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<document id="7C120873F8E9144F2198E8EFEDE66FFE" ID-DOI="10.11646/zootaxa.4751.1.6" ID-GBIF-Dataset="c76f145d-543d-48ea-ba43-91b5dc4528e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3711860" ID-ZooBank="D8104E82-8754-484F-9D03-51FA65F481B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584349247993" checkinUser="plazi" docAuthor="Rodrigues, Patrícia E. S., Golovatch, Sergei I., Ott, Ricardo &amp; Ro- Drigues, Everton N. L." docDate="2020" docId="03DE9004FC5F547F4BA993A7FE68F851" docLanguage="en" docName="zootaxa.4751.1.6.pdf" docOrigin="Zootaxa 4751 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catharosoma pedritense Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues 2020, new species" docType="treatment" docUuid="8185375E-63EC-4F62-B84A-6D2BB574679C" docUuidSource="ZooBank" docVersion="6" lastPageNumber="125" masterDocId="FFE7E87CFC5954794B3E917FFFF7FFA3" masterDocTitle="Three new species of the millipede genus Catharosoma Silvestri, 1897 from southern Brazil, with new records and a clarified identity of Catharosoma intermedium (Carl, 1902) (Diplopoda: Polydesmida: Paradoxosomatidae)" masterLastPageNumber="130" masterPageNumber="119" pageNumber="125" updateTime="1698729150640" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C69F48B24B04F3422E2261B85E37CCB0">Three new species of the millipede genus Catharosoma Silvestri, 1897 from southern Brazil, with new records and a clarified identity of Catharosoma intermedium (Carl, 1902) (Diplopoda: Polydesmida: Paradoxosomatidae)</mods:title>
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<mods:namePart id="D06E500857F0B1563F28ED0512337AC3">Rodrigues, Patrícia E. S.</mods:namePart>
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<mods:namePart id="C1F7A413D175661218A1AD43106BA511">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="CCD2EFE760E9B072FE00945FBEA59E16">Ott, Ricardo</mods:namePart>
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<mods:namePart id="000A594D69E611F2AA00F321BEB0ACFB">Ro- Drigues, Everton N. L.</mods:namePart>
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<mods:title id="7284CBE693C1138075AD146E55844FAE">Zootaxa</mods:title>
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<mods:date id="4CDB2EA93DE78248432E78D18D8DFC78">2020</mods:date>
<mods:detail id="A7DDC447540F53EE4D031944C6B2DD90" type="pubDate">
<mods:number id="EA6B7D4A9803CB6CCC72A18A6754EDEF">2020-03-16</mods:number>
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<mods:number id="7C776DB329722B50148FB08D522DC11F">4751</mods:number>
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<mods:identifier id="49865F855B739E68158241F5311A6B4D" type="DOI">10.11646/zootaxa.4751.1.6</mods:identifier>
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<treatment id="03DE9004FC5F547F4BA993A7FE68F851" ID-DOI="http://doi.org/10.5281/zenodo.3718093" ID-GBIF-Taxon="162575796" ID-Zenodo-Dep="3718093" LSID="urn:lsid:zoobank.org:act:8185375E-63EC-4F62-B84A-6D2BB574679C" httpUri="http://treatment.plazi.org/id/03DE9004FC5F547F4BA993A7FE68F851" lastPageNumber="125" pageId="6" pageNumber="125">
<subSubSection id="C36D7299FC5F547F4BA993A7FEE3FCB4" pageId="6" pageNumber="125" type="nomenclature">
<paragraph id="8BC82112FC5F547F4BA993A7FEE3FCB4" blockId="6.[151,1437,585,791]" pageId="6" pageNumber="125">
<emphasis id="B903FD00FC5F547F4BA993A7FDA2FD51" bold="true" box="[151,597,728,755]" pageId="6" pageNumber="125">
<taxonomicName id="4C775A91FC5F547F4BA993A7FE49FD50" ID-CoL="32a85c3c-b550-4d70-b5c9-cc5c0a6affd5" authority="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues, 2020" authorityName="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues" authorityYear="2020" box="[151,446,728,755]" class="Diplopoda" family="Paradoxosomatidae" genus="Catharosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="pedritense" status="sp. nov.">
<emphasis id="B903FD00FC5F547F4BA993A7FE49FD50" bold="true" box="[151,446,728,755]" italics="true" pageId="6" pageNumber="125">Catharosoma pedritense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230407BFC5F547F4AFB93A6FDA2FD51" box="[453,597,728,755]" pageId="6" pageNumber="125" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation id="134C3D97FC5F547F4BA99383FEE3FCB4" box="[151,276,764,791]" captionStart-0="FIGURES 18, 19" captionStart-1="FIGURES 24, 25" captionStartId-0="6.[151,264,1469,1494]" captionStartId-1="7.[151,264,1874,1899]" captionTargetBox-0="[296,1292,1065,1442]" captionTargetBox-1="[151,1436,152,1848]" captionTargetId-0="figure@6.[295,1293,1064,1444]" captionTargetId-1="figure@7.[253,1334,272,1324]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 18, 19. Catharosoma pedritense n. sp., holotype. 18, Anterior part of body, lateral view. 19, Posterior part of body, lateral view." captionText-1="FIGURES 24, 25. Catharosoma pedritense n. sp., holotype. 24, Left gonopod, dorsal view. 25, Structural details of parabasal lobe of solenophore. Scale bars: 0.1 mm (24), 0.01 mm (25)." figureDoi-0="http://doi.org/10.5281/zenodo.3711872" figureDoi-1="http://doi.org/10.5281/zenodo.3711874" httpUri-0="https://zenodo.org/record/3711872/files/figure.png" httpUri-1="https://zenodo.org/record/3711874/files/figure.png" pageId="6" pageNumber="125">Figs 1825</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D7299FC5F547F4BA9923AFDDFFC21" pageId="6" pageNumber="125" type="materials_examined">
<paragraph id="8BC82112FC5F547F4BA9923AFDDFFC21" blockId="6.[151,1437,836,1043]" pageId="6" pageNumber="125">
<emphasis id="B903FD00FC5F547F4BA9923AFEB7FCFC" bold="true" box="[151,320,836,863]" pageId="6" pageNumber="125">Type material</emphasis>
: Male
<typeStatus id="54CC9FB0FC5F547F4AAD923BFE02FCFD" box="[403,501,836,862]" pageId="6" pageNumber="125" type="holotype">holotype</typeStatus>
from Dom Pedrito (
<geoCoordinate id="EE4347D5FC5F547F49DF923BFC98FCFC" box="[737,879,836,863]" degrees="54" direction="west" minutes="40" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="23" value="-54.673058">54º4023”W</geoCoordinate>
,
<geoCoordinate id="EE4347D5FC5F547F4843923BFBF5FCFC" box="[893,1026,836,863]" degrees="30" direction="south" minutes="58" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="58" value="-30.982779">30º5858”S</geoCoordinate>
),
<collectingRegion id="49B3EFF0FC5F547F4F26923BFB04FCFD" box="[1048,1267,836,862]" country="Brazil" name="Rio Grande do Sul" pageId="6" pageNumber="125">Rio Grande do Sul</collectingRegion>
,
<collectingCountry id="F3606182FC5F547F4E3E923BFAB0FCFD" box="[1280,1351,836,862]" name="Brazil" pageId="6" pageNumber="125">Brazil</collectingCountry>
,
<date id="FFC907D2FC5F547F4E6B923BFF25FC21" pageId="6" pageNumber="125" value="1978-06-01">
01.
<collectionCode id="ED66B9D7FC5F547F4E48923BFA6FFCFD" box="[1398,1432,836,862]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="125">VI</collectionCode>
<materialsCitation id="3B1F2B4FFC5F547F4EA6923BFF25FC21" ID-GBIF-Occurrence="2576149817" collectionCode="MCN" collectorName="C. J. Becker" pageId="6" pageNumber="125" specimenCode="MCN 131" specimenCount="1">. 1978</materialsCitation>
</date>
,
<collectorName id="268244C4FC5F547F4BE39217FE91FC21" box="[221,358,872,899]" pageId="6" pageNumber="125">C. J. Becker</collectorName>
leg. (
<specimenCode id="DBD18969FC5F547F4A9B9217FDEDFC21" box="[421,538,872,898]" collectionCode="MCN" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="6" pageNumber="125" type="Herbarium">MCN 131</specimenCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D7299FC5F547F4BF992F3FADDFBB0" pageId="6" pageNumber="125" type="diagnosis">
<paragraph id="8BC82112FC5F547F4BF992F3FADDFBB0" blockId="6.[151,1437,836,1043]" pageId="6" pageNumber="125">
<emphasis id="B903FD00FC5F547F4BF992F3FECDFC04" bold="true" box="[199,314,908,935]" pageId="6" pageNumber="125">Diagnosis</emphasis>
:
<taxonomicName id="4C775A91FC5F547F4A7492F3FDA9FC05" authority="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues, 2020" authorityName="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues" authorityYear="2020" box="[330,606,908,934]" class="Diplopoda" family="Paradoxosomatidae" genus="Catharosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="pedritense" status="sp. nov.">
<emphasis id="B903FD00FC5F547F4A7492F3FDA9FC05" box="[330,606,908,934]" italics="true" pageId="6" pageNumber="125">Catharosoma pedritense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230407BFC5F547F495B92F2FD55FC04" box="[613,674,909,935]" pageId="6" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
strongly resembles
<taxonomicName id="4C775A91FC5F547F48BB92F2FBDFFC05" authority="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues, 2020" authorityName="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues" authorityYear="2020" box="[901,1064,909,935]" class="Diplopoda" family="Paradoxosomatidae" genus="Catharosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="promatense" status="sp. nov.">
<emphasis id="B903FD00FC5F547F48BB92F2FBDFFC05" box="[901,1064,909,935]" italics="true" pageId="6" pageNumber="125">C. promatense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230407BFC5F547F4F1192F2FB9BFC04" box="[1071,1132,909,935]" pageId="6" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
by the shape of the sternal structures (Figs 12, 20), but differs by the paramedian process and tufts of setae on sternum between coxae of leg pair 6 longer, colour pattern without two paramedian stripes and by the shape of the parabasal lobe (p) of the solenophore which is quadrate (Figs 23-25)—this shape being different from all species known in the genus.
</paragraph>
</subSubSection>
<caption id="DF08719AFC5F547F4BA994C2FEE3FA59" ID-DOI="http://doi.org/10.5281/zenodo.3711872" ID-Zenodo-Dep="3711872" httpUri="https://zenodo.org/record/3711872/files/figure.png" pageId="6" pageNumber="125" startId="6.[151,264,1469,1494]" targetBox="[296,1292,1065,1442]" targetPageId="6">
<paragraph id="8BC82112FC5F547F4BA994C2FEE3FA59" blockId="6.[151,1437,1469,1530]" pageId="6" pageNumber="125">
<emphasis id="B903FD00FC5F547F4BA994C2FEB9FA75" bold="true" box="[151,334,1469,1494]" pageId="6" pageNumber="125">FIGURES 18, 19</emphasis>
.
<taxonomicName id="4C775A91FC5F547F4A6594C2FDA2FA76" authority="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues, 2020" authorityName="Rodrigues &amp; Golovatch &amp; Ott &amp; Ro- Drigues" authorityYear="2020" box="[347,597,1469,1493]" class="Diplopoda" family="Paradoxosomatidae" genus="Catharosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="pedritense" status="sp. nov.">
<emphasis id="B903FD00FC5F547F4A6594C2FDA2FA76" box="[347,597,1469,1493]" italics="true" pageId="6" pageNumber="125">Catharosoma pedritense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230407BFC5F547F496594C1FD63FA76" box="[603,660,1470,1493]" pageId="6" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
, holotype.
<emphasis id="B903FD00FC5F547F483B94C2FCE9FA75" bold="true" box="[773,798,1469,1494]" pageId="6" pageNumber="125">18</emphasis>
, Anterior part of body, lateral view.
<emphasis id="B903FD00FC5F547F4FAC94C2FB5CFA75" bold="true" box="[1170,1195,1469,1494]" pageId="6" pageNumber="125">19</emphasis>
, Posterior part of body, lateral view.
</paragraph>
</caption>
<subSubSection id="C36D7299FC5F547F4BF99758FE68F851" pageId="6" pageNumber="125" type="description">
<paragraph id="8BC82112FC5F547F4BF99758FECBF86D" blockId="6.[151,1437,1575,2034]" pageId="6" pageNumber="125">
<emphasis id="B903FD00FC5F547F4BF99758FEA6F9E1" bold="true" box="[199,337,1575,1602]" pageId="6" pageNumber="125">Description</emphasis>
: Male
<typeStatus id="54CC9FB0FC5F547F4AA09757FDF7F9E1" box="[414,512,1576,1602]" pageId="6" pageNumber="125" type="holotype">holotype</typeStatus>
(
<figureCitation id="134C3D97FC5F547F49309757FD7AF9E1" box="[526,653,1576,1602]" captionStart="FIGURES 18, 19" captionStartId="6.[151,264,1469,1494]" captionTargetBox="[296,1292,1065,1442]" captionTargetId="figure@6.[295,1293,1064,1444]" captionTargetPageId="6" captionText="FIGURES 18, 19. Catharosoma pedritense n. sp., holotype. 18, Anterior part of body, lateral view. 19, Posterior part of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3711872" httpUri="https://zenodo.org/record/3711872/files/figure.png" pageId="6" pageNumber="125">Figs 18, 19</figureCitation>
). Length
<quantity id="4C8F8CF7FC5F547F49CB9757FCB4F9E1" box="[757,835,1576,1602]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="125" unit="mm" value="23.0">23 mm</quantity>
and maximum width of midbody
<quantity id="4C8F8CF7FC5F547F4F849757FAE5F9E1" box="[1210,1298,1576,1602]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="125" unit="mm" value="1.9">1.9 mm</quantity>
.
<emphasis id="B903FD00FC5F547F4E259758FA6BF9E1" bold="true" box="[1307,1436,1575,1602]" pageId="6" pageNumber="125">Coloration</emphasis>
(in alcohol): generally yellowish, but legs and antennae pale yellow. White, axial, dorsal line present. Legs densely setose, tibiae swollen ventrally with brushes starting in leg pair 10 until two last pairs. Male coxae of leg pair 2 with a prominent gonapophysis.
<emphasis id="B903FD00FC5F547F4AF397ECFDB7F90D" bold="true" box="[461,576,1683,1710]" pageId="6" pageNumber="125">Antennae</emphasis>
relatively long,
<emphasis id="B903FD00FC5F547F49C997EAFCCBF90D" box="[759,828,1685,1710]" italics="true" pageId="6" pageNumber="125">in situ</emphasis>
reaching behind body ring 4 dorsally. Frons and clypeus smooth and setose.
<emphasis id="B903FD00FC5F547F4A9D97C7FDDAF971" bold="true" box="[419,557,1720,1746]" pageId="6" pageNumber="125">Integument</emphasis>
smooth and shining. Ozopores lateral, poorly visible. Limbus thin and entire. Paraterga strongly reduced, arcuate sulci on body rings 24, missing thereafter. Epiproct pointed, rather long, with a dorsal and a ventral pair of spinnerets. Hypoproct subtriangular and relatively short.
<emphasis id="B903FD00FC5F547F4F5B9780FA6AF8B9" bold="true" box="[1125,1437,1791,1818]" pageId="6" pageNumber="125">Sternal structures of male</emphasis>
(Fig. 20). Sternal structures with longer paramedian process and tufts of setae on sternum between coxae of leg pair 6. Postgonopodal sterna with paramedian spines.
<emphasis id="B903FD00FC5F547F49CF9638FC9DF8C1" bold="true" box="[753,874,1863,1890]" pageId="6" pageNumber="125">Gonopods</emphasis>
(Figs 2125) complex, subcircular and elongated. Gonocoxite subcylindrical. A usual, densely setose, prefemoral portion somewhat shorter than a large femorite. Postfemoral portion excavated. Solenophore (sph) with a parabasal lobe (p) unciform in ventral view and quadrate in dorsal view.
</paragraph>
<paragraph id="8BC82112FC5F547F4BF996A7FE68F851" blockId="6.[151,1437,1575,2034]" box="[199,415,2008,2034]" pageId="6" pageNumber="125">Female: Unknown.</paragraph>
</subSubSection>
</treatment>
</document>