52: 8, 10</bibRefCitation>
). Coryphe elongate, with distinct median carina running from its anterior margin to its middle; anterior margin acutely angulately convex (
<bibRefCitation id="EFE658FD747DFFF7FD73FA7BFBA9FA5A" box="[752,1065,1486,1511]" pageId="5" pageNumber="163" refId="ref5415" refString="Doering, K. C. (1938) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part 2. The University of Kansas Science Bulletin, 25 (20), 447 - 575." type="journal article">Doering 1938, Pl. 51: 2, 5</bibRefCitation>
). Second and third segments of rostrum are almost equal in length; third segment narrowing apically (
<figureCitation id="134C3989747DFFF7FC43FA47FB96F9B6" box="[960,1046,1522,1547]" captionStart="FIGURES 1718" captionStartId="9.[151,264,1811,1834]" captionTargetBox="[598,999,917,1788]" captionTargetId="figure@9.[588,1011,914,1800]" captionTargetPageId="9" captionText="FIGURES 1718. Thioniina, rostrum in lateral view. 17—Cheiloceps musca Uhler; 18—Fowlerium naso (Fowler)." httpUri="https://zenodo.org/record/1290896/files/figure.png" pageId="5" pageNumber="163">Fig. 18</figureCitation>
). Fore wings elongate, surpassing abdomen, with narrow hypocostal plate. Forewing vein sequence: R 2, furcating near to basal cell; M 2, furcating before wing middle; CuΑ 1. Clavus long, open—Pcu+Α1 runs into apex of clavus. Pcu fusing Α1 after middle of clavus. Hind wings 3-lobed, with deep cubital and anal clefts and without coupling lobe on its anterior margin. CuΑ2 and CuP not fused apically, but connected by several transverse veins (
<figureCitation id="134C3989747DFFF7FB85F930FB0BF923" box="[1030,1163,1669,1694]" captionStart="FIGURES 1213" captionStartId="7.[151,264,1555,1578]" captionTargetBox="[487,1103,495,1517]" captionTargetId="figure@7.[464,1123,477,1534]" captionTargetPageId="7" captionText="FIGURES 1213. Fowlerium spp., hind wing. 12—F. naso (Fowler); 13—F. productum (Van Duzee)." httpUri="https://zenodo.org/record/1290892/files/figure.png" pageId="5" pageNumber="163">Figs 12, 13</figureCitation>
). Pcu and Α1.1 not fused medially, free. Hindwing sequence: R 2 r-m 1 M 1 m-cua 1 CuΑ 2 cua-cup 2-5 CuP 1 cup-pcu 1 Pcu 3 pcu-a1 1 Α1 2 Α2 24. Hind tibia with 2 lateral spines. First metatarsomere with a whole row of spines arranged in arc. Ventral aedeagal hooks slightly furcating subapically (
<bibRefCitation id="EFE658FD747DFFF7FD20F94CFC71F8AF" box="[675,1009,1785,1810]" pageId="5" pageNumber="163" refId="ref5415" refString="Doering, K. C. (1938) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part 2. The University of Kansas Science Bulletin, 25 (20), 447 - 575." type="journal article">Doering, 1938, Pl. 55: 1, 7, 8</bibRefCitation>
). Ovipositor with rounded gonoplacs. Hind margin of VII sternum with large semicircular process.
</paragraph>
</subSubSection>
<subSubSection id="C36D7687747DFFF7FF44F8F4FEFFF87A" pageId="5" pageNumber="163" type="etymology">
<paragraph id="8BC8250C747DFFF7FF44F8F4FAB0F8C3" blockId="5.[151,1437,1414,1991]" pageId="5" pageNumber="163">
<emphasis id="B903F91E747DFFF7FF44F8F4FECEF8E7" bold="true" box="[199,334,1857,1882]" pageId="5" pageNumber="163">Etymology.</emphasis>
Genus named in honor of famous British entomologist William Weeks Fowler (
<date id="FFC903CC747DFFF7FB58F8F7FADCF8E7" box="[1243,1372,1858,1882]" pageId="5" pageNumber="163" value="1849" valueMax="1923">18491923</date>
) who describing
<taxonomicName id="4C775E8F747DFFF7FE91F8D0FE25F8C3" authorityName="Fowler" authorityYear="1905" box="[274,421,1893,1918]" class="Insecta" family="Issidae" genus="Thionia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="species" species="naso">
<emphasis id="B903F91E747DFFF7FE91F8D0FE25F8C3" box="[274,421,1893,1918]" italics="true" pageId="5" pageNumber="163">Thionia naso</emphasis>
</taxonomicName>
had doubts on its generic position already (
<bibRefCitation id="EFE658FD747DFFF7FC0DF8D0FBDBF8C3" box="[910,1115,1893,1918]" pageId="5" pageNumber="163" refId="ref5620" refString="Fowler, W. W. (1905) Family Issidae. Order Rhynchota. Suborder Hemiptera-Homoptera. Insecta (Continued). Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, 1, 113 - 129." type="journal article">Fowler 1905: 125</bibRefCitation>
). Neuter in gender.
</paragraph>
<paragraph id="8BC8250C747DFFF7FF44F83CFEFFF87A" blockId="5.[151,1437,1414,1991]" pageId="5" pageNumber="163">
<emphasis id="B903F91E747DFFF7FF44F83CFEE7F81F" bold="true" box="[199,359,1929,1954]" pageId="5" pageNumber="163">Composition.</emphasis>
In addition to type species
<taxonomicName id="4C775E8F747DFFF7FD1DF83CFBABF81F" authority="Van Duzee, 1908" authorityName="Van Duzee" authorityYear="1908" box="[670,1067,1929,1954]" class="Insecta" family="Issidae" genus="Thionia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="B903F91E747DFFF7FD1DF83CFCE1F81F" box="[670,865,1929,1954]" italics="true" pageId="5" pageNumber="163">Thionia producta</emphasis>
<bibRefCitation id="EFE658FD747DFFF7FCEAF83FFBABF81F" box="[873,1067,1930,1954]" pageId="5" pageNumber="163" refId="ref6870" refString="Van Duzee, E. P. (1908) Studies in North American Fulgoridae. Proceedings of the Academy of Natural Sciences of Philadelphia, 1907, 467 - 498." type="proceedings paper">Van Duzee, 1908</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C775E8F747DFFF7FBE0F83EFAE7F81F" authority="Doering, 1941" authorityName="Doering" authorityYear="1941" box="[1123,1383,1929,1954]" class="Insecta" family="Issidae" genus="Thionia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B903F91E747DFFF7FBE0F83EFB3DF81F" box="[1123,1213,1931,1954]" italics="true" pageId="5" pageNumber="163">T. acuta</emphasis>
<bibRefCitation id="EFE658FD747DFFF7FB47F83CFAE7F81F" box="[1220,1383,1929,1954]" pageId="5" pageNumber="163" refId="ref5463" refString="Doering, K. C. (1941) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part 4. The University of Kansas Science Bulletin, 27 (10), 185 - 233." type="journal article">Doering, 1941</bibRefCitation>
</taxonomicName>
also belong to this genus.
</paragraph>
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</treatment>
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