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<document id="05299F73AE93E076D10B1D491DA96D64" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFF9AAF35D1DBDA0BFE1A1661" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Trichillum Harold 1868" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="47" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="949A3F2E33E90A6C80B8581E9FBC0C57">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
</mods:titleInfo>
<mods:name id="6382D4E5A182BA3B2A7656C7D937DE11" type="personal">
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<mods:namePart id="5DD9DF4E3735AF4C4ED19613581B7440">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="308C971A3FC07F23E3ADFF220B4C31C8">Zootaxa</mods:title>
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<mods:part id="6124EAA89CDE56C89A3DC13865DDB2F5">
<mods:date id="6CAA60BD838562FD8913AA8176F830D5">2008</mods:date>
<mods:detail id="03BCA1D30863CCCCC845338ABFA7645C" type="pubDate">
<mods:number id="90BBB9A4E152670671ED908ACEBF00C3">2008-12-05</mods:number>
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<mods:number id="E867D8D2A42056E2F8A06EFDF6E0D7EB">1955</mods:number>
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<mods:classification id="46E91F657A36707829FA922677BBF8A9">journal article</mods:classification>
<mods:identifier id="684C74F1DD7701958718A941CCCB2B4C" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03D287CEFF9AAF35D1DBDA0BFE1A1661" ID-DOI="http://doi.org/10.5281/zenodo.5242051" ID-Zenodo-Dep="5242051" LSID="urn:lsid:plazi:treatment:03D287CEFF9AAF35D1DBDA0BFE1A1661" httpUri="http://treatment.plazi.org/id/03D287CEFF9AAF35D1DBDA0BFE1A1661" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C3616553FF9AAF34D1DBDA0BFE4B16C1" box="[151,493,1407,1433]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BC436D8FF9AAF34D1DBDA0BFE4B16C1" blockId="46.[151,493,1407,1433]" box="[151,493,1407,1433]" pageId="46" pageNumber="47">
<heading id="D08C81B4FF9AAF34D1DBDA0BFE4B16C1" bold="true" box="[151,493,1407,1433]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B90FEACAFF9AAF34D1DBDA0BFE4B16C1" bold="true" box="[151,493,1407,1433]" pageId="46" pageNumber="47">
17.
<taxonomicName id="4C7B4D5BFF9AAF34D18FDA0BFE4B16C1" ID-CoL="7Z67" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[195,493,1407,1433]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9AAF34D18FDA0BFEE216C1" bold="true" box="[195,324,1407,1433]" italics="true" pageId="46" pageNumber="47">Trichillum</emphasis>
<bibRefCitation id="EFEA4B29FF9AAF34D007DA0BFE4B16C1" author="Harold, E." box="[331,493,1407,1433]" pageId="46" pageNumber="47" pagination="33 - 55" refId="ref48745" refString="Harold, E. (1868) Die Choeridiiden-Gattungen Uroxys und Trichillum. Coleopterologische Hefte, 3, 33 - 55." type="journal article" year="1868">Harold, 1868</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9AAF34D1DBDABEFE46158C" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BC436D8FF9AAF34D1DBDABEFE46158C" blockId="46.[151,1436,1482,2028]" pageId="46" pageNumber="47">
<emphasis id="B90FEACAFF9AAF34D1DBDABEFEBF16BC" bold="true" box="[151,281,1482,1508]" pageId="46" pageNumber="47">Diagnosis:</emphasis>
Body form oval (
<figureCitation id="13402A5DFF9AAF34D0BEDABEFD2216BC" box="[498,644,1482,1508]" captionStart="FIGURES 31–38" captionStartId="58.[151,269,1935,1959]" captionTargetBox="[222,1364,194,1911]" captionTargetId="figure-57@58.[222,1365,194,1911]" captionTargetPageId="58" captionText="FIGURES 31–38. Dorsal views. 31. Pereiraidium almeidai, ♀; 32. Scatimus ovatus; 33. Scatrichus bicarinatus; 34. Silvinha unica; 35. Trichillidium quadridens; 36. Trichillidium brevisetosum; 37. Trichillum heydeni; 38. Trichillum pauliani." pageId="46" pageNumber="47">Figs. 37–38</figureCitation>
); color grey to black, shiny or not, always lacking metallic sheen. Clypeal teeth variable in shape, but always two (or indicated by anterior sinuosity) and always continuing clypeal border (Figs. 98, 100, 102). Pseudoepipleura strongly sinuate on anterior half and strongly angled near metacoxa; basal half with longitudinal carina parallel to pseudoepipleural-epipleural division (
<figureCitation id="13402A5DFF9AAF34D5A5D936FA961504" box="[1257,1328,1602,1628]" captionStart="FIGURES 5–11" captionStartId="54.[151,269,1929,1953]" captionTargetBox="[223,1320,197,1895]" captionTargetId="figure-60@54.[203,1372,197,1917]" captionTargetPageId="54" captionText="FIGURES 5–11. Left epipleuron and pseudoepipleuron, ventral view. 5. Trichillum externepunctatum; 6. Onoreidium cristatum; 7. Scatrichus bicarinatus; 8. Pedaridium hirsutum; 9. Silvinha unica; 10. Besourenga minutus; 11. Degallieridium lilliputanum. Scale = 1.0 mm." pageId="46" pageNumber="47">Fig. 5</figureCitation>
). Aedeagus with short subrectangular parameres (Figs. 99, 101) and phallobase very short (about twice as long as wide); spermatheca particularly spiral shaped, spermathecal duct sclerotized and feebly differentiated from spermatheca itself (
<figureCitation id="13402A5DFF9AAF34D037D9CEFE74158C" box="[379,466,1722,1748]" captionStart="FIGURES 12–14" captionStartId="55.[151,269,1373,1397]" captionTargetBox="[173,1369,194,1301]" captionTargetId="figure-46@55.[169,1419,194,1350]" captionTargetPageId="55" captionText="FIGURES 12–14. Spermatheca (receptaculum seminis), and attached part of spermathecal duct, and spermathecal gland; 12. Pedaridium hirsutum; 13. Trichillum externepunctatum; 14. Eutrichillum hirsutum. Scale = 0.2 mm." pageId="46" pageNumber="47">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9AAF35D18AD996FE4F12AA" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="8BC436D8FF9AAF35D18AD996FD1012FA" blockId="46.[151,1436,1482,2028]" lastBlockId="47.[151,1436,152,1337]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B90FEACAFF9AAF34D18AD996FEC615A4" bold="true" box="[198,352,1762,1788]" pageId="46" pageNumber="47">Description:</emphasis>
Size 2.2–5.0 mm, body oval (
<figureCitation id="13402A5DFF9AAF34D39FD996FCC315A4" box="[723,869,1762,1788]" captionStart="FIGURES 31–38" captionStartId="58.[151,269,1935,1959]" captionTargetBox="[222,1364,194,1911]" captionTargetId="figure-57@58.[222,1365,194,1911]" captionTargetPageId="58" captionText="FIGURES 31–38. Dorsal views. 31. Pereiraidium almeidai, ♀; 32. Scatimus ovatus; 33. Scatrichus bicarinatus; 34. Silvinha unica; 35. Trichillidium quadridens; 36. Trichillidium brevisetosum; 37. Trichillum heydeni; 38. Trichillum pauliani." pageId="46" pageNumber="47">Figs. 37–38</figureCitation>
), color grey or brown to black, always lacking metallic sheen. Clypeus with two strong teeth separated by U- or V-shaped emargination. Clypeogenal and frontoclypeal sutures indistinct. Clypeal margin lateral to clypeal teeth strongly curved or angled, becoming straight nearing clypeogenal border, which is straight or slightly sinuate. Gena straight or slightly curved mesally to lateral angle (Figs. 98, 100, 102). Pronotum lacking anterior or posterior beads (Fig. 102); disc with indistinct lateral callosity. Hypomeron posteriorly with longitudinal lateral carina; mesoepimera with transverse anterior carina. Metasternal disc glabrous. Elytral interstriae with uniseriate setose punctures, mesal striae slightly widened and deepened at apical declivity. Pseudoepipleura strongly narrowed to apex, angled near metacoxa, anterior half sinuous; glabrous, with longitudinal carina on anterior half, ending in shallow medial concavity (
<figureCitation id="13402A5DFF9BAF35D03CDF9CFE1F125A" box="[368,441,232,258]" captionStart="FIGURES 5–11" captionStartId="54.[151,269,1929,1953]" captionTargetBox="[223,1320,197,1895]" captionTargetId="figure-60@54.[203,1372,197,1917]" captionTargetPageId="54" captionText="FIGURES 5–11. Left epipleuron and pseudoepipleuron, ventral view. 5. Trichillum externepunctatum; 6. Onoreidium cristatum; 7. Scatrichus bicarinatus; 8. Pedaridium hirsutum; 9. Silvinha unica; 10. Besourenga minutus; 11. Degallieridium lilliputanum. Scale = 1.0 mm." pageId="47" pageNumber="48">Fig. 5</figureCitation>
). Protibiae with three strong lateral teeth, distributed along apical four fifths of tibial length; with scale-like setae ventrally to teeth base. Parameres shorter than half of phallobase length; rectangular, apically flattened (Figs. 99, 101); phallobase only twice as long as its width. Coxites large, laminar, asymmetrical or centrally fused. Spermatheca spiral shaped, spermathecal duct sclertotized and difficult to differentiate from spermatheca itself (
<figureCitation id="13402A5DFF9BAF35D31DDEFCFD0E12FA" box="[593,680,392,418]" captionStart="FIGURES 12–14" captionStartId="55.[151,269,1373,1397]" captionTargetBox="[173,1369,194,1301]" captionTargetId="figure-46@55.[169,1419,194,1350]" captionTargetPageId="55" captionText="FIGURES 12–14. Spermatheca (receptaculum seminis), and attached part of spermathecal duct, and spermathecal gland; 12. Pedaridium hirsutum; 13. Trichillum externepunctatum; 14. Eutrichillum hirsutum. Scale = 0.2 mm." pageId="47" pageNumber="48">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BC436D8FF9BAF35D18ADEDBFE4F12AA" blockId="47.[151,1436,152,1337]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADEDBFE621291" bold="true" box="[198,452,431,457]" pageId="47" pageNumber="48">Sexual dimorphism:</emphasis>
Males with ventrally directed apicomesal tooth on protibiae and concave metasternal disc (convex in females).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9BAF35D18ADE8BFC241142" box="[198,898,511,538]" pageId="47" pageNumber="48" type="type_taxon">
<paragraph id="8BC436D8FF9BAF35D18ADE8BFC241142" blockId="47.[151,1436,152,1337]" box="[198,898,511,538]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADE8BFECC1141" bold="true" box="[198,362,511,537]" pageId="47" pageNumber="48">
<typeStatus id="54C0887AFF9BAF35D18ADE8BFEA51141" box="[198,259,511,537]" pageId="47" pageNumber="48">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7B4D5BFF9BAF35D03EDD74FD561142" ID-CoL="5893D" authority="Harold, 1868" authorityName=": Harold" authorityYear="1868" box="[370,752,512,538]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="heydeni">
<emphasis id="B90FEACAFF9BAF35D03EDD74FDE81141" box="[370,590,512,537]" italics="true" pageId="47" pageNumber="48">Trichillum heydeni</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D31ADD74FD561142" author="Harold, E." box="[598,752,512,538]" pageId="47" pageNumber="48" pagination="33 - 55" refId="ref48745" refString="Harold, E. (1868) Die Choeridiiden-Gattungen Uroxys und Trichillum. Coleopterologische Hefte, 3, 33 - 55." type="journal article" year="1868">Harold, 1868</bibRefCitation>
</taxonomicName>
(monotypy)
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9BAF35D18ADD53FC6D11CA" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BC436D8FF9BAF35D18ADD53FC6D11CA" blockId="47.[151,1436,152,1337]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADD53FECF1119" bold="true" box="[198,361,551,577]" pageId="47" pageNumber="48">Distribution:</emphasis>
<collectingCountry id="F36C7648FF9BAF35D03FDD5CFE18111A" box="[371,446,552,578]" name="Brazil" pageId="47" pageNumber="48">Brazil</collectingCountry>
, French
<collectingCountry id="F36C7648FF9BAF35D36ADD5CFD22111A" box="[550,644,552,578]" name="Guyana" pageId="47" pageNumber="48">Guyana</collectingCountry>
,
<collectingCountry id="F36C7648FF9BAF35D3DEDD5CFCAC111A" box="[658,778,552,578]" name="Argentina" pageId="47" pageNumber="48">Argentina</collectingCountry>
,
<collectingCountry id="F36C7648FF9BAF35D254DD5CFCD7111A" box="[792,881,552,578]" name="Bolivia" pageId="47" pageNumber="48">Bolivia</collectingCountry>
,
<collectingCountry id="F36C7648FF9BAF35D233DD5CFC4A111A" box="[895,1004,552,578]" name="Paraguay" pageId="47" pageNumber="48">Paraguay</collectingCountry>
and
<collectingCountry id="F36C7648FF9BAF35D565DD5CFB36111A" box="[1065,1168,552,578]" name="Uruguay" pageId="47" pageNumber="48">Uruguay</collectingCountry>
, always eastern of the Andes. Comprises the Humid
<collectingCountry id="F36C7648FF9BAF35D0B2DD24FDFA1132" box="[510,604,592,618]" name="Guyana" pageId="47" pageNumber="48">Guyana</collectingCountry>
, Amapa, Para, Pantanal, Tapajos-Xingu and Roraima provinces of the Amazonian subregion, and the entire Chacoan and Parana subregions.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9BAF35D18ADDEBFBFE164A" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8BC436D8FF9BAF35D18ADDEBFE1B10F2" blockId="47.[151,1436,152,1337]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADDEBFEE711E1" bold="true" box="[198,321,671,697]" pageId="47" pageNumber="48">Remarks:</emphasis>
<taxonomicName id="4C7B4D5BFF9BAF35D005DDD4FE6511E1" box="[329,451,672,697]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9BAF35D005DDD4FE6511E1" box="[329,451,672,697]" italics="true" pageId="47" pageNumber="48">Trichillum</emphasis>
</taxonomicName>
in its present definition is undoubtedly sister to
<taxonomicName id="4C7B4D5BFF9BAF35D2B4DDD4FB2511E1" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1016,1155,672,697]" class="Insecta" family="Scarabaeidae" genus="Besourenga" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9BAF35D2B4DDD4FB2511E1" box="[1016,1155,672,697]" italics="true" pageId="47" pageNumber="48">Besourenga</emphasis>
</taxonomicName>
(see remarks under that genus), and is supported by the following apomorphies: presence of a pseudoepipleural carina parallel to the pseudoepipleuron-epipleuron delimitation on anterior half; mesotibiae and metatibiae strongly widened apically, with apical width less than third of tibial length; coxites large and asymmetrical, or if symmetrical, centrally fused as single piece; spermathecal duct uniformly sclerotized, not differentiated from the spermatheca itself. The strong sinuosity of the anterior half of pseudoepipleura is shared with
<taxonomicName id="4C7B4D5BFF9BAF35D502DC1CFB0810D9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1102,1198,872,897]" class="Insecta" family="Scarabaeidae" genus="Silvinha" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9BAF35D502DC1CFB0810D9" box="[1102,1198,872,897]" italics="true" pageId="47" pageNumber="48">Silvinha</emphasis>
</taxonomicName>
, however it is probably a symplesiomorphy.
</paragraph>
<paragraph id="8BC436D8FF9BAF35D18ADCC3FBFE164A" blockId="47.[151,1436,152,1337]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADCC3FEC91089" bold="true" box="[198,367,951,977]" pageId="47" pageNumber="48">Composition:</emphasis>
<taxonomicName id="4C7B4D5BFF9BAF35D034DCCCFE541089" box="[376,498,952,977]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9BAF35D034DCCCFE541089" box="[376,498,952,977]" italics="true" pageId="47" pageNumber="48">Trichillum</emphasis>
</taxonomicName>
in the present sense is almost equivalent to the subgenus
<taxonomicName id="4C7B4D5BFF9BAF35D5EDDCCCFABD1089" box="[1185,1307,952,977]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF9BAF35D5EDDCCCFABD1089" box="[1185,1307,952,977]" italics="true" pageId="47" pageNumber="48">Trichillum</emphasis>
</taxonomicName>
as defined by Martínez, except for the inclusion of
<taxonomicName id="4C7B4D5BFF9BAF35D321DC94FD4D10A1" authorityName="Balthasar" authorityYear="1939" box="[621,747,992,1017]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B90FEACAFF9BAF35D321DC94FD4D10A1" box="[621,747,992,1017]" italics="true" pageId="47" pageNumber="48">T. pauliani</emphasis>
</taxonomicName>
, which Martínez placed (1969) in
<emphasis id="B90FEACAFF9BAF35D533DC94FEB51779" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="4C7B4D5BFF9BAF35D533DC94FAB110A1" authorityName="Martinez" authorityYear="1969" box="[1151,1303,992,1017]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">Eutrichillum</taxonomicName>
.
<taxonomicName id="4C7B4D5BFF9BAF35D46EDC94FEB51779" authorityName=": Martinez" authorityYear="1969" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="adjunctum">Trichillum adjunctum</taxonomicName>
</emphasis>
and
<taxonomicName id="4C7B4D5BFF9BAF35D003DB7CFE691779" authorityName="Balthasar" authorityYear="1939" box="[335,463,1032,1057]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B90FEACAFF9BAF35D003DB7CFE691779" box="[335,463,1032,1057]" italics="true" pageId="47" pageNumber="48">T. pauliani</emphasis>
</taxonomicName>
form a somewhat isolated group within the genus, distinguished by the dull grey color and the uniquely centrally fused coxites; that form a large vaginal plaque continuously sclerotized with the spermathecal duct and spermatheca. Besides the
<typeStatus id="54C0887AFF9BAF35D25ADB2CFCEE172A" box="[790,840,1112,1138]" pageId="47" pageNumber="48">type</typeStatus>
species, this genus contains
<taxonomicName id="4C7B4D5BFF9BAF35D5E6DB2CFE5E17C2" authority="Preudhomme de Borre, 1880" authorityName=": Preudhomme de Borre" authorityYear="1880" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="externepunctatum">
<emphasis id="B90FEACAFF9BAF35D5E6DB2CFA3A1729" box="[1194,1436,1112,1137]" italics="true" pageId="47" pageNumber="48">T. externepunctatum</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D1DBDBF4FE5E17C2" author="Preudhomme de Borre, F. P. C. A." box="[151,504,1152,1178]" pageId="47" pageNumber="48" pagination="xxvii - xxviii" refId="ref50508" refString="Preudhomme de Borre, F. P. C. A. (1880) Description d'une espece nouvelle du genre Trichillum Harold (Coprides; Choeridiides). Compte Rendue de la Sceance de la Societe Entomologique de Belgique (Annales de la Societe Entomologique de Belgique), 23, xxvii - xxviii." type="journal article" year="1880">Preudhomme de Borre, 1880</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D34BDBF4FCBB17C2" authority="Saylor, 1935" authorityName=": Saylor" authorityYear="1935" box="[519,797,1152,1178]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B90FEACAFF9BAF35D34BDBF4FDDE17C1" box="[519,632,1152,1177]" italics="true" pageId="47" pageNumber="48">T. arrowi</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D3CDDBF4FCBB17C2" author="Saylor, L. W." box="[641,797,1152,1178]" pageId="47" pageNumber="48" pagination="207 - 208" refId="ref50785" refString="Saylor, L. W. (1935) New species of Trichillum (Col. Copridae). Stylops, 4, 207 - 208." type="journal article" year="1935">Saylor, 1935</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D260DBF4FBD117C2" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[812,1143,1152,1178]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B90FEACAFF9BAF35D260DBF4FC0817C1" box="[812,942,1152,1177]" italics="true" pageId="47" pageNumber="48">T. pauliani</emphasis>
Balthasar, 1939
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D5CADBF4FF73179A" authority="Balthasar, 1942" authorityName=": Balthasar" authorityYear="1942" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="depilatum">
<emphasis id="B90FEACAFF9BAF35D5CADBF4FABA17C1" box="[1158,1308,1152,1177]" italics="true" pageId="47" pageNumber="48">T. depilatum</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D46BDBF4FF73179A" author="Balthasar, V." pageId="47" pageNumber="48" pagination="36 - 44" refId="ref47394" refString="Balthasar, V. (1942) Neue Scarabeiden aus Sud-Amerika. 72. Beitrag zur Kenntnis der Scarabeiden (Col.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 39, 36 - 44." type="journal article" year="1942">Balthasar, 1942</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D1A8DBDCFDE2179A" authority="Martinez, 1969" authorityName=": Martinez" authorityYear="1969" box="[228,580,1192,1218]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="adjunctum">
<emphasis id="B90FEACAFF9BAF35D1A8DBDCFE261799" box="[228,384,1192,1217]" italics="true" pageId="47" pageNumber="48">T. adjunctum</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D0C6DBDCFDE2179A" author="Martinez, A." box="[394,580,1192,1218]" pageId="47" pageNumber="48" pagination="119 - 147" refId="ref49692" refString="Martinez, A. (1969) Notas para una monografia del genero Trichillum Harold, 1868 (Col. Scarabaeinae-Coprini). Revista de la Sociedad Mexicana de Historia Natural, 28, 119 - 147." type="journal article" year="1969">Martínez, 1969</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D31FDBDCFC33179A" authority="Martinez, 1969" authorityName=": Martinez" authorityYear="1969" box="[595,917,1192,1218]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="halffteri">
<emphasis id="B90FEACAFF9BAF35D31FDBDCFD771799" box="[595,721,1192,1217]" italics="true" pageId="47" pageNumber="48">T. halffteri</emphasis>
<bibRefCitation id="EFEA4B29FF9BAF35D390DBDCFC33179A" author="Martinez, A." box="[732,917,1192,1218]" pageId="47" pageNumber="48" pagination="119 - 147" refId="ref49692" refString="Martinez, A. (1969) Notas para una monografia del genero Trichillum Harold, 1868 (Col. Scarabaeinae-Coprini). Revista de la Sociedad Mexicana de Historia Natural, 28, 119 - 147." type="journal article" year="1969">Martínez, 1969</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D2E8DBDCFAE1179A" authority="Verdu &amp; Galante, 1998" authorityName="Verdu &amp; Galante" authorityYear="1998" box="[932,1351,1192,1218]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="morellii">
<emphasis id="B90FEACAFF9BAF35D2E8DBDCFB861799" box="[932,1056,1192,1217]" italics="true" pageId="47" pageNumber="48">T. morellii</emphasis>
Verdú &amp; Galante, 1998
</taxonomicName>
;
<taxonomicName id="4C7B4D5BFF9BAF35D41ADBDCFEA117B1" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tischechkini">
<emphasis id="B90FEACAFF9BAF35D41ADBDCFEA117B1" italics="true" pageId="47" pageNumber="48">T. tischechkini</emphasis>
</taxonomicName>
<bibRefCitation id="EFEA4B29FF9BAF35D043DBA4FDD217B2" author="Vaz-de-Mello, F. Z. &amp; Genier, F." box="[271,628,1232,1258]" pageId="47" pageNumber="48" pagination="41 - 52" refId="ref51259" refString="Vaz-de-Mello, F. Z. &amp; Genier, F. (2005) Lectotype designations, new synonymies, and new species in the genera Trichillum Harold and Pedaridium Harold (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 1038, 41 - 52." type="journal article" year="2005">Vaz-de-Mello &amp; Génier, 2005</bibRefCitation>
;
<taxonomicName id="4C7B4D5BFF9BAF35D3CDDBA4FCE617B1" authorityName="Vaz-de-Mello &amp; Genier" authorityYear="2005" box="[641,832,1232,1257]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pseudoarrowi">
<emphasis id="B90FEACAFF9BAF35D3CDDBA4FCE617B1" box="[641,832,1232,1257]" italics="true" pageId="47" pageNumber="48">T. pseudoarrowi</emphasis>
</taxonomicName>
<bibRefCitation id="EFEA4B29FF9BAF35D205DBA4FB0B17B2" author="Vaz-de-Mello, F. Z. &amp; Genier, F." box="[841,1197,1232,1258]" pageId="47" pageNumber="48" pagination="41 - 52" refId="ref51259" refString="Vaz-de-Mello, F. Z. &amp; Genier, F. (2005) Lectotype designations, new synonymies, and new species in the genera Trichillum Harold and Pedaridium Harold (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 1038, 41 - 52." type="journal article" year="2005">Vaz-de-Mello &amp; Génier, 2005</bibRefCitation>
;
<taxonomicName id="4C7B4D5BFF9BAF35D5F6DBA4FE65164A" authority="Vazde-Mello &amp; Genier, 2005" authorityName="Vazde-Mello &amp; Genier" authorityYear="2005" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cordobense">
<emphasis id="B90FEACAFF9BAF35D5F6DBA4FAF817B1" box="[1210,1374,1232,1257]" italics="true" pageId="47" pageNumber="48">T. cordobense</emphasis>
Vazde-Mello &amp; Génier, 2005
</taxonomicName>
; and at least 30 new species currently under description.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF9BAF35D18ADA6BFE1A1661" box="[198,444,1311,1337]" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BC436D8FF9BAF35D18ADA6BFE1A1661" blockId="47.[151,1436,152,1337]" box="[198,444,1311,1337]" pageId="47" pageNumber="48">
<emphasis id="B90FEACAFF9BAF35D18ADA6BFE1A1661" bold="true" box="[198,444,1311,1337]" pageId="47" pageNumber="48">Material examined:</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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