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<document id="AEABF26B3482F290DE0627C78553CE0D" ID-DOI="10.5252/zoosystema2024v46a29" ID-ISSN="1638-9387" ID-Zenodo-Dep="14132095" ID-ZooBank="B8FB5A89-4811-4DB1-819B-442067652F3E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1731487024113" checkinUser="plazi" docAuthor="L., Peter K., Castro, Ng Peter, Rodríguez, Paula A. &amp; Moreno" docDate="2024" docId="03D287C27D5F9C152A38FDA4FA1DF812" docLanguage="en" docName="zoosystema.2024.46.29.pdf" docOrigin="Zoosystema 46 (29)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a29.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Diuplax L., Castro, Rodríguez &amp; Moreno, 2024, n. gen." docType="treatment" docVersion="2" lastPageNumber="741" masterDocId="FFEBFFBA7D559C1F2E22FFB2FFB2FFFB" masterDocTitle="Notes on the taxonomy of Frevillea A. Milne-Edwards, 1880 (Crustacea, Brachyura, Goneplacoidea), with the description of a new genus for F. sigsbei A. Milne-Edwards, 1880" masterLastPageNumber="748" masterPageNumber="733" pageNumber="741" updateTime="1731487453058" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B3E308CF1B7B6B039E6E8EC0387F5B69">Notes on the taxonomy of Frevillea A. Milne-Edwards, 1880 (Crustacea, Brachyura, Goneplacoidea), with the description of a new genus for F. sigsbei A. Milne-Edwards, 1880</mods:title>
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<mods:namePart id="4359A3C8DC6F03834C2B56303F2ADAFB">L., Peter K.</mods:namePart>
<mods:affiliation id="1E0DD4A6ED342F9354C50D5460A6B581">Lee Kong Chian Natural History Museum, Conservatory Drive, National University of Singapore, Singapore 117377 (Republic of Singapore) Biological Sciences Department, California State Polytechnic University, Pomona, CA 91768 (United States)</mods:affiliation>
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<mods:affiliation id="538806FB384D3B8D1420547B2785DFBB">Direction générale déléguée aux case postale 53, 57 rue Cuvier, F- 75231 Paris cedex paula. martin-lefevre @ mnhn. fr (corresponding</mods:affiliation>
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<taxonomicName id="4C7B4D577D5F9C152A38FDA4FBC2FDCB" authorityName="L. &amp; Castro &amp; Rodríguez &amp; Moreno" authorityYear="2024" box="[1050,1136,534,560]" class="Malacostraca" family="Goneplacidae" genus="Diuplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEAC67D5F9C152A38FDA4FBC2FDCB" box="[1050,1136,534,560]" italics="true" pageId="10" pageNumber="741">Diuplax</emphasis>
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. —
<taxonomicName id="4C7B4D577D5F9C152DF4FD3CFAD2FD5C" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[982,1376,654,679]" class="Malacostraca" family="Euryplacidae" genus="Frevillea" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="species" species="sigsbei">
<emphasis id="B90FEAC67D5F9C152DF4FD3CFBD4FD5D" box="[982,1126,654,678]" italics="true" pageId="10" pageNumber="741">Frevillea sigsbei</emphasis>
A.
<bibRefCitation id="EFEA4B257D5F9C152AAAFD3CFAD2FD5C" author="MILNE-EDWARDS A." box="[1160,1376,654,679]" pageId="10" pageNumber="741" pagination="1 - 68" refId="ref13222" refString="MILNE-EDWARDS A. 1880. - Etudes preliminaires sur les Crustaces 1 ere partie, in Reports on the results of dredging under the supervision of Alexander Agassiz in the Gulf of Mexico and in the Caribbean Sea 1877 ' 78 ' 79 by the U. S. Coast Survey Steamer &quot; Blake &quot; Lieut. - Commander C. D. Sigsbee U. S. N. and Commander ̿. R. Bartlett U. S. N. commanding VIII. Bulletin of the Museum of Comparative Zoology at Harvard College 8: 1 - 68, pls 1, 2. https: // www. biodiversitylibrary. org / page / 30184918" type="book chapter" year="1880">Milne-Edwards, 1880</bibRefCitation>
</taxonomicName>
, by present designation.
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<subSubSection id="C361655F7D5F9C152D0FFD50FA30FCB2" pageId="10" pageNumber="741" type="etymology">
<paragraph id="8BC436D47D5F9C152D0FFD50FA30FCB2" blockId="10.[813,1455,738,841]" pageId="10" pageNumber="741">
<smallCapsWord id="8D22A0087D5F9C152D0FFD50FC14FD03" baselines="755,756" box="[813,934,738,761]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="10" pageNumber="741">ETYMOLOGY</smallCapsWord>
. — The name is derived from the Latin
<emphasis id="B90FEAC67D5F9C152B10FD50FAC1FD01" box="[1330,1395,738,762]" italics="true" pageId="10" pageNumber="741">diutius</emphasis>
for “a long time,” in arbitrary combination with “-
<emphasis id="B90FEAC67D5F9C152AC1FD4EFAB8FCEF" box="[1251,1290,764,788]" italics="true" pageId="10" pageNumber="741">plax</emphasis>
”, the ending for many goneplacid taxa, alluding to the long time it took to discover the true generic affinities of
<taxonomicName id="4C7B4D577D5F9C152A1CFC80FB3DFCB2" authorityName="A. Milne-Edwards" authorityYear="1880" box="[1086,1167,817,841]" class="Malacostraca" family="Euryplacidae" genus="Frevillea" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="species" species="sigsbei">
<emphasis id="B90FEAC67D5F9C152A1CFC80FB3DFCB2" box="[1086,1167,817,841]" italics="true" pageId="10" pageNumber="741">F. sigsbei</emphasis>
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. The gender is feminine.
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<paragraph id="8BC436D47D5F9C152D0FFCD8FBB7FABF" blockId="10.[811,1457,874,1348]" pageId="10" pageNumber="741">
<smallCapsWord id="8D22A0087D5F9C152D0FFCD8FC28FC7B" baselines="891,892" box="[813,922,874,897]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="10" pageNumber="741">DIAGNOSIS</smallCapsWord>
. — Carapace broadly trapezoidal (
<figureCitation id="13402A517D5F9C152AC8FCD8FA11FC79" box="[1258,1443,874,898]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5A, B, D, F, G</figureCitation>
); anterolateral tooth adjacent to pronounced external orbital tooth, separated by deep cleft (
<figureCitation id="13402A517D5F9C152A33FC2DFB7EFC4C" box="[1041,1228,926,951]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Figs 5A, B, D, F, G</figureCitation>
;
<figureCitation id="13402A517D5F9C152AF7FC2DFAA2FC4C" box="[1237,1296,927,951]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">6A, B</figureCitation>
); posterior margin of epistome relatively wide transversely, margin almost straight (
<figureCitation id="13402A517D5F9C152D10FC61FC28FC17" box="[818,922,979,1004]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">Fig. 6A, B</figureCitation>
); ocular peduncle relatively short,
<emphasis id="B90FEAC67D5F9C152AD2FC66FAB2FC10" box="[1264,1280,980,1003]" italics="true" pageId="10" pageNumber="741">c.</emphasis>
2 times length of cornea (
<figureCitation id="13402A517D5F9C152D5FFC5CFBF5FBFD" box="[893,1095,1006,1030]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Figs 5A, B, D, F, G</figureCitation>
;
<figureCitation id="13402A517D5F9C152A70FC5CFB3DFBFD" box="[1106,1167,1006,1030]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">6A, B</figureCitation>
); male thoracic sternites 1-4 relatively wide (
<figureCitation id="13402A517D5F9C152DEBFBBAFBA7FBDB" box="[969,1045,1032,1056]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5E</figureCitation>
); male thoracic sternite 8 visible as triangular plate when pleon closed; pleonal somite 1 reaching coxa of fourth ambulatory leg; episternite 7 partially overlaps episternite
<quantity id="4C839B317D5F9C152D0FFBEAFCE7FB8B" box="[813,853,1112,1136]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="741" unit="in" value="8.0">8 in</quantity>
ventral view, penis visible in oblique view, between narrow channel between episternites 7 and 8 (
<figureCitation id="13402A517D5F9C152A9CFBC0FB4EFB71" box="[1214,1276,1138,1162]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">Fig. 6</figureCitation>
C-E); male pleon relatively narrower transversely, especially somites 3-6 (
<figureCitation id="13402A517D5F9C152B7AFB3FFA16FB5E" box="[1368,1444,1165,1189]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5E</figureCitation>
); adult male cheliped short, relatively stout (
<figureCitation id="13402A517D5F9C152ACAFB15FAE8FB44" box="[1256,1370,1191,1215]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5D, E</figureCitation>
); fourth ambulatory dactylus spatuliform (
<figureCitation id="13402A517D5F9C152AABFB70FAABFB21" box="[1161,1305,1217,1242]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5A, B, D</figureCitation>
); G1 relatively short, broadly C-shaped, evenly distally tapering gradually to sharp tip, opening subdistal, large (
<figureCitation id="13402A517D5F9C152AB7FB45FAB2FAF4" box="[1173,1280,1270,1295]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">Fig. 6F, G</figureCitation>
); G2 longer than G1; flagellum long, longer than basal article, distal part strongly recurved (
<figureCitation id="13402A517D5F9C152DB3FA99FC4BFABF" box="[913,1017,1323,1348]" captionStart="FIG" captionStartId="14.[132,143,1108,1126]" captionTargetBox="[136,1403,235,1045]" captionTargetId="figure-530@14.[132,1426,215,1058]" captionTargetPageId="14" captionText="FIG. 6. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); B-I, ♂ (13.0 × 8.5 mm) (MNHN- IU-2014-11828); A, frontal view of cephalothorax;B, left side of carapace showing orbit and third maxilliped (left side);C, ♂ thoracic sternite 8 and pleonal somites 1 and 2; D, sternopleonal cavity; E, penis and coxa of fourth ambulatory leg (tilted at angle to see penis); F, left G1; G, distal part of left G1; H, left G2; I, distal part of left G2. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: fig. 3); B, D, F-I, after Guinot (1969b: figs 63, 68, 71, 72); C, E, by P. K. L. Ng. Abbreviations: cx5, coxa of fourth ambulatory leg; pe, penis; so1, 2, pleonal somites 1 and 2, respectively; st5-8, thoracic sternites 5-8." figureDoi="http://doi.org/10.5281/zenodo.14132115" httpUri="https://zenodo.org/record/14132115/files/figure.png" pageId="10" pageNumber="741">Fig. 6H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361655F7D5F9C152D0FFAC6FA1DF812" pageId="10" pageNumber="741" type="discussion">
<paragraph id="8BC436D47D5F9C152D0FFAC6FC23FA77" blockId="10.[812,1458,1396,1772]" box="[813,913,1396,1422]" pageId="10" pageNumber="741">
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</paragraph>
<paragraph id="8BC436D47D5F9C152D0FFA21FB13F917" blockId="10.[812,1458,1396,1772]" pageId="10" pageNumber="741">
The differences between
<taxonomicName id="4C7B4D577D5F9C152A72FA21FB1EFA56" authorityName="L. &amp; Castro &amp; Rodríguez &amp; Moreno" authorityYear="2024" box="[1104,1196,1427,1453]" class="Malacostraca" family="Goneplacidae" genus="Diuplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEAC67D5F9C152A72FA21FB1EFA56" box="[1104,1196,1427,1453]" italics="true" pageId="10" pageNumber="741">Diuplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57BD7D5F9C152A94FA26FABAFA55" box="[1206,1288,1428,1454]" pageId="10" pageNumber="741" rank="genus">n. gen.</taxonomicNameLabel>
and
<emphasis id="B90FEAC67D5F9C152B64FA21FCD1FA36" italics="true" pageId="10" pageNumber="741">
<taxonomicName id="4C7B4D577D5F9C152B64FA21FA1DFA56" authorityName="Leach" authorityYear="1814" box="[1350,1455,1427,1453]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" sensu="stricto">Goneplax</taxonomicName>
<taxonomicNameLabel id="A23C57BD7D5F9C152D0FFA06FCD1FA36" box="[813,867,1460,1485]" pageId="10" pageNumber="741" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
are pronounced (see
<tableCitation id="C6F9036F7D5F9C152A71FA01FB16FA36" box="[1107,1188,1459,1485]" captionStart="TABLE" captionStartId="12.[510,521,219,237]" captionText="TABLE 1. — Differences between Goneplax s. str. and Diuplax n. gen." pageId="10" pageNumber="741">Table 1</tableCitation>
). Secondary differences observed between
<taxonomicName id="4C7B4D577D5F9C152DDAFA61FBE3FA16" authorityName="L. &amp; Castro &amp; Rodríguez &amp; Moreno" authorityYear="2024" box="[1016,1105,1491,1517]" class="Malacostraca" family="Goneplacidae" genus="Diuplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEAC67D5F9C152DDAFA61FBE3FA16" box="[1016,1105,1491,1517]" italics="true" pageId="10" pageNumber="741">Diuplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57BD7D5F9C152A7AFA61FB16FA16" box="[1112,1188,1491,1517]" pageId="10" pageNumber="741" rank="genus">n. gen.</taxonomicNameLabel>
and
<emphasis id="B90FEAC67D5F9C152AF9FA61FACFFA16" box="[1243,1405,1491,1517]" italics="true" pageId="10" pageNumber="741">
<taxonomicName id="4C7B4D577D5F9C152AF9FA61FAF2FA16" authorityName="Leach" authorityYear="1814" box="[1243,1344,1491,1517]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" sensu="stricto">Goneplax</taxonomicName>
<taxonomicNameLabel id="A23C57BD7D5F9C152B65FA66FACFFA16" box="[1351,1405,1492,1517]" pageId="10" pageNumber="741" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
that may be of generic value are: the cheliped merus of the former is short, stout, dorsal margin of merus possessing a low submedian tubercle (
<figureCitation id="13402A517D5F9C152A4BF980FB72F9B6" box="[1129,1216,1586,1613]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5B</figureCitation>
) (elongated, slender, with a distinct short submedian spine on dorsal margin in
<taxonomicName id="4C7B4D577D5F9C152D0EF9C0FC23F977" authorityName="Leach" authorityYear="1814" box="[812,913,1650,1676]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEAC67D5F9C152D0EF9C0FC23F977" box="[812,913,1650,1676]" italics="true" pageId="10" pageNumber="741">Goneplax</emphasis>
</taxonomicName>
,
<figureCitation id="13402A517D5F9C152D80F9C0FC44F977" box="[930,1014,1650,1676]" captionStart="FIG" captionStartId="15.[133,144,1358,1376]" captionTargetBox="[132,1455,213,1311]" captionTargetId="figure-345@15.[132,1455,213,1316]" captionTargetPageId="15" captionText="FIG. 7. — Goneplax rhomboides (Linnaeus, 1758), ♂ (36.5 × 22.6 mm) (ZRC 2010.0069), Scotland. A, dorsal view of carapace; B, frontal view of cephalothorax; C, left third maxilliped; D, anterior thoracic sternum and pleon; E, posterior thoracic sternum and pleon; F, right merus of cheliped (inner view); G, left fourth ambulatory leg. Credits: photographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132121" httpUri="https://zenodo.org/record/14132121/files/figure.png" pageId="10" pageNumber="741">Fig. 7F</figureCitation>
); and the dorsal margin of the meri of the ambulatory legs have each show a subdistal angle but is not spiniform (
<figureCitation id="13402A517D5F9C152DD2F900FB39F937" box="[1008,1163,1714,1741]" captionStart="FIG" captionStartId="13.[132,143,1939,1957]" captionTargetBox="[132,1455,228,1885]" captionTargetId="figure-17@13.[132,1455,215,1887]" captionTargetPageId="13" captionText="FIG. 5. — Diuplax sigsbei (A. Milne-Edwards, 1880) n. comb.: A-C, lectotype ovigerous ♀ (15.3 × 9.5 mm) (MCZ-IZ-CRU-10991); D, E, paralectotype ♂ (14.1 × 8.4 mm) (MCZ-IZ-CRU-10992); F, ♂ (13.0 × 8.5 mm) (MNHN-IU-2014-11828); G, ♀ (10.8 × 7.3 mm) (MNHN-IU-2014-11828): A, B, D, F, G, overall dorsal view; C, E, ventral view of cephalothorax and pleon. Credits: A, after A. Milne-Edwards &amp; Bouvier (1923: pl. 6, fig. 2); B, C, photographs by Alana Rivera, MCZ; D-G, pho- tographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132113" httpUri="https://zenodo.org/record/14132113/files/figure.png" pageId="10" pageNumber="741">Fig. 5A, B, D</figureCitation>
) (with a distinct subdistal spine in
<emphasis id="B90FEAC67D5F9C152DA9F960FB9FF917" box="[907,1069,1746,1772]" italics="true" pageId="10" pageNumber="741">
<taxonomicName id="4C7B4D577D5F9C152DA9F960FC42F917" authorityName="Leach" authorityYear="1814" box="[907,1008,1746,1772]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus" sensu="stricto">Goneplax</taxonomicName>
<taxonomicNameLabel id="A23C57BD7D5F9C152DDAF961FB9FF917" box="[1016,1069,1747,1772]" pageId="10" pageNumber="741" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="13402A517D5F9C152A19F960FB21F917" box="[1083,1171,1746,1772]" captionStart="FIG" captionStartId="15.[133,144,1358,1376]" captionTargetBox="[132,1455,213,1311]" captionTargetId="figure-345@15.[132,1455,213,1316]" captionTargetPageId="15" captionText="FIG. 7. — Goneplax rhomboides (Linnaeus, 1758), ♂ (36.5 × 22.6 mm) (ZRC 2010.0069), Scotland. A, dorsal view of carapace; B, frontal view of cephalothorax; C, left third maxilliped; D, anterior thoracic sternum and pleon; E, posterior thoracic sternum and pleon; F, right merus of cheliped (inner view); G, left fourth ambulatory leg. Credits: photographs by P. K. L. Ng." figureDoi="http://doi.org/10.5281/zenodo.14132121" httpUri="https://zenodo.org/record/14132121/files/figure.png" pageId="10" pageNumber="741">Fig. 7G</figureCitation>
).
</paragraph>
<paragraph id="8BC436D47D5F9C152D0FF8AAFA1DF812" blockId="10.[811,1456,1816,2025]" pageId="10" pageNumber="741">
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<smallCapsWord id="8D22A0087D5F9C152D99F8AFFBAAF8D5" baselines="1834" box="[955,1048,1821,1838]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="10" pageNumber="741">MATERIAL</smallCapsWord>
. —
<taxonomicName id="4C7B4D577D5F9C152A63F8AAFA1DF8CB" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1089,1455,1816,1840]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="species" species="rhomboides">
<emphasis id="B90FEAC67D5F9C152A63F8AAFAB4F8CB" box="[1089,1286,1816,1840]" italics="true" pageId="10" pageNumber="741">Goneplax rhomboides</emphasis>
(
<bibRefCitation id="EFEA4B257D5F9C152B33F8AAFA1AF8CB" author="LINNAEUS C." box="[1297,1448,1816,1840]" pageId="10" pageNumber="741" refId="ref12756" refString="LINNAEUS C. 1758. - Systema Naturae per Regna Tria Naturae Secundum Classes Ordines Genera Species cum Characteribus Differentiis Synonymis Locis Tomus I. Laurentii Salvii, Holmi., [iv] + 824 p. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
<specimenCount id="9D7DFD5D7D5F9C152D18F880FCD2F8B6" box="[826,864,1842,1869]" count="3" pageId="10" pageNumber="741" type="male">3 ♂</specimenCount>
(largest 36.5 ×
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); Kames Bay, Isle of Cumbray,
<collectingCountry id="F36C76447D5F9C152B5EF880FCE8F89E" name="United Kingdom" pageId="10" pageNumber="741">Scotland</collectingCountry>
; coll. S. De Grave;
<date id="FFC510147D5F9C152A37F8FEFBCEF89F" box="[1045,1148,1868,1892]" pageId="10" pageNumber="741" value="2005-06-02">2.VI.2005</date>
; ZRC 2010.0069 •
<specimenCount id="9D7DFD5D7D5F9C152B1DF8FFFAD0F893" box="[1343,1378,1869,1896]" count="1" pageId="10" pageNumber="741" type="male">1♂</specimenCount>
(22.0 ×
<quantity id="4C839B317D5F9C152D0EF8D5FC37F884" box="[812,901,1895,1919]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="10" pageNumber="741" unit="mm" value="35.8">35.8 mm</quantity>
); Grand Bassam,
<collectingRegion id="49BFF8367D5F9C152A01F8D5FBC2F884" box="[1059,1136,1895,1919]" country="Ivory Coast" name="Abidjan" pageId="10" pageNumber="741">Abidjan</collectingRegion>
,
<collectingCountry id="F36C76447D5F9C152A55F8DAFB56F884" box="[1143,1252,1895,1920]" name="Ivory Coast" pageId="10" pageNumber="741">Ivory Coast</collectingCountry>
, trawl;
<quantity id="4C839B317D5F9C152B07F8D5FAD6F884" box="[1317,1380,1895,1919]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="741" unit="m" value="250.0">250 m</quantity>
; coll. Le Loeuff &amp; Intès;
<date id="FFC510147D5F9C152DF2F833FBFEF862" box="[976,1100,1921,1945]" pageId="10" pageNumber="741" value="1966-12-23">23.XII.1966</date>
(det.
<taxonomicName id="4C7B4D577D5F9C152AA8F833FAF3F862" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[1162,1345,1921,1945]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B90FEAC67D5F9C152AA8F833FAF3F862" box="[1162,1345,1921,1945]" italics="true" pageId="10" pageNumber="741">Goneplax angulata</emphasis>
</taxonomicName>
) MNHN- IU-2024-5374 (= MNHN-B19812).
<taxonomicName id="4C7B4D577D5F9C152A8BF82EFC1FF834" authority="Guinot &amp; Castro, 2007" authorityName="Guinot &amp; Castro" authorityYear="2007" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="741" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B90FEAC67D5F9C152A8BF82EFAF0F84F" box="[1193,1346,1948,1972]" italics="true" pageId="10" pageNumber="741">Goneplax clevai</emphasis>
<bibRefCitation id="EFEA4B257D5F9C152B68F82EFC1FF834" author="GUINOT D. &amp; CASTRO P." pageId="10" pageNumber="741" pagination="17 - 31" refId="ref12422" refString="GUINOT D. &amp; CASTRO P. 2007. - A new species of Goneplax Leach 1814 (Crustacea Decapoda Brachyura Goneplacidae) from the south Atlantic and western limits of the Indo-West Pacific region long confused with G. rhomboides (Linnaeus 1758). Zootaxa 1577: 17 - 31. https: // doi. org / 10.11646 / zootaxa. 1577.1.3" type="journal article" year="2007">Guinot &amp; Castro, 2007</bibRefCitation>
</taxonomicName>
<specimenCount id="9D7DFD5D7D5F9C152D9DF805FC53F829" box="[959,993,1975,2002]" count="1" pageId="10" pageNumber="741" type="male">1♂</specimenCount>
(21.3 ×
<quantity id="4C839B317D5F9C152A13F804FB3EF834" box="[1073,1164,1974,1999]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="10" pageNumber="741" unit="mm" value="35.3">35.3 mm</quantity>
); Port Elizabeth,
<collectingCountry id="F36C76447D5F9C152B12F804FA19F835" box="[1328,1451,1974,1998]" name="South Africa" pageId="10" pageNumber="741">South Africa</collectingCountry>
; coll. S. Fennessy, from local trawlers;
<date id="FFC510147D5F9C152AACF863FB4CF812" box="[1166,1278,2001,2025]" pageId="10" pageNumber="741" value="2003-12-01">1.XII.2003</date>
; ZRC 2004.0700.
</paragraph>
</subSubSection>
</treatment>
</document>