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<document id="284C627B6882BDAF875099DA2F89F5DB" ID-DOI="10.11646/zootaxa.5357.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10012545" ID-ZooBank="5DDC347A-3DE3-45F0-8BB5-7EAD1DE87E7C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1697528556513" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Wiesner, Jürgen" docDate="2023" docId="03BB87D4FFBB2E2EFF0391EEFA3A0A6B" docLanguage="en" docName="zootaxa.5357.1.3.pdf" docOrigin="Zootaxa 5357 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5357.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symplecthyma (Symplecthyma) Rivalier 1964" docType="key" docVersion="3" lastPageNumber="73" masterDocId="FF82FFACFFB92E2CFF94905BFFA60F62" masterDocTitle="Philippine species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae). Part 2. Subgenus Symplecthyma Rivalier, 1964" masterLastPageNumber="99" masterPageNumber="71" pageNumber="73" updateTime="1697651972978" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="F88EBF1B6828831BCA64FB9F93E2D809">Philippine species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae). Part 2. Subgenus Symplecthyma Rivalier, 1964</mods:title>
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<mods:namePart id="B3A1421703EDC5A255C774E9C63E7C58">Wiesner, Jürgen</mods:namePart>
<mods:affiliation id="4FD1AEDF66236472FDF92C3D7B4CACA9">Dresdener Ring 11, D- 38444 Wolfsburg, Germany</mods:affiliation>
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<mods:date id="353F1F629DEC943892CB74C4F42C98B4">2023</mods:date>
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<treatment id="03BB87D4FFBB2E2EFF0391EEFA3A0A6B" LSID="urn:lsid:plazi:treatment:03BB87D4FFBB2E2EFF0391EEFA3A0A6B" httpUri="http://treatment.plazi.org/id/03BB87D4FFBB2E2EFF0391EEFA3A0A6B" lastPageNumber="73" pageId="2" pageNumber="73">
<subSubSection id="C3086549FFBB2E2EFF0391EEFC0C0EAD" box="[151,938,436,463]" pageId="2" pageNumber="73" type="nomenclature">
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<heading id="D0E581AEFFBB2E2EFF0391EEFC0C0EAD" bold="true" box="[151,938,436,463]" fontSize="11" level="1" pageId="2" pageNumber="73" reason="1">
<emphasis id="B966EAD0FFBB2E2EFF0391EEFC0C0EAD" bold="true" box="[151,938,436,463]" pageId="2" pageNumber="73">
Key to the species of the subgenus
<taxonomicName id="4C124D41FFBB2E2EFDD291EFFC0C0EAD" authority="Rivalier, 1964" authorityName="Rivalier" authorityYear="1964" box="[582,938,436,463]" class="Insecta" family="Cicindelidae" genus="Symplecthyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD0FFBB2E2EFDD291EFFD500EAD" bold="true" box="[582,758,436,463]" italics="true" pageId="2" pageNumber="73">Symplecthyma</emphasis>
<bibRefCitation id="EF834B33FFBB2E2EFD6A91EEFC0C0EAD" author="Rivalier, E." box="[766,938,437,463]" pageId="2" pageNumber="73" pagination="127 - 164" refId="ref16974" refString="Rivalier, E. (1964) Le genre Prothyma Hope. Revision et description de quatre especes nouvelles. Revue francaise d'Entomologie, 31 (3), 127 - 164." type="journal article" year="1964">Rivalier, 1964</bibRefCitation>
</taxonomicName>
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<paragraph id="8BAD36C2FFBB2E2EFF0391A1FA3A0D6D" blockId="2.[151,1437,506,1290]" box="[151,1436,506,528]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF0391A1FA3A0D6D" box="[151,1436,506,528]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF0391A1FF050D6D" bold="true" box="[151,163,506,527]" pageId="2" pageNumber="73">1</emphasis>
Abdominal sternites light brown (
<figureCitation id="13292A47FFBB2E2EFDB291A1FD200D72" box="[550,646,506,528]" captionStart="FIGURE 1" captionStartId="5.[152,255,1553,1577]" captionTargetBox="[181,1415,198,1509]" captionTargetId="figure-17@5.[151,1436,181,1529]" captionTargetPageId="5" captionText="FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10012549" httpUri="https://zenodo.org/record/10012549/files/figure.png" pageId="2" pageNumber="73">Figs 1a, b</figureCitation>
)................................................................
<emphasis id="B966EAD0FFBB2E2EFA0491A1FA3A0D6D" bold="true" box="[1424,1436,506,527]" pageId="2" pageNumber="73">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF03924CFA3A0D4E" blockId="2.[151,1437,506,1290]" box="[151,1436,535,557]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF03924CFA3A0D4E" box="[151,1436,535,557]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF03924CFF390D4E" bold="true" box="[151,159,535,556]" pageId="2" pageNumber="73">-</emphasis>
Abdominal sternites entirely metallic (
<figureCitation id="13292A47FFBB2E2EFDC0924CFD310D4E" box="[596,663,535,557]" captionStart="FIGURE 1" captionStartId="5.[152,255,1553,1577]" captionTargetBox="[181,1415,198,1509]" captionTargetId="figure-17@5.[151,1436,181,1529]" captionTargetPageId="5" captionText="FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10012549" httpUri="https://zenodo.org/record/10012549/files/figure.png" pageId="2" pageNumber="73">Fig. 1c</figureCitation>
)..............................................................
<emphasis id="B966EAD0FFBB2E2EFA04924CFA3A0D4E" bold="true" box="[1424,1436,535,556]" pageId="2" pageNumber="73">4</emphasis>
</keyLead>
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<keyStep id="30E62E67FFBB2E2EFF03926FFA3A0DE6" pageId="2" pageNumber="73">
<paragraph id="8BAD36C2FFBB2E2EFF03926FFA3A0D28" blockId="2.[151,1437,506,1290]" box="[151,1436,564,586]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF03926FFA3A0D28" box="[151,1436,564,586]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF03926FFF050D2B" bold="true" box="[151,163,564,585]" pageId="2" pageNumber="73">2</emphasis>
Meso-, metasternum, metepisternum and abdomen light brown (
<figureCitation id="13292A47FFBB2E2EFCDE926FFC2E0D28" box="[842,904,564,586]" captionStart="FIGURE 1" captionStartId="5.[152,255,1553,1577]" captionTargetBox="[181,1415,198,1509]" captionTargetId="figure-17@5.[151,1436,181,1529]" captionTargetPageId="5" captionText="FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10012549" httpUri="https://zenodo.org/record/10012549/files/figure.png" pageId="2" pageNumber="73">Fig 1a</figureCitation>
).......................................... 3
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF039209FA3A0DE6" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF039209FA3A0DE6" pageId="2" pageNumber="73">
- Mesosternum, all episterna and sides of metasternum metallic, central part of metasternum and abdomen light brown (
<figureCitation id="13292A47FFBB2E2EFAE19209FF5C0DE7" captionStart="FIGURE 1" captionStartId="5.[152,255,1553,1577]" captionTargetBox="[181,1415,198,1509]" captionTargetId="figure-17@5.[151,1436,181,1529]" captionTargetPageId="5" captionText="FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10012549" httpUri="https://zenodo.org/record/10012549/files/figure.png" pageId="2" pageNumber="73">Fig. 1b</figureCitation>
)................................................................................
<emphasis id="B966EAD0FFBB2E2EFB2B9234FA3A0DE6" bold="true" box="[1215,1436,623,645]" pageId="2" pageNumber="73">
<taxonomicName id="4C124D41FFBB2E2EFB2B9234FAEF0DE7" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[1215,1353,623,645]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="kapuka" status="sp. nov." subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFB2B9234FB740DE6" bold="true" box="[1215,1234,623,644]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB4B9234FB540DE6" bold="true" box="[1247,1266,623,644]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFA949234FAEF0DE7" bold="true" box="[1280,1353,623,645]" italics="true" pageId="2" pageNumber="73">kapuka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFBB2E2EFAC49234FA3A0DE6" box="[1360,1436,623,644]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<keyStep id="30E62E67FFBB2E2EFF0392D7FA3A0D9B" pageId="2" pageNumber="73">
<paragraph id="8BAD36C2FFBB2E2EFF0392D7FA3A0DDD" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF0392D7FA3A0DDD" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF0392D7FF050DC3" bold="true" box="[151,163,652,673]" pageId="2" pageNumber="73">3</emphasis>
Striae on vertex strong; pronotum gently transversally rugate; elytra coarsely punctate; yellow apical band of elytra reaching suture along margins (
<figureCitation id="13292A47FFBB2E2EFE2C92F1FE580DDD" box="[440,510,682,704]" captionStart="FIGURE 11" captionStartId="21.[152,255,1847,1871]" captionTargetBox="[168,1403,195,1796]" captionTargetId="figure-17@21.[151,1436,181,1823]" captionTargetPageId="21" captionText="FIGURE 11. Prothyma (Symplecthyma) inopinata sp. nov. a) Dorsal habitus holotype female. b) Dorsal habitus paratype female. Scale = 5 mm. c) Labrum. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012583" httpUri="https://zenodo.org/record/10012583/files/figure.png" pageId="2" pageNumber="73">Fig. 11</figureCitation>
). Head as wide as elytra (mean EW/HW = 1.0)...................
<emphasis id="B966EAD0FFBB2E2EFB3E92F1FA3A0DDD" bold="true" box="[1194,1436,681,703]" pageId="2" pageNumber="73">
<taxonomicName id="4C124D41FFBB2E2EFB3E92F1FAEC0DDD" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[1194,1354,681,703]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="inopinata" status="sp. nov." subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFB3E92F1FB1A0DDD" bold="true" box="[1194,1212,682,703]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB5D92F1FB7B0DDD" bold="true" box="[1225,1245,682,703]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFB7F92F1FAEC0DDD" bold="true" box="[1259,1354,682,703]" italics="true" pageId="2" pageNumber="73">inopinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFBB2E2EFAC492F1FA3A0DDD" box="[1360,1436,682,703]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF03929CFA3A0D9B" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF03929CFA3A0D9B" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF03929CFF390DBE" bold="true" box="[151,159,711,732]" pageId="2" pageNumber="73">-</emphasis>
Striae on vertex shallow, pronotum smooth; elytra shiny, sparsely and weakly punctate; yellow apical band of elytra rounded (
<figureCitation id="13292A47FFBB2E2EFF7D92BFFE970D98" box="[233,305,740,762]" captionStart="FIGURE 10" captionStartId="19.[152,255,1944,1968]" captionTargetBox="[268,1307,187,1900]" captionTargetId="figure-17@19.[244,1342,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Prothyma (Symplecthyma) incerata Rivalier, 1964. a) Dorsal habitus holotype, male. Scale = 5 mm. b) Aedeagus. c) Endophallus. Drawings from (Rivalier, 1964)." figureDoi="http://doi.org/10.5281/zenodo.10012581" httpUri="https://zenodo.org/record/10012581/files/figure.png" pageId="2" pageNumber="73">Fig. 10</figureCitation>
). Head wider than elytra (EW/HW = 0.89)....................................
<taxonomicName id="4C124D41FFBB2E2EFBEC92BEFA3A0D9B" authority="Rivalier, 1964" authorityName="Rivalier" authorityYear="1964" box="[1144,1436,740,762]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="incerata" subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFBEC92BEFA3A0D9B" bold="true" box="[1144,1436,740,762]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFBEC92BEFB2C0D98" bold="true" box="[1144,1162,741,762]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB0392BFFB0C0D9B" bold="true" box="[1175,1194,740,761]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFB2C92BFFAAF0D9B" bold="true" box="[1208,1289,740,761]" italics="true" pageId="2" pageNumber="73">incerata</emphasis>
<bibRefCitation id="EF834B33FFBB2E2EFA8492BFFA3A0D9B" author="Rivalier, E." box="[1296,1436,740,762]" pageId="2" pageNumber="73" pagination="127 - 164" refId="ref16974" refString="Rivalier, E. (1964) Le genre Prothyma Hope. Revision et description de quatre especes nouvelles. Revue francaise d'Entomologie, 31 (3), 127 - 164." type="journal article" year="1964">Rivalier, 1964</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyStep id="30E62E67FFBB2E2EFF039359FA3A0C0D" pageId="2" pageNumber="73">
<paragraph id="8BAD36C2FFBB2E2EFF039359FA3A0C57" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF039359FA3A0C57" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF039359FF050C75" bold="true" box="[151,163,770,791]" pageId="2" pageNumber="73">4</emphasis>
Punctures on elytral surface rounded at base and behind humeri; transverse, narrow and often confluent on the disc; becoming squamiform in posterior part............................................................................ 5
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF039366FA3A0C0D" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF039366FA3A0C0D" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF039366FF390C30" bold="true" box="[151,159,829,850]" pageId="2" pageNumber="73">-</emphasis>
Elytra uniformly sculptured laterally and on the disc. Punctures deeply impressed and rounded, slightly transverse in apical part; never squamiform. Body slender, upper side mate, blue or coppery red (
<figureCitation id="13292A47FFBB2E2EFC519301FBBD0C12" box="[965,1051,858,880]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="9.[152,255,1874,1898]" captionStartId-1="10.[152,255,1943,1967]" captionTargetBox-0="[253,1325,187,1839]" captionTargetBox-1="[341,1249,194,1910]" captionTargetId-0="figure-17@9.[251,1337,181,1850]" captionTargetId-1="figure-25@10.[322,1266,181,1920]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 4. Prothyma (Symplecthyma) coerulea Horn, 1920. a) Dorsal habitus Lectotype, male. b) Dorsal habitus Holotype P. bella Dheurle &amp; Colas, 2018, syn. nov. = P. coerulea opacosericea Horn, 1929 comb. nov.; c) Dorsal habitus Lectotype, male P. coerulea opacosericea Horn, 1929 comb. nov. Scale = 5 mm. d–e Labrum P. coerulea s.str. ST. Scale = 1 mm.: d) Male, e) Female. f–h. Aedeagus. Scale = 1 mm. f) P. coerulea Lectotype. g) Lectotype, male P. coerulea opacosericea Horn, 1929 comb. nov.. h) Holotype P. bella Dheurle &amp; Colas, 2018, syn. nov. = P. coerulea opacosericea Horn, 1929 comb. nov." captionText-1="FIGURE 5. Head of Prothyma (Symplecthyma) coerulea Horn, 1920. a) P. coerulea s.str. b) P. bella Dheurle &amp; Colas, 2018, syn. nov., c) P. coerulea opacosericea Horn, 1929. Scale = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10012563" figureDoi-1="http://doi.org/10.5281/zenodo.10012567" httpUri-0="https://zenodo.org/record/10012563/files/figure.png" httpUri-1="https://zenodo.org/record/10012567/files/figure.png" pageId="2" pageNumber="73">Figs 4–5</figureCitation>
)........
<taxonomicName id="4C124D41FFBB2E2EFB199301FA3A0C0D" authority="Horn, 1920" authorityName="W. Horn" authorityYear="1920" box="[1165,1436,857,879]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="coerulea" subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFB199301FA3A0C0D" bold="true" box="[1165,1436,857,879]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFB199301FB060C0D" bold="true" box="[1165,1184,858,879]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB399301FB660C0D" bold="true" box="[1197,1216,858,879]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFB5A9302FA850C0D" bold="true" box="[1230,1315,857,879]" italics="true" pageId="2" pageNumber="73">coerulea</emphasis>
<bibRefCitation id="EF834B33FFBB2E2EFABE9301FA3A0C0D" author="Horn, W." box="[1322,1436,858,879]" pageId="2" pageNumber="73" pagination="1 - 21" refId="ref16698" refString="Horn, W. (1920) Cicindelinen-Studien aus dem schwedischen Reichsmuseum nebst einigen Bemerkungen neuer Arten etc. Arkiv for Zoologi, 13 (11), 1 - 21." type="journal article" year="1920">Horn, 1920</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="30E62E67FFBB2E2EFF03932CFA3A0CC8" pageId="2" pageNumber="73">
<paragraph id="8BAD36C2FFBB2E2EFF03932CFA3A0CEF" blockId="2.[151,1437,506,1290]" box="[151,1436,887,909]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF03932CFA3A0CEF" box="[151,1436,887,909]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF03932CFF050CEE" bold="true" box="[151,163,887,908]" pageId="2" pageNumber="73">5</emphasis>
Aedeagus elongate, narrow in basal third and gradually widened on dorsal side to the middle (
<figureCitation id="13292A47FFBB2E2EFBFC932CFB010CEF" box="[1128,1191,887,909]" captionStart="FIGURE 7" captionStartId="14.[152,255,1222,1246]" captionTargetBox="[163,1420,194,1185]" captionTargetId="figure-25@14.[151,1436,181,1198]" captionTargetPageId="14" captionText="FIGURE 7. Aedeagus of Prothyma (Symplecthyma) eureka sp. nov. a–b) Mt. Apo; c–d) Mt. Kitanglad; e–f) San Miguel; g–i) Agusan del Sur, Esperanza. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012573" httpUri="https://zenodo.org/record/10012573/files/figure.png" pageId="2" pageNumber="73">Figs 7</figureCitation>
,
<figureCitation id="13292A47FFBB2E2EFB25932CFB180CEE" box="[1201,1214,887,908]" captionStart="FIGURE 9" captionStartId="18.[152,255,1351,1375]" captionTargetBox="[161,1424,208,1300]" captionTargetId="figure-25@18.[151,1436,181,1327]" captionTargetPageId="18" captionText="FIGURE 9. Aedeagus of Prothyma (Symplecthyma) heteromallicollis Horn, 1909. a–b) Holotype; c–e) Luzon, Cagayan, St. Ana; f) Mindanao, Luagon; g–h) Mindanao, Nabunturan; i–j) Mindanao, Davao de Oro. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012579" httpUri="https://zenodo.org/record/10012579/files/figure.png" pageId="2" pageNumber="73">9</figureCitation>
)................ 6
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF0393CEFA3A0CC8" blockId="2.[151,1437,506,1290]" box="[151,1436,916,938]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF0393CEFA3A0CC8" box="[151,1436,916,938]" pageId="2" pageNumber="73">
- Aedeagus short, abruptly widened immediately from the base (
<figureCitation id="13292A47FFBB2E2EFCA993CFFCDD0CC8" box="[829,891,916,938]" captionStart="FIGURE 3" captionStartId="7.[152,255,1908,1932]" captionTargetBox="[255,1335,196,1876]" captionTargetId="figure-17@7.[238,1350,181,1884]" captionTargetPageId="7" captionText="FIGURE 3. Aedeagus of Prothyma (Symplecthyma) aeneoparva Horn, 1934, left lateral view. a) Syntype, Biliran; b) Lope DeVega, Samar, lateral and ventral view; c) Syntype, Samar; d) Hinabangan, Samar; e) P. nitida Rivalier 1964 syn. nov., Type, drawing from (Rivalier, 1964); f) Segod, Leyte; g) Park Tayabas, Luzon; h) Surigao, Mindanao; i) Wao, Mindanao; j) Sibagat; k) Cawa-Cawa, Mindanao; l) Gutalac, Mindanao; m) Cabanglasan, Mindanao; n) Esperanza, Mindanao. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012559" httpUri="https://zenodo.org/record/10012559/files/figure.png" pageId="2" pageNumber="73">Figs 3</figureCitation>
,
<figureCitation id="13292A47FFBB2E2EFC1293CEFC380CC8" box="[902,926,917,938]" captionStart="FIGURE 13" captionStartId="25.[152,255,1943,1967]" captionTargetBox="[241,1335,203,1890]" captionTargetId="figure-17@25.[228,1360,181,1919]" captionTargetPageId="25" captionText="FIGURE 13. Prothyma (Symplecthyma) sciakyi sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, male; d) Head, female; e) Aedeagus, left lateral view, Governor Generoso; f) Aedeagus, left lateral view, San Francisco Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012589" httpUri="https://zenodo.org/record/10012589/files/figure.png" pageId="2" pageNumber="73">13</figureCitation>
)........................................ 7
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<keyStep id="30E62E67FFBB2E2EFF0393E9FA3A0B3B" pageId="2" pageNumber="73">
<paragraph id="8BAD36C2FFBB2E2EFF0393E9FA3A0B60" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF0393E9FA3A0B60" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF0393E9FF050CA5" bold="true" box="[151,163,946,967]" pageId="2" pageNumber="73">6</emphasis>
Dorsal side of body shiny, with bright copper and green reflections. Anterior half of pronotum smooth and shiny (
<figureCitation id="13292A47FFBB2E2EFA8893E9FAF10CAA" box="[1308,1367,946,968]" captionStart="FIGURE 6" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[184,1402,196,1906]" captionTargetId="figure-17@13.[169,1418,181,1919]" captionTargetPageId="13" captionText="FIGURE 6. Prothyma (Symplecthyma) eureka sp. nov. a) Dorsal habitus. Holotype male; b) Elytra, paratype, female. Scale = 5 mm. c) Labrum, female; d) Labrum, male. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012571" httpUri="https://zenodo.org/record/10012571/files/figure.png" pageId="2" pageNumber="73">Fig. 6</figureCitation>
). Body relatively robust. Apex of aedeagus narrow, with rounded hook spiral sclerite of endophallus with two full coils at base (
<figureCitation id="13292A47FFBB2E2EFAE19394FF490B60" captionStart="FIGURE 7" captionStartId="14.[152,255,1222,1246]" captionTargetBox="[163,1420,194,1185]" captionTargetId="figure-25@14.[151,1436,181,1198]" captionTargetPageId="14" captionText="FIGURE 7. Aedeagus of Prothyma (Symplecthyma) eureka sp. nov. a–b) Mt. Apo; c–d) Mt. Kitanglad; e–f) San Miguel; g–i) Agusan del Sur, Esperanza. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012573" httpUri="https://zenodo.org/record/10012573/files/figure.png" pageId="2" pageNumber="73">Fig. 7</figureCitation>
).................................................................................
<emphasis id="B966EAD0FFBB2E2EFB5193B6FA3A0B60" bold="true" box="[1221,1436,1004,1026]" pageId="2" pageNumber="73">
<taxonomicName id="4C124D41FFBB2E2EFB5193B6FAEC0B60" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[1221,1354,1004,1026]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="eureka" status="sp. nov." subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFB5193B6FB710B60" bold="true" box="[1221,1239,1005,1026]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB7093B7FB510B63" bold="true" box="[1252,1271,1004,1025]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFA9293B7FAEC0B60" bold="true" box="[1286,1354,1004,1026]" italics="true" pageId="2" pageNumber="73">eureka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFBB2E2EFAC493B6FA3A0B60" box="[1360,1436,1005,1026]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF039451FA3A0B3B" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF039451FA3A0B3B" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF039451FF390B7D" bold="true" box="[151,159,1034,1055]" pageId="2" pageNumber="73">-</emphasis>
Dorsal side of body dull, dark copper. Pronotum evenly, transversely rugate throughout (
<figureCitation id="13292A47FFBB2E2EFBD89451FB2E0B7D" box="[1100,1160,1034,1056]" captionStart="FIGURE 8" captionStartId="17.[152,255,1943,1967]" captionTargetBox="[266,1337,191,1905]" captionTargetId="figure-17@17.[224,1364,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Prothyma (Symplecthyma) heteromallicollis Horn, 1909. a) Dorsal habitus, Holotype, male. b) Elytra, female. Scale = 5 mm. c) Head and labrum, male; d) Head and labrum, female. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012575" httpUri="https://zenodo.org/record/10012575/files/figure.png" pageId="2" pageNumber="73">Fig. 8</figureCitation>
). Apex of aedeagus slightly wider, with down turned tip; spiral sclerite of endophallus wide, with two and half full coils at base (
<figureCitation id="13292A47FFBB2E2EFB33947CFB790B5E" box="[1191,1247,1063,1085]" captionStart="FIGURE 9" captionStartId="18.[152,255,1351,1375]" captionTargetBox="[161,1424,208,1300]" captionTargetId="figure-25@18.[151,1436,181,1327]" captionTargetPageId="18" captionText="FIGURE 9. Aedeagus of Prothyma (Symplecthyma) heteromallicollis Horn, 1909. a–b) Holotype; c–e) Luzon, Cagayan, St. Ana; f) Mindanao, Luagon; g–h) Mindanao, Nabunturan; i–j) Mindanao, Davao de Oro. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012579" httpUri="https://zenodo.org/record/10012579/files/figure.png" pageId="2" pageNumber="73">Fig. 9</figureCitation>
)......................................................................................
<taxonomicName id="4C124D41FFBB2E2EFBA9941EFA3A0B3B" authority="Horn, 1909" authorityName="Horn" authorityYear="1909" box="[1085,1436,1092,1114]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="heteromallicollis" subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFBA9941EFA3A0B3B" bold="true" box="[1085,1436,1092,1114]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFBA9941EFBF60B38" bold="true" box="[1085,1104,1093,1114]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFBC9941FFBD60B3B" bold="true" box="[1117,1136,1092,1113]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFBEA941FFA850B38" bold="true" box="[1150,1315,1092,1114]" italics="true" pageId="2" pageNumber="73">heteromallicollis</emphasis>
<bibRefCitation id="EF834B33FFBB2E2EFABE941EFA3A0B3B" author="Horn, W." box="[1322,1436,1092,1114]" pageId="2" pageNumber="73" pagination="311 - 314" refId="ref16633" refString="Horn, W. (1909) Zwei neue Philippinen- Prothymae. Deutsche Entomologische Zeitschrift, 1902, 311 - 314." type="journal article" year="1909">Horn, 1909</bibRefCitation>
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<keyLead id="30E395F7FFBB2E2EFF039439FA3A0BD0" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF039439FF050B15" bold="true" box="[151,163,1122,1143]" pageId="2" pageNumber="73">7</emphasis>
Pronotum dull, densely transversely rugate. Disc of elytra with coarse transverse punctures, becoming confluent in apical half. Aedeagus short and robust; apex short and wide, with thick hook; spiral sclerite of endophallus wide, with one and half full coils at base (
<figureCitation id="13292A47FFBB2E2EFEF094C7FE0F0BD0" box="[356,425,1180,1202]" captionStart="FIGURE 13" captionStartId="25.[152,255,1943,1967]" captionTargetBox="[241,1335,203,1890]" captionTargetId="figure-17@25.[228,1360,181,1919]" captionTargetPageId="25" captionText="FIGURE 13. Prothyma (Symplecthyma) sciakyi sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, male; d) Head, female; e) Aedeagus, left lateral view, Governor Generoso; f) Aedeagus, left lateral view, San Francisco Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012589" httpUri="https://zenodo.org/record/10012589/files/figure.png" pageId="2" pageNumber="73">Fig. 13</figureCitation>
)..................................................................
<emphasis id="B966EAD0FFBB2E2EFB5F94C6FA3A0BD0" bold="true" box="[1227,1436,1180,1202]" pageId="2" pageNumber="73">
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<emphasis id="B966EAD0FFBB2E2EFB5F94C6FB780BD0" bold="true" box="[1227,1246,1181,1202]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB7F94C7FB580BD3" bold="true" box="[1259,1278,1180,1201]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFA9894C7FAEF0BD0" bold="true" box="[1292,1353,1180,1202]" italics="true" pageId="2" pageNumber="73">sciaky</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFBB2E2EFAC494C6FA3A0BD0" box="[1360,1436,1181,1202]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAD36C2FFBB2E2EFF0394E1FA3A0A6B" blockId="2.[151,1437,506,1290]" pageId="2" pageNumber="73">
<keyLead id="30E395F7FFBB2E2EFF0394E1FA3A0A6B" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFF0394E1FF390BAD" bold="true" box="[151,159,1210,1231]" pageId="2" pageNumber="73">-</emphasis>
Pronotum shiny, moderately transversally rugate. Apex of aedeagus not thickened, with more or less pointed hook-shaped denticle; spiral sclerite of endophallus long and narrow, with three full coils at base (
<figureCitation id="13292A47FFBB2E2EFB99948CFBC50B8E" box="[1037,1123,1239,1261]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[152,255,1909,1933]" captionStartId-1="7.[152,255,1908,1932]" captionTargetBox-0="[245,1340,188,1881]" captionTargetBox-1="[255,1335,196,1876]" captionTargetId-0="figure-25@6.[239,1348,181,1884]" captionTargetId-1="figure-17@7.[238,1350,181,1884]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Prothyma (Symplecthyma) aeneoparva Horn, 1934. a–f: Dorsal habitus. Scale = 5 mm. a) Female, Leyte. b) P. nitida Rivalier 1964 syn. nov., Type, male. c) P. boholensis Dheurle 2012 syn. nov., Holotype, female. d) Male from Cawa- Cawa, Mindanao. e) Male, Park Tayabas, Luzon. f) Lectotype, male, Samar. g) Labrum, Male. h) Labrum, Female. Scale = 1 mm." captionText-1="FIGURE 3. Aedeagus of Prothyma (Symplecthyma) aeneoparva Horn, 1934, left lateral view. a) Syntype, Biliran; b) Lope DeVega, Samar, lateral and ventral view; c) Syntype, Samar; d) Hinabangan, Samar; e) P. nitida Rivalier 1964 syn. nov., Type, drawing from (Rivalier, 1964); f) Segod, Leyte; g) Park Tayabas, Luzon; h) Surigao, Mindanao; i) Wao, Mindanao; j) Sibagat; k) Cawa-Cawa, Mindanao; l) Gutalac, Mindanao; m) Cabanglasan, Mindanao; n) Esperanza, Mindanao. Scale = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10012553" figureDoi-1="http://doi.org/10.5281/zenodo.10012559" httpUri-0="https://zenodo.org/record/10012553/files/figure.png" httpUri-1="https://zenodo.org/record/10012559/files/figure.png" pageId="2" pageNumber="73">Figs 2–3</figureCitation>
)....................................................................................................
<taxonomicName id="4C124D41FFBB2E2EFBE494AEFA3A0A6B" authority="Horn, 1934" authorityName="W. Horn" authorityYear="1934" baseAuthorityName=": Wiesner" baseAuthorityYear="1980" box="[1136,1436,1268,1290]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="aeneoparva" subGenus="Symplecthyma">
<emphasis id="B966EAD0FFBB2E2EFBE494AEFA3A0A6B" bold="true" box="[1136,1436,1268,1290]" pageId="2" pageNumber="73">
<emphasis id="B966EAD0FFBB2E2EFBE494AEFB240A68" bold="true" box="[1136,1154,1269,1290]" italics="true" pageId="2" pageNumber="73">P.</emphasis>
(
<emphasis id="B966EAD0FFBB2E2EFB0494AFFB050A6B" bold="true" box="[1168,1187,1268,1289]" italics="true" pageId="2" pageNumber="73">S.</emphasis>
)
<emphasis id="B966EAD0FFBB2E2EFB2594AEFA820A68" bold="true" box="[1201,1316,1269,1290]" italics="true" pageId="2" pageNumber="73">aeneoparva</emphasis>
<bibRefCitation id="EF834B33FFBB2E2EFABE94AEFA3A0A6B" author="Horn, W." box="[1322,1436,1268,1290]" pageId="2" pageNumber="73" pagination="144 - 146" refId="ref16811" refString="Horn, W. (1934) Neues uber das Genus Prothyma (Cicind.: Col.). Natuurhistorisch Maandblad, 23, 144 - 146." type="journal article" year="1934">Horn, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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