293 lines
35 KiB
XML
293 lines
35 KiB
XML
<document id="6FF796731474C1D4355A16256C1ACE69" ID-DOI="10.11646/zootaxa.3670.2.9" ID-GBIF-Dataset="492d3ca4-105c-4c92-8a7d-6cb180d00b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="223576" ID-ZooBank="33F668E4-95BC-4203-B892-80365B4F3B89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633857078" checkinUser="plazi" docAuthor="Filippini, Valentina, Micó, Estefania & Galante, Eduardo" docDate="2013" docId="03D28781FFFEFFD95AA660E7FB1EFD33" docLanguage="en" docName="zt03670p273.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala flavacoma Filippini, Mico" docType="treatment" docVersion="8" lastPageNumber="266" masterDocId="FFEBFFF9FFF7FFD25A31622DFFA8FFB7" masterDocTitle="Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" masterLastPageNumber="273" masterPageNumber="255" pageNumber="264" updateTime="1698334918277" updateUser="plazi">
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<mods:titleInfo id="3BDA428917DAFAC95643FD481C483470">
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<mods:title id="AD3FE391F82F2F361F57B2B678DCC60E">Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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<mods:name id="A41EA4EF18A22D8C87E4564BAD32FD69" type="personal">
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<mods:namePart id="4F7E3BAC5600575814214398ED27B51E">Filippini, Valentina</mods:namePart>
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<mods:roleTerm id="9BC08C34969899EB269BE9A37ED7183F">Author</mods:roleTerm>
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<mods:namePart id="D039CC64421DF3D028D1BBFF6F17C02F">Micó, Estefania</mods:namePart>
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</mods:name>
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<mods:name id="17126D2E210C30FB2638785252A06F9E" type="personal">
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</mods:role>
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<mods:namePart id="9B62D97A38CB4BF322F2FFADA4F771BF">Galante, Eduardo</mods:namePart>
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</mods:name>
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<mods:title id="F0655C6EB2F6095A6111795969DFE2C8">Zootaxa</mods:title>
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<mods:part id="1FF66FB4D38956ED852CD8A12D067547">
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<mods:date id="688E666C97FC167AE6A9C63AC99DCD70">2013</mods:date>
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<mods:number id="228446CA0F1B9F8386B0F0271B3CBA84">3670</mods:number>
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<mods:identifier id="7939E75EDBEF012B14A1CE5CF7008959" type="DOI">10.11646/zootaxa.3670.2.9</mods:identifier>
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<mods:identifier id="370252878234F3C27D779D53197B60FC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0017719B25851C32EEBDFBDDC730619C" type="Zenodo-Dep">223576</mods:identifier>
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<treatment id="03D28781FFFEFFD95AA660E7FB1EFD33" ID-DOI="http://doi.org/10.5281/zenodo.6159035" ID-GBIF-Taxon="119570127" ID-Zenodo-Dep="6159035" LSID="urn:lsid:plazi:treatment:03D28781FFFEFFD95AA660E7FB1EFD33" httpUri="http://treatment.plazi.org/id/03D28781FFFEFFD95AA660E7FB1EFD33" lastPageId="11" lastPageNumber="266" pageId="9" pageNumber="264">
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<subSubSection id="C361651CFFFEFFDB5AA660E7FF70FCB2" pageId="9" pageNumber="264" type="nomenclature">
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<paragraph id="8BC43697FFFEFFDB5AA660E7FCD3FD53" blockId="9.[151,891,714,773]" box="[151,891,714,740]" pageId="9" pageNumber="264">
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<heading id="D08C81FBFFFEFFDB5AA660E7FCD3FD53" bold="true" box="[151,891,714,740]" fontSize="11" level="1" pageId="9" pageNumber="264" reason="1">
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<emphasis id="B90FEA85FFFEFFDB5AA660E7FCD3FD53" bold="true" box="[151,891,714,740]" pageId="9" pageNumber="264">
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<taxonomicName id="4C7B4D14FFFEFFDB5AA660E7FDE5FD53" authority="Filippini, Mico" authorityName="Filippini, Mico" box="[151,589,714,740]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="flavacoma">
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<emphasis id="B90FEA85FFFEFFDB5AA660E7FE22FD53" bold="true" box="[151,394,714,740]" italics="true" pageId="9" pageNumber="264">Anomala flavacoma</emphasis>
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<collectingCountry id="F36C7607FFFEFFDB5BA360E7FE57FD53" box="[402,511,714,740]" name="Philippines" pageId="9" pageNumber="264">Filippini</collectingCountry>
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, Micó
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</taxonomicName>
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, & Galante, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC43697FFFEFFDB5AA660C0FF70FCB2" blockId="9.[151,891,714,773]" box="[151,216,749,773]" pageId="9" pageNumber="264">
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<figureCitation id="13402A12FFFEFFDB5AA660C0FF70FCB2" box="[151,216,749,773]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1929,1953]" captionTargetBox="[193,1406,206,1890]" captionTargetId="figure@4.[184,1420,194,1919]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Habitus images. 2: Anomala eucoma (paralectotype, Las Mercedes, Guatemala). 3: A. flavaeucoma (female paratype, Estación Hitoy Cerere, Costa Rica). 4. A. megaparamera (holotype). 5: A. pseudoeucoma (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/223578/files/figure.png" pageId="9" pageNumber="264">Fig. 3</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C361651CFFFEFFDB5AA66103FA99FA31" pageId="9" pageNumber="264" type="materials_examined">
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<paragraph id="8BC43697FFFEFFDB5AA66103FC3FFCDD" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">
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<emphasis id="B90FEA85FFFEFFDB5AA66103FE2EFCF0" bold="true" box="[151,390,814,839]" pageId="9" pageNumber="264">Material examined:</emphasis>
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<typeStatus id="54C08835FFFEFFDB5BA06103FE56FCF1" box="[401,510,814,838]" pageId="9" pageNumber="264" type="holotype">Holotype</typeStatus>
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: ɞ "Limon,
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<collectingCountry id="F36C7607FFFEFFDB58A26103FCB8FCF1" box="[659,784,814,838]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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Reserva Hitoy–Cerere Rio Cerere 100 msnm
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<date id="FFC51057FFFEFFDB5F1E6103FF67FCDD" pageId="9" pageNumber="264" value="1987-03-24">24 marzo 1987</date>
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Angel Solis / 56 /
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<collectingCountry id="F36C7607FFFEFFDB5B90617FFDB3FCDD" box="[417,539,850,874]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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INBIOCRI002517822" (
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<collectionCode id="ED6AAE52FFFEFFDB5906617FFC23FCDD" box="[823,907,850,874]" pageId="9" pageNumber="264">INBIO</collectionCode>
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).
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</paragraph>
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<paragraph id="8BC43697FFFEFFDB5AF6615AFA99FA31" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">
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<typeStatus id="54C08835FFFEFFDB5AF6615AFE9CFC38" box="[199,308,887,911]" pageId="9" pageNumber="264" type="paratype">Paratypes</typeStatus>
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(11): 1ɞ "Est. Hitoy Cerere,
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<quantity id="4C839B72FFFEFFDB58BD615BFD67FC39" box="[652,719,886,910]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="100.0">100m</quantity>
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, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon,
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<collectingCountry id="F36C7607FFFEFFDB5F29615BFA30FC39" box="[1304,1432,886,910]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
,
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<date id="FFC51057FFFEFFDB5AA661B6FEA0FC05" box="[151,264,922,946]" pageId="9" pageNumber="264" value="1992-11">Nov 1992</date>
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G. Carballo L- N /
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<collectingCountry id="F36C7607FFFEFFDB5BDD61B7FDC0FC05" box="[492,616,922,946]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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INBIOCRI000926676"; 1ɞ "Est. Hitoy Cerere,
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<quantity id="4C839B72FFFEFFDB5EA761B7FB48FC05" box="[1174,1248,922,946]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="100.0">100 m</quantity>
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, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon,
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<collectingCountry id="F36C7607FFFEFFDB58196193FD7DFC61" box="[552,725,958,982]" name="Costa Rica" pageId="9" pageNumber="264">COSTA RICA</collectingCountry>
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.
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<date id="FFC51057FFFEFFDB58D56193FCFFFC61" box="[740,855,958,982]" pageId="9" pageNumber="264" value="1993-04">Abr 1993</date>
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. G. Carballo. L- N 184200_643300 /
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<collectingCountry id="F36C7607FFFEFFDB5F2D6193FA34FC61" box="[1308,1436,958,982]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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INBIOCRI000975444"; 1Ƥ "Est. Hitoy Cerere,
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||
<quantity id="4C839B72FFFEFFDB58F561CFFCAFFC4D" box="[708,775,994,1018]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="100.0">100m</quantity>
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||
, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon,
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||
<collectingCountry id="F36C7607FFFEFFDB5F6D61CFFF65FBA9" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
,
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||
<date id="FFC51057FFFEFFDB5AEB662AFEE3FBA9" box="[218,331,1030,1054]" pageId="9" pageNumber="264" value="1992-11">Nov 1992</date>
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||
G. Carballo L- N /
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||
<collectingCountry id="F36C7607FFFEFFDB581E662BFD03FBA9" box="[559,683,1030,1054]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
INBIOCRI000926686"; 1Ƥ "Est. Hitoy Cerere,
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||
<quantity id="4C839B72FFFEFFDB5EE9662BFAB3FBA9" box="[1240,1307,1030,1054]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="100.0">100m</quantity>
|
||
, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon,
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||
<collectingCountry id="F36C7607FFFEFFDB58726607FD17FBF5" box="[579,703,1066,1090]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
,
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||
<date id="FFC51057FFFEFFDB58FB6606FC91FBF5" box="[714,825,1066,1090]" pageId="9" pageNumber="264" value="1992-11">Nov 1992</date>
|
||
G. Carballo L- N /
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||
<collectingCountry id="F36C7607FFFEFFDB5E3F6607FB2FFBF5" box="[1038,1159,1066,1090]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
INBIOCRI000926680"; 1ɞ "Cuatro esquinas, P.N. Tortuguero, Prov. Limon
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<collectingCountry id="F36C7607FFFEFFDB58DF6663FC3FFBD1" box="[750,919,1102,1126]" name="Costa Rica" pageId="9" pageNumber="264">COSTA RICA</collectingCountry>
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||
. 0m.
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||
<date id="FFC51057FFFEFFDB59E66663FB6AFBD1" box="[983,1218,1102,1127]" pageId="9" pageNumber="264" value="1989-08-26" valueMax="1989-09-06" valueMin="1989-08-26">26 Aug– 6 Set 1989</date>
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. J. Solano 280000, 590500 /
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<collectingCountry id="F36C7607FFFEFFDB5B35665FFE28FB3D" box="[260,384,1138,1162]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
INBIOCRI000020620"; 1ɞ "Albergue Heliconias, Bijagua (
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||
<collectingCountry id="F36C7607FFFEFFDB5E06665FFB12FB3D" box="[1079,1210,1138,1162]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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).
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<quantity id="4C839B72FFFEFFDB5EFF665FFAA6FB3D" box="[1230,1294,1138,1162]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="264" unit="m" value="750.0">750m</quantity>
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<date id="FFC51057FFFEFFDB5F26665FFA30FB3D" box="[1303,1432,1138,1162]" pageId="9" pageNumber="264" value="2006-01-30">30–1–2006</date>
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. Luz casa (18:45–22:00). Leg.: Micó, García, Galante. /
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<collectionCode id="ED6AAE52FFFEFFDB592D66BBFCC3FB19" box="[796,875,1174,1198]" pageId="9" pageNumber="264">CEUA</collectionCode>
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00003306"; 1ɞ "Volcán Tenorio. Alajuela.
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||
<collectingCountry id="F36C7607FFFEFFDB5F6D66BBFF66FB65" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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.
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||
<quantity id="4C839B72FFFEFFDB5AEC6697FE85FB65" box="[221,301,1210,1234]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="9" pageNumber="264" unit="m" value="1160.0">1160m</quantity>
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.
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<date id="FFC51057FFFEFFDB5B0C6697FE0CFB65" box="[317,420,1210,1234]" pageId="9" pageNumber="264" value="2008-04-07">7/4/2008</date>
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. Leg. J.A. Azofeifa /
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<collectionCode id="ED6AAE52FFFEFFDB589F6697FD55FB65" box="[686,765,1210,1234]" pageId="9" pageNumber="264">CEUA</collectionCode>
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||
00003320"; 1ɞ "Estación Cabro Muco, Repr. ICE – Z.P. Miravalles, Guanacaste.
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||
<collectingCountry id="F36C7607FFFEFFDB5B8166F3FD98FB41" box="[432,560,1246,1270]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
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||
.
|
||
<quantity id="4C839B72FFFEFFDB580D66F3FD22FB41" box="[572,650,1246,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="1000.0">1000m</quantity>
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<date id="FFC51057FFFEFFDB58A266F3FCBEFB41" box="[659,790,1246,1270]" pageId="9" pageNumber="264" value="2010-06-12">12/06/2010</date>
|
||
. L.
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||
<collectingCountry id="F36C7607FFFEFFDB597266F3FC0EFB41" box="[835,934,1246,1270]" name="Philippines" pageId="9" pageNumber="264">Filippini</collectingCountry>
|
||
, Moraga. /
|
||
<collectionCode id="ED6AAE52FFFEFFDB5E1D66F3FBD3FB41" box="[1068,1147,1246,1270]" pageId="9" pageNumber="264">CEUA</collectionCode>
|
||
00105076"; 1Ƥ "Estación Cabro Muco, Repr. ICE – Z.P. Miravalles, Guanacaste.
|
||
<collectingCountry id="F36C7607FFFEFFDB5911672FFC09FAAD" box="[800,929,1282,1306]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
|
||
.
|
||
<quantity id="4C839B72FFFEFFDB599E672FFC54FAAD" box="[943,1020,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="264" unit="m" value="1000.0">1000m</quantity>
|
||
<date id="FFC51057FFFEFFDB5E36672FFB22FAAD" box="[1031,1162,1282,1306]" pageId="9" pageNumber="264" value="2010-06-12">12/06/2010</date>
|
||
. L.
|
||
<collectingCountry id="F36C7607FFFEFFDB5E8B672FFAB5FAAD" box="[1210,1309,1282,1306]" name="Philippines" pageId="9" pageNumber="264">Filippini</collectingCountry>
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||
, Moraga. /
|
||
<collectionCode id="ED6AAE52FFFEFFDB5AA6670BFF4EFA89" box="[151,230,1318,1342]" pageId="9" pageNumber="264">CEUA</collectionCode>
|
||
00105077"; 1Ƥ "Dos de Tilarán (San Ramón), Prov. Guana.
|
||
<collectingCountry id="F36C7607FFFEFFDB59F8670BFBDFFA89" box="[969,1143,1318,1342]" name="Costa Rica" pageId="9" pageNumber="264">COSTA RICA</collectingCountry>
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||
.
|
||
<quantity id="4C839B72FFFEFFDB5EB9670BFB70FA89" box="[1160,1240,1318,1342]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="264" unit="m" value="1100.0">1100m</quantity>
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||
.
|
||
<date id="FFC51057FFFEFFDB5EDB670AFADAFA89" box="[1258,1394,1318,1343]" pageId="9" pageNumber="264" value="1995-05">MAY 1995</date>
|
||
. G. Rodriguez. L_N_262600_437500 #5891 / INBIOCRI002337626"; 1Ƥ "Dos de Tilarán (San Ramón), Prov. Guana.
|
||
<collectingCountry id="F36C7607FFFEFFDB5AA66743FE96FA31" box="[151,318,1390,1414]" name="Costa Rica" pageId="9" pageNumber="264">COSTA RICA</collectingCountry>
|
||
.
|
||
<quantity id="4C839B72FFFEFFDB5B796743FE30FA31" box="[328,408,1390,1414]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="264" unit="m" value="1100.0">1100m</quantity>
|
||
.
|
||
<date id="FFC51057FFFEFFDB5B926742FD8CFA31" box="[419,548,1390,1415]" pageId="9" pageNumber="264" value="1995-05">MAY 1995</date>
|
||
. G. Rodriguez. L_N_262600_437500 #5891/ INBIOCRI002337602".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361651CFFFEFFD85AF667BFFE74FC13" lastPageId="10" lastPageNumber="265" pageId="9" pageNumber="264" type="description">
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<paragraph id="8BC43697FFFEFFDB5AF667BFFC96F9A1" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">
|
||
<emphasis id="B90FEA85FFFEFFDB5AF667BFFEF1FA1C" bold="true" box="[199,345,1426,1451]" pageId="9" pageNumber="264">Description.</emphasis>
|
||
Male. Body shape oval. Length
|
||
<quantity id="4C839B72FFFEFFDB58EA67BFFCFBFA1D" box="[731,851,1426,1450]" metricMagnitude="-2" metricUnit="m" metricValue="1.121" pageId="9" pageNumber="264" unit="mm" value="11.21">11.21 mm</quantity>
|
||
. Width
|
||
<quantity id="4C839B72FFFEFFDB598067BFFBB3FA1D" box="[945,1051,1426,1450]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="264" unit="mm" value="6.3">6.30 mm</quantity>
|
||
. Head, pronotum, scutellum, and pygidium dark brown; legs and underside from reddish brown to dark orange; elytra light brown with suture, humerus, and basal half of lateral margins dark brown. The dark stripe on suture broadens mid elytra, forming a small expansion. Surface all covered with fine, blonde setae.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFEFFDB5AF6640FFBEEF935" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">Clypeus trapezoidal, densely punctate-reticulate; ratio width/length 1.88. Anterior angles rounded. Frons densely punctate, flat; vertex with apparent boundary. Ocular canthum long, thin, and rounded at apex. Interocular ratio (interocular width/width of eye): 3.20. Male antenna: ratio funiculus/club 0.76.</paragraph>
|
||
<paragraph id="8BC43697FFFEFFDB5AF664A2FD8BF959" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">Pronotum trapezoidal, width 1.71 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and sharp. Basal margin thin and complete, sinuate. Entire surface with deep punctures.</paragraph>
|
||
<paragraph id="8BC43697FFFEFFDB5AF664D7FE0CF881" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">Scutellum subpentagonal to subtriangular in shape, with rounded sides and acute apex. Ratio width/length 1.38. Sparsely punctate.</paragraph>
|
||
<paragraph id="8BC43697FFFEFFDB5AF6656FFB97F815" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">Elytra with striae defined by dark punctures. Subsutural interstice with 2–3 rows of sparse punctures, 2nd–4th interstices with one irregular secondary striae. Surface covered with shallow setigerous punctures. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</paragraph>
|
||
<paragraph id="8BC43697FFFEFFDB5AF66583FE42F85D" blockId="9.[151,1437,814,2026]" pageId="9" pageNumber="264">Pygidium finely granulate, with longer setae at hind margins. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.69.</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF662BAFC58FF63" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">
|
||
Metasternal disc slightly sulcated, densely punctate. Space between the mesocoxae narrow, slightly convex; width
|
||
<quantity id="4C839B72FFFDFFD85AEC6291FEECFF63" box="[221,324,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="265" unit="mm" value="0.22">0.22 mm</quantity>
|
||
. Mesometasternal suture well defined, at base of mesocoxae.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF662F2FB27FEAB" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">Abdominal sternites completely covered with dense setae, arranged in about 7–10 irregular rows, with strigate surface. Male last sternite strigate-imbricate, with apical margin well defined and rounded.</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF6630AFC9FFE78" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">
|
||
Protibia with 2 teeth (
|
||
<figureCitation id="13402A12FFFDFFD85B8E630AFE55FE88" box="[447,509,295,319]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,269,1156,1180]" captionTargetBox="[174,1395,662,1128]" captionTargetId="figure@5.[151,1436,658,1144]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Shape of protibia in 7: Anomala eucoma (paralectotype, Las Mercedes, Guatemala); 8: A. flavacoma (3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9: A. megaparamera (3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10: A. pseudoeucoma (3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="10" pageNumber="265">Fig 8</figureCitation>
|
||
); apical tooth long and curved; second tooth at the same level of internal apex of protibia, triangular in shape, obtuse. Metatibia stout, slightly narrower subapically. Ratio length/width 2.95. First carina well developed. Surface with elongate punctures above second carina and rugose below. Protarsal claws: external claw strongly curved; ratio length/height: 1.37; internal claw bifurcate, with upper branch slightly shorter and half the width of the lower one. Inferior margin sinuate.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF663F1FC5EFDE8" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">
|
||
Aedeagus (
|
||
<figureCitation id="13402A12FFFDFFD85B7463F1FE61FE43" box="[325,457,476,500]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="10" pageNumber="265">Figs 17–19</figureCitation>
|
||
): parameres pointed and narrow. Endophallus (
|
||
<figureCitation id="13402A12FFFDFFD859EC63F1FB87FE43" box="[989,1071,476,500]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="10" pageNumber="265">Fig. 30</figureCitation>
|
||
) wide, with 2 short sacculi. The wider, ventral one has a strip of thick spines, with defined margins that extend frontally from side to side in a horseshoe shape. The dorsal sacculus is covered with fine setation, slightly thicker than the microsetation that commonly covers the entire surface of endophallus. Median lobe sclerotized.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF66041FC4CFD7B" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">
|
||
Female. Similar to male. Scutellum with blunt apex. Internal protarsal claw narrower than in male, inferior margin straight. Protibia (
|
||
<figureCitation id="13402A12FFFDFFD85B8460A2FE5FFD10" box="[437,503,655,679]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,269,1156,1180]" captionTargetBox="[174,1395,662,1128]" captionTargetId="figure@5.[151,1436,658,1144]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Shape of protibia in 7: Anomala eucoma (paralectotype, Las Mercedes, Guatemala); 8: A. flavacoma (3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9: A. megaparamera (3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10: A. pseudoeucoma (3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="10" pageNumber="265">Fig. 8</figureCitation>
|
||
) with apical tooth wider and longer than in male; second tooth above internal apex. Metatibia wider than in males. Last abdominal sternite with curved margin.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFDFFD85AF660FAFE74FC13" blockId="10.[151,1437,151,932]" pageId="10" pageNumber="265">
|
||
<emphasis id="B90FEA85FFFDFFD85AF660FAFE93FD47" bold="true" box="[199,315,727,752]" pageId="10" pageNumber="265">Variation.</emphasis>
|
||
Dark pigmentation of elytra margin can extend to the tip of elytra; the expansions of sutural dark stripe mid length of elytra can be inconspicuous or form a complete band across the elytra (as in specimens from P.N. Tortuguero, Est. Cabro Muco, Heliconias). Body length
|
||
<quantity id="4C839B72FFFDFFD859716132FBA8FC8F" box="[832,1024,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="1.1035" metricValueMax="1.121" metricValueMin="1.0859999999999999" pageId="10" pageNumber="265" unit="mm" value="11.035" valueMax="11.21" valueMin="10.86">10.86–11.21 mm</quantity>
|
||
, body width
|
||
<quantity id="4C839B72FFFDFFD85EA16132FA9EFC8F" box="[1168,1334,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="6.17" metricValueMax="6.3" metricValueMin="6.04" pageId="10" pageNumber="265" unit="mm" value="6.17" valueMax="6.3" valueMin="6.04">6.04–6.30 mm</quantity>
|
||
. Clypeus w/l: 1.88–2.04. Interocular ratio (interocular width/width of eye): 3.15–3.32. Pronotum w/l: 1.56–1.71. Scutellum w/l: 1.33–1.49. Pygidium w/l: 1.66–1.77. Width between mesocoxae:
|
||
<quantity id="4C839B72FFFDFFD859F3614AFBC4FC37" box="[962,1132,871,896]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="10" pageNumber="265" unit="mm" value="0.205" valueMax="0.22" valueMin="0.19">0.19–0.22 mm</quantity>
|
||
. Metatibia w/l: 2.68–2.95. External claw l/h: 1.37–1.58.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF04661FFFFDFFD85AA66582FD06F871" box="[151,686,1967,1991]" httpUri="https://zenodo.org/record/223584/files/figure.png" pageId="10" pageNumber="265" targetBox="[173,1418,996,1917]" targetPageId="10">
|
||
<paragraph id="8BC43697FFFDFFD85AA66582FD06F871" blockId="10.[151,686,1967,1991]" box="[151,686,1967,1991]" pageId="10" pageNumber="265">
|
||
<emphasis id="B90FEA85FFFDFFD85AA66582FE88F870" bold="true" box="[151,288,1967,1991]" pageId="10" pageNumber="265">FIGURE 34</emphasis>
|
||
. Distribution of
|
||
<taxonomicName id="4C7B4D14FFFDFFD85BE0659DFD0FF870" box="[465,679,1968,1991]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="species" species="flavacoma">
|
||
<emphasis id="B90FEA85FFFDFFD85BE0659DFD0FF870" box="[465,679,1968,1991]" italics="true" pageId="10" pageNumber="265">Anomala flavacoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C361651CFFFCFFD95AF662BAFBFAFD8B" pageId="11" pageNumber="266" type="diagnosis">
|
||
<paragraph id="8BC43697FFFCFFD95AF662BAFD19FEAB" blockId="11.[151,1436,151,644]" pageId="11" pageNumber="266">
|
||
<emphasis id="B90FEA85FFFCFFD95AF662BAFEE9FF07" bold="true" box="[199,321,151,176]" pageId="11" pageNumber="266">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C7B4D14FFFCFFD95B7F62B5FD9EFF07" box="[334,566,152,176]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="flavacoma">
|
||
<emphasis id="B90FEA85FFFCFFD95B7F62B5FD9EFF07" box="[334,566,152,176]" italics="true" pageId="11" pageNumber="266">Anomala flavacoma</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from similar species using the combination of the following characters: medium size (
|
||
<quantity id="4C839B72FFFCFFD9581A6291FD0FFF63" box="[555,679,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="11" pageNumber="266" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
) elytra light brown, with a dark central spot to a complete medium transversal band; metatibia with oblong punctures above second carina, abdominal sternites with dense setae, endophallus with two short and swollen sacculi.
|
||
</paragraph>
|
||
<paragraph id="8BC43697FFFCFFD95AF6630AFBFAFD8B" blockId="11.[151,1436,151,644]" pageId="11" pageNumber="266">
|
||
Slightly smaller and more slender than
|
||
<taxonomicName id="4C7B4D14FFFCFFD958B56305FD53FEF7" box="[644,763,296,320]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="eucoma">
|
||
<emphasis id="B90FEA85FFFCFFD958B56305FD53FEF7" box="[644,763,296,320]" italics="true" pageId="11" pageNumber="266">A. eucoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D14FFFCFFD959056305FC52FEF7" box="[820,1018,296,320]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="pseudoeucoma">
|
||
<emphasis id="B90FEA85FFFCFFD959056305FC52FEF7" box="[820,1018,296,320]" italics="true" pageId="11" pageNumber="266">A. pseudoeucoma</emphasis>
|
||
</taxonomicName>
|
||
, it can be distinguished by the much lighter colour of elytra, pronotum, and underside; slender apex of parameres and endophallus that is shorter and wider. The specimens that develop a horizontal dark band on elytra may look similar to the lighter specimens of
|
||
<taxonomicName id="4C7B4D14FFFCFFD95FB5635DFF46FE1B" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="eucoma">
|
||
<emphasis id="B90FEA85FFFCFFD95FB5635DFF46FE1B" italics="true" pageId="11" pageNumber="266">A. eucoma</emphasis>
|
||
</taxonomicName>
|
||
, but in
|
||
<taxonomicName id="4C7B4D14FFFCFFD95B7F63B9FE4CFE1B" box="[334,484,404,428]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="flavacoma">
|
||
<emphasis id="B90FEA85FFFCFFD95B7F63B9FE4CFE1B" box="[334,484,404,428]" italics="true" pageId="11" pageNumber="266">A. flavacoma</emphasis>
|
||
</taxonomicName>
|
||
the band is much darker than in
|
||
<taxonomicName id="4C7B4D14FFFCFFD9594163B9FC42FE1B" box="[880,1002,404,428]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="eucoma">
|
||
<emphasis id="B90FEA85FFFCFFD9594163B9FC42FE1B" box="[880,1002,404,428]" italics="true" pageId="11" pageNumber="266">A. eucoma</emphasis>
|
||
</taxonomicName>
|
||
, and there is just one. Moreover, the puncture on the elytra is shallow and not coalescent in
|
||
<taxonomicName id="4C7B4D14FFFCFFD959356395FC30FE67" box="[772,920,440,464]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="flavacoma">
|
||
<emphasis id="B90FEA85FFFCFFD959356395FC30FE67" box="[772,920,440,464]" italics="true" pageId="11" pageNumber="266">A. flavacoma</emphasis>
|
||
</taxonomicName>
|
||
, with clear differentiation between setigerous punctures and striae punctures, the latter being larger and dark coloured; whereas in
|
||
<taxonomicName id="4C7B4D14FFFCFFD95E4D63F1FB5EFE43" box="[1148,1270,476,500]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="eucoma">
|
||
<emphasis id="B90FEA85FFFCFFD95E4D63F1FB5EFE43" box="[1148,1270,476,500]" italics="true" pageId="11" pageNumber="266">A. eucoma</emphasis>
|
||
</taxonomicName>
|
||
both
|
||
<typeStatus id="54C08835FFFCFFD95F0F63F0FAD1FE42" box="[1342,1401,477,501]" pageId="11" pageNumber="266">types</typeStatus>
|
||
of punctures are deep, and coalescent.
|
||
<taxonomicName id="4C7B4D14FFFCFFD9581C602DFD50FDAF" box="[557,760,512,536]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="species" species="megaparamera">
|
||
<emphasis id="B90FEA85FFFCFFD9581C602DFD50FDAF" box="[557,760,512,536]" italics="true" pageId="11" pageNumber="266">A. megaparamera</emphasis>
|
||
</taxonomicName>
|
||
is much smaller and darker in colour, with a proportionally wider pygidium, wider parameres and proportionally longer and thinner endophallus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361651CFFFCFFD95AF6606AFCFEFDE8" box="[199,854,583,608]" pageId="11" pageNumber="266" type="distribution">
|
||
<paragraph id="8BC43697FFFCFFD95AF6606AFCFEFDE8" blockId="11.[151,1436,151,644]" box="[199,854,583,608]" pageId="11" pageNumber="266">
|
||
<emphasis id="B90FEA85FFFCFFD95AF6606AFEC8FDD7" bold="true" box="[199,352,583,608]" pageId="11" pageNumber="266">Distribution.</emphasis>
|
||
Caribbean lowlands of
|
||
<collectingCountry id="F36C7607FFFCFFD9585A606AFD4DFDE8" box="[619,741,583,607]" name="Costa Rica" pageId="11" pageNumber="266">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="13402A12FFFCFFD958C4606AFCE1FDE8" box="[757,841,583,607]" captionStart="FIGURE 34" captionStartId="10.[151,255,1967,1991]" captionTargetBox="[173,1418,996,1917]" captionTargetId="figure@10.[151,1436,966,1946]" captionTargetPageId="10" captionText="FIGURE 34. Distribution of Anomala flavacoma." httpUri="https://zenodo.org/record/223584/files/figure.png" pageId="11" pageNumber="266">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361651CFFFCFFD95AF66041FB1EFD33" box="[199,1206,620,644]" pageId="11" pageNumber="266" type="etymology">
|
||
<paragraph id="8BC43697FFFCFFD95AF66041FB1EFD33" blockId="11.[151,1436,151,644]" box="[199,1206,620,644]" pageId="11" pageNumber="266">
|
||
<emphasis id="B90FEA85FFFCFFD95AF66041FEE6FD33" bold="true" box="[199,334,620,644]" pageId="11" pageNumber="266">Etymology.</emphasis>
|
||
From Latin "flava", blonde, and "coma", hair, for its pale colour and setation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |