treatments-xml/data/03/BB/87/03BB87BEFFDAAE23FED19677FB29F89F.xml

<document id="7AC2A3DD6115415D43397134E0B808F5" ID-CLB-Dataset="27022" ID-DOI="10.5281/zenodo.2646265" ID-GBIF-Dataset="9cfb392b-d6e9-431d-bcc4-598698ad1763" ID-ISSN="1175-5326" ID-Zenodo-Dep="2646265" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555587978888" checkinUser="plazi" docAuthor="Holzenthal, Ralph W. &amp; Blahnik, Roger J." docDate="2006" docId="03BB87BEFFDAAE23FED19677FB29F89F" docLanguage="en" docName="zt01197p037.pdf" docOrigin="Zootaxa 1197" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Protoptila chitaria Holzenthal &amp; Blahnik 2006, new species" docType="treatment" docVersion="11" lastPageNumber="18" masterDocId="FF82FFC6FFD6AE32FFD99323FF85FFEE" masterDocTitle="The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="13" updateTime="1698722985916" updateUser="ExternalLinkService">
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<mods:title id="4DD8125D6EF4342A9B911F0DB3937CF4">The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)</mods:title>
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<mods:namePart id="7C2FA8DBBFF349ED5CAC5EBCCFAF5F77">Holzenthal, Ralph W.</mods:namePart>
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<mods:namePart id="FB2E8342711F69D7F50E9CC82275F5BD">Blahnik, Roger J.</mods:namePart>
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<mods:date id="C0A213EA56D996B9EB276945902A4827">2006</mods:date>
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<mods:number id="A3D46F33D34F75AEE74957154B1915F1">2006-05-08</mods:number>
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<treatment id="03BB87BEFFDAAE23FED19677FB29F89F" ID-DOI="http://doi.org/10.5281/zenodo.6260777" ID-GBIF-Taxon="156789595" ID-Zenodo-Dep="6260777" LSID="urn:lsid:plazi:treatment:03BB87BEFFDAAE23FED19677FB29F89F" httpUri="http://treatment.plazi.org/id/03BB87BEFFDAAE23FED19677FB29F89F" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13">
<subSubSection id="C3086523FFDAAE3EFED19677FD01FA80" box="[264,644,1364,1390]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BAD36A8FFDAAE3EFED19677FD01FA80" blockId="12.[264,644,1364,1430]" box="[264,644,1364,1390]" pageId="12" pageNumber="13">
<heading id="D0E581C4FFDAAE3EFED19677FD01FA80" bold="true" box="[264,644,1364,1390]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B966EABAFFDAAE3EFED19677FD01FA80" bold="true" box="[264,644,1364,1390]" pageId="12" pageNumber="13">
<taxonomicName id="4C124D2BFFDAAE3EFED19677FE63FA80" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[264,486,1364,1390]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="chitaria" status="sp. nov.">
<emphasis id="B966EABAFFDAAE3EFED19677FE63FA80" bold="true" box="[264,486,1364,1390]" italics="true" pageId="12" pageNumber="13">Protoptila chitaria</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFDAAE3EFE2C9677FD01FA80" box="[501,644,1364,1390]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086523FFDAAE3CFED1965FFDBBFD98" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="8BAD36A8FFDAAE3EFED1965FFECEFA78" blockId="12.[264,644,1364,1430]" box="[264,331,1404,1430]" pageId="12" pageNumber="13">
<figureCitation id="13292A2DFFDAAE3EFED1965FFECEFA78" box="[264,331,1404,1430]" captionStart="FIGURE 8" captionStartId="15.[264,368,1695,1719]" captionTargetBox="[266,1320,569,1666]" captionTargetId="figure@15.[266,1321,569,1677]" captionTargetPageId="15" captionText="FIGURE 8. Protoptila chitaria, new species. Male genitalia: A—lateral (inset: apex of segment X, caudal, enlarged); B—segment VIII, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646285/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFDAAE3EFED196EFFCE5F940" blockId="12.[264,1325,1484,1790]" pageId="12" pageNumber="13">
This species is very similar to 
<taxonomicName id="4C124D2BFFDAAE3EFDAD96EFFD4EFA0B" box="[628,715,1484,1510]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="B966EABAFFDAAE3EFDAD96EFFD07FA0B" box="[628,642,1484,1509]" italics="true" pageId="12" pageNumber="13">P</emphasis>
. 
<emphasis id="B966EABAFFDAAE3EFD4B96EFFD4EFA0B" box="[658,715,1484,1509]" italics="true" pageId="12" pageNumber="13">cana</emphasis>
</taxonomicName>
. They differ in the shape of the apex of the ventral margin of segment IX, which is divided, with each branch partially fused to the lateral margin of sternum VIII. In 
<taxonomicName id="4C124D2BFFDAAE3EFDBC953FFD60F9DB" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[613,741,1564,1590]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="chitaria">
<emphasis id="B966EABAFFDAAE3EFDBC953FFDF6F9DB" box="[613,627,1564,1589]" italics="true" pageId="12" pageNumber="13">P</emphasis>
. 
<emphasis id="B966EABAFFDAAE3EFD51953FFD60F9DB" box="[648,741,1564,1589]" italics="true" pageId="12" pageNumber="13">chitaria</emphasis>
</taxonomicName>
this apex is shorter than the ventral apices of sternum VIII and curved apically, whereas in 
<taxonomicName id="4C124D2BFFDAAE3EFC909567FC21F9B3" box="[841,932,1604,1629]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="B966EABAFFDAAE3EFC909567FC21F9B3" box="[841,932,1604,1629]" italics="true" pageId="12" pageNumber="13">P. cana</emphasis>
</taxonomicName>
it is more elongate and nearly straight. The shapes of the apices of the apical sections of tergum X of the two species are also slightly different, as shown in the illustrations.
</paragraph>
<paragraph id="8BAD36A8FFDAAE3EFEE1959FFD26F910" blockId="12.[264,1325,1484,1790]" pageId="12" pageNumber="13">
Adult. Length of forewing: male 
<quantity id="4CEA9B4DFFDAAE3EFD15959FFCDAF938" box="[716,863,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="12" pageNumber="13" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.2–3.9 mm</quantity>
; female 
<quantity id="4CEA9B4DFFDAAE3EFC11959FFBD8F938" box="[968,1117,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="12" pageNumber="13" unit="mm" value="3.85" valueMax="4.2" valueMin="3.5">3.5–4.2 mm</quantity>
. Color brown (in alcohol), wing markings indistinct.
</paragraph>
<caption id="DF6D6620FFDBAE3FFED1965CFDB2FA57" httpUri="https://zenodo.org/record/2646281/files/figure.svg" pageId="13" pageNumber="14" startId="13.[264,368,1407,1431]" targetBox="[94,1286,286,1395]" targetPageId="13">
<paragraph id="8BAD36A8FFDBAE3FFED1965CFDB2FA57" blockId="13.[264,1322,1407,1465]" pageId="13" pageNumber="14">
<emphasis id="B966EABAFFDBAE3FFED1965CFE09FA79" bold="true" box="[264,396,1407,1431]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
<taxonomicName id="4C124D2BFFDBAE3FFE4F96A3FD1EFA79" authority=", Flint." authorityName="Flint." box="[406,667,1408,1431]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="burica">
<emphasis id="B966EABAFFDBAE3FFE4F96A3FDD4FA79" box="[406,593,1408,1431]" italics="true" pageId="13" pageNumber="14">Protoptila burica</emphasis>
, Flint.
</taxonomicName>
Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
<paragraph id="8BAD36A8FFDBAE3CFEE196CEFDBBFD98" blockId="13.[264,1324,1517,1903]" lastBlockId="14.[264,1323,284,630]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Male genitalia. Sternum VI process prominent, slightly curved, longer than wide, apex subacute. Tergum VIII posterior margin with row of elongate setae; sternum VIII produced ventrally, distinctly bifurcate apically, branches acute apically, broad basally, relatively short. Segment IX with anterior margin broadly rounded; sternum IX laterally produced posteriorly, forming paired sinuous processes, partially fused to lateral margin of sternum VIII, processes not extending beyond posterior margin of sternum VIII; posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X divided mesally to form sclerotized lateral parts with basal and apical sections; basal section subquadrate, about as long as wide, apical section longer than basal section, apically divided to form broadly rounded ventral lobe and acute dorsal and mesal lobes. Phallobase dorsally with large, semicircular, laterally compressed apodeme; ventrally with short, rodlike, articulated appendages with apical setae, appendages fitting into sclerotized pockets on ventral margin of phallobase; posteroventral margin of phallobase with short, paired, upturned, apically acute basal processes and also dorsolateral processes; dorsolateral processes very short, straight, acute; paramere absent; phallicata short, narrowest basally, apex enlarged, membranous laterally, sclerotized ventrolaterally to form paired, acute, ventrally curved projection; endophallus membranous, enlarged when everted, with small apical and subapical spines; phallotremal sclerite spine­like, sinuous, sclerotized apically, somewhat divided basally.</paragraph>
</subSubSection>
<caption id="DF6D6620FFD8AE3CFED1959BFCD3F91C" httpUri="https://zenodo.org/record/2646283/files/figure.png" pageId="14" pageNumber="15" startId="14.[264,367,1720,1744]" targetBox="[283,1307,681,1693]" targetPageId="14">
<paragraph id="8BAD36A8FFD8AE3CFED1959BFAAEF921" blockId="14.[264,1323,1720,1778]" box="[264,1323,1720,1744]" pageId="14" pageNumber="15">
<emphasis id="B966EABAFFD8AE3CFED1959BFE0EF93E" bold="true" box="[264,395,1720,1744]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
<taxonomicName id="4C124D2BFFD8AE3CFE4D959AFD00F921" authority=", Flint." authorityName="Flint." box="[404,645,1720,1744]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="B966EABAFFD8AE3CFE4D959AFDB9F93E" box="[404,572,1721,1744]" italics="true" pageId="14" pageNumber="15">Protoptila cana</emphasis>
, Flint.
</taxonomicName>
Male genitalia: A—lateral (inset: apex of segment X, caudal);
</paragraph>
<paragraph id="8BAD36A8FFD8AE3CFED195F8FCD3F91C" blockId="14.[264,1323,1720,1778]" box="[264,854,1755,1778]" pageId="14" pageNumber="15">B—segment VIII, ventral; C—phallic complex, lateral.</paragraph>
</caption>
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<paragraph id="8BAD36A8FFD8AE3DFEE19406FCE7FED8" blockId="14.[264,1323,1829,1896]" lastBlockId="15.[264,1321,284,470]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<materialsCitation id="3B7A3CF5FFD8AE3CFEE19406FAAEF8AE" ID-GBIF-Occurrence="2236573119" box="[312,1323,1829,1856]" collectorName="NW Rio" country="Costa Rica" location="Rio Chitaria" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<emphasis id="B966EABAFFD8AE3CFEE19406FD4FF8D1" bold="true" box="[312,714,1829,1855]" pageId="14" pageNumber="15">
<typeStatus id="54A9880AFFD8AE3CFEE19406FE2FF8D1" box="[312,426,1829,1855]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="F3057638FFD8AE3CFDD49406FD4FF8D1" box="[525,714,1829,1855]" name="Costa Rica" pageId="14" pageNumber="15">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFD8AE3CFD389406FCC2F8D1" bold="true" box="[737,839,1829,1855]" pageId="14" pageNumber="15">
<collectingRegion id="49D6F84AFFD8AE3CFD389406FCC2F8D1" box="[737,839,1829,1855]" country="Costa Rica" name="Cartago" pageId="14" pageNumber="15">Cartago</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFD8AE3CFC879405FC7CF8AE" LSID="urn:lsid:plazi:treatment:03BB87BEFFDAAE23FED19677FB29F89F:8ECD6073FFD8AE3CFC879405FC7CF8AE" box="[862,1017,1830,1856]" country="Costa Rica" name="Rio Chitaria" pageId="14" pageNumber="15" stateProvince="Cartago">Río Chitaría</location>
, rt 10, 10 km 
<collectorName id="26E7537EFFD8AE3CFB189405FAAEF8AE" box="[1217,1323,1830,1856]" pageId="14" pageNumber="15">NW Río</collectorName>
</materialsCitation>
Reventazón, 
<geoCoordinate id="EE26506FFFD8AE3CFE73946DFDB9F886" box="[426,572,1870,1896]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="9.92">09°55’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFD8AE3CFD88946DFD70F886" box="[593,757,1870,1896]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="-83.60389">083°36’14”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFD8AE3CFCEE946DFC09F886" box="[823,908,1870,1896]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="14" pageNumber="15" unit="m" value="740.0">740 m</quantity>
, 
<date id="FFAC1068FFD8AE3CFC78946DFBA6F886" box="[929,1059,1870,1896]" pageId="14" pageNumber="15" value="1991-03-21">21.iii.1991</date>
, Holzenthal, Muñoz, Huisman (in alcohol) (UMSP000208545) (UMSP).
</paragraph>
<paragraph id="8BAD36A8FFD9AE3DFEE19260FB2AFE68" blockId="15.[264,1321,284,470]" pageId="15" pageNumber="16">
<emphasis id="B966EABAFFD9AE3DFEE19260FE30FEB3" bold="true" box="[312,437,323,349]" pageId="15" pageNumber="16">
<typeStatus id="54A9880AFFD9AE3DFEE19260FE30FEB3" box="[312,437,323,349]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as 
<typeStatus id="54A9880AFFD9AE3DFDAD9267FD5EFEB0" box="[628,731,324,350]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
— 
<materialsCitation id="3B7A3CF5FFD9AE3DFCD69267FA8EFEB0" ID-GBIF-Occurrence="2236573158" box="[783,1291,324,350]" collectionCode="UMSP" pageId="15" pageNumber="16" specimenCount="71" specimenCount-female="66" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="9D14FD21FFD9AE3DFCD69267FCF5FEB0" box="[783,880,324,350]" pageId="15" pageNumber="16" type="male">5 males</specimenCount>
, 
<specimenCount id="9D14FD21FFD9AE3DFCA69267FB86FEB0" box="[895,1027,324,350]" pageId="15" pageNumber="16" type="female">66 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFD9AE3DFB749267FA87FEB0" box="[1197,1282,324,350]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="15" pageNumber="16" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFD9AE3DFAC39267FD57FE68" ID-GBIF-Occurrence="2236573113" collectionCode="NMNH" pageId="15" pageNumber="16" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D14FD21FFD9AE3DFAC39267FECAFE68" pageId="15" pageNumber="16" type="male">2 males</specimenCount>
, 
<specimenCount id="9D14FD21FFD9AE3DFE82924FFE49FE68" box="[347,460,364,390]" pageId="15" pageNumber="16" type="female">4 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFD9AE3DFDB3924FFD4FFE68" box="[618,714,364,390]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFD9AE3DFD07924FFB2FFE68" ID-GBIF-Occurrence="2236573153" box="[734,1194,364,390]" collectionCode="INBIO" pageId="15" pageNumber="16" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D14FD21FFD9AE3DFD07924FFCB5FE68" box="[734,816,364,390]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
, 
<specimenCount id="9D14FD21FFD9AE3DFCE2924FFC29FE68" box="[827,940,364,390]" pageId="15" pageNumber="16" type="female">3 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFD9AE3DFB93924FFB27FE68" box="[1098,1186,364,390]" pageId="15" pageNumber="16">INBIO</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36A8FFD9AE3DFEE192B7FC04FE38" blockId="15.[264,1321,284,470]" pageId="15" pageNumber="16">
Etymology. This species is named for Chitaría, the town and river in 
<collectingCountry id="F3057638FFD9AE3DFBB492B7FB6BFE40" box="[1133,1262,404,430]" name="Costa Rica" pageId="15" pageNumber="16">Costa Rica</collectingCountry>
near which the 
<typeStatus id="54A9880AFFD9AE3DFE5B929FFE30FE38" box="[386,437,444,470]" pageId="15" pageNumber="16">type</typeStatus>
material for this species was collected.
</paragraph>
<caption id="DF6D6620FFD9AE3DFED195BCFB9EF937" httpUri="https://zenodo.org/record/2646285/files/figure.png" pageId="15" pageNumber="16" startId="15.[264,368,1695,1719]" targetBox="[266,1320,569,1666]" targetPageId="15">
<paragraph id="8BAD36A8FFD9AE3DFED195BCFB9EF937" blockId="15.[264,1322,1695,1753]" pageId="15" pageNumber="16">
<emphasis id="B966EABAFFD9AE3DFED195BCFE0EF959" bold="true" box="[264,395,1695,1719]" pageId="15" pageNumber="16">FIGURE 8.</emphasis>
<taxonomicName id="4C124D2BFFD9AE3DFE489583FDDDF959" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[401,600,1696,1719]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chitaria" status="sp. nov.">
<emphasis id="B966EABAFFD9AE3DFE489583FDDDF959" box="[401,600,1696,1719]" italics="true" pageId="15" pageNumber="16">Protoptila chitaria</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFD9AE3DFDBF9583FD6CF959" box="[614,745,1696,1719]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral (inset: apex of segment X, caudal, enlarged); B—segment VIII, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
<caption id="DF6D6620FFC6AE22FED1940CFDB2F884" httpUri="https://zenodo.org/record/2646287/files/figure.svg" pageId="16" pageNumber="17" startId="16.[264,368,1839,1863]" targetBox="[302,1493,285,1824]" targetPageId="16">
<paragraph id="8BAD36A8FFC6AE22FED1940CFAAEF8A9" blockId="16.[264,1323,1839,1898]" box="[264,1323,1839,1863]" pageId="16" pageNumber="17">
<emphasis id="B966EABAFFC6AE22FED1940CFE0CF8A9" bold="true" box="[264,393,1839,1863]" pageId="16" pageNumber="17">FIGURE 9.</emphasis>
<taxonomicName id="4C124D2BFFC6AE22FE499413FDD1F8A9" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[400,596,1840,1863]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cristula" status="sp. nov.">
<emphasis id="B966EABAFFC6AE22FE499413FDD1F8A9" box="[400,596,1840,1863]" italics="true" pageId="16" pageNumber="17">Protoptila cristula</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFC6AE22FDB89413FD60F8A9" box="[609,741,1840,1863]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral; B—segment VIII, ventral;
</paragraph>
<paragraph id="8BAD36A8FFC6AE22FED19470FDB2F884" blockId="16.[264,1323,1839,1898]" box="[264,567,1875,1898]" pageId="16" pageNumber="17">C—phallic complex, lateral.</paragraph>
</caption>
<paragraph id="8BAD36A8FFC7AE23FED1923FFD07FEDB" blockId="17.[264,642,283,345]" box="[264,642,283,310]" pageId="17" pageNumber="18">
<emphasis id="B966EABAFFC7AE23FED1923FFD07FEDB" bold="true" box="[264,642,283,310]" pageId="17" pageNumber="18">
<taxonomicName id="4C124D2BFFC7AE23FED1923FFE66FED8" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[264,483,284,310]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cristula" status="sp. nov.">
<emphasis id="B966EABAFFC7AE23FED1923FFE66FED8" bold="true" box="[264,483,284,310]" italics="true" pageId="17" pageNumber="18">Protoptila cristula</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFC7AE23FE2B9238FD07FEDB" box="[498,642,283,309]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FED1921CFECEFEB7" blockId="17.[264,642,283,345]" box="[264,331,319,345]" pageId="17" pageNumber="18">
<figureCitation id="13292A2DFFC7AE23FED1921CFECEFEB7" box="[264,331,319,345]" captionStart="FIGURE 9" captionStartId="16.[264,368,1839,1863]" captionTargetBox="[302,1493,285,1824]" captionTargetId="figure@16.[302,1279,307,1824]" captionTargetPageId="16" captionText="FIGURE 9. Protoptila cristula, new species. Male genitalia: A—lateral; B—segment VIII, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646287/files/figure.svg" pageId="17" pageNumber="18">Fig. 9</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FED192ACFE02FD2F" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">
This species is closely related to 
<taxonomicName id="4C124D2BFFC7AE23FD4592ACFD6CFE46" authorityName="Mosely" authorityYear="1937" box="[668,745,399,424]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rota">
<emphasis id="B966EABAFFC7AE23FD4592ACFD6CFE46" box="[668,745,399,424]" italics="true" pageId="17" pageNumber="18">P. rota</emphasis>
</taxonomicName>
, and both are similar to 
<taxonomicName id="4C124D2BFFC7AE23FBC592ACFB2AFE46" box="[1052,1199,399,424]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="B966EABAFFC7AE23FBC592ACFB2AFE46" box="[1052,1199,399,424]" italics="true" pageId="17" pageNumber="18">P. bicornuta</emphasis>
</taxonomicName>
. All three have horn­like processes or “ 
<emphasis id="B966EABAFFC7AE23FDBF9294FD3FFE3E" box="[614,698,439,464]" italics="true" pageId="17" pageNumber="18">cornuti</emphasis>
” projecting from the phallicata. In 
<taxonomicName id="4C124D2BFFC7AE23FBBC9294FB58FE3E" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[1125,1245,439,464]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cristula">
<emphasis id="B966EABAFFC7AE23FBBC9294FB58FE3E" box="[1125,1245,439,464]" italics="true" pageId="17" pageNumber="18">P. cristula</emphasis>
</taxonomicName>
, these structures are more upright and the spine­like structures at the base of these horns are much more elongate than in either 
<taxonomicName id="4C124D2BFFC7AE23FD449124FD6FFDCE" authorityName="Mosely" authorityYear="1937" box="[669,746,519,544]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rota">
<emphasis id="B966EABAFFC7AE23FD449124FD6FFDCE" box="[669,746,519,544]" italics="true" pageId="17" pageNumber="18">P. rota</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C124D2BFFC7AE23FCC99124FC1BFDCE" box="[784,926,519,544]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="B966EABAFFC7AE23FCC99124FC1BFDCE" box="[784,926,519,544]" italics="true" pageId="17" pageNumber="18">P. bicornuta</emphasis>
</taxonomicName>
. In both 
<taxonomicName id="4C124D2BFFC7AE23FBDF9124FBF9FDCE" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[1030,1148,519,544]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cristula">
<emphasis id="B966EABAFFC7AE23FBDF9124FBF9FDCE" box="[1030,1148,519,544]" italics="true" pageId="17" pageNumber="18">P. cristula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C124D2BFFC7AE23FB6D9124FB79FDCE" authorityName="Mosely" authorityYear="1937" box="[1204,1276,519,544]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rota">
<emphasis id="B966EABAFFC7AE23FB6D9124FB79FDCE" box="[1204,1276,519,544]" italics="true" pageId="17" pageNumber="18">P. rota</emphasis>
</taxonomicName>
the ventral margin of sternum VIII is much more elongate and upcurved apically than 
<taxonomicName id="4C124D2BFFC7AE23FACC910CFEFFFD9E" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="B966EABAFFC7AE23FACC910CFEFFFD9E" italics="true" pageId="17" pageNumber="18">P. bicornuta</emphasis>
</taxonomicName>
, as well as more deeply bifurcate apicomesally. 
<taxonomicName id="4C124D2BFFC7AE23FC629174FB10FD9E" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[955,1173,599,624]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cristula">
<emphasis id="B966EABAFFC7AE23FC629174FB10FD9E" box="[955,1173,599,624]" italics="true" pageId="17" pageNumber="18">Protoptila cristula</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C124D2BFFC7AE23FED1915CFEDDFD76" authorityName="Mosely" authorityYear="1937" box="[264,344,639,664]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rota">
<emphasis id="B966EABAFFC7AE23FED1915CFEDDFD76" box="[264,344,639,664]" italics="true" pageId="17" pageNumber="18">P. rota</emphasis>
</taxonomicName>
in that the lateral processes of tergum X are also much more elongate and projecting.
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FEE191ECFBA3FCFF" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">
Adult. Length of forewing: male 
<quantity id="4CEA9B4DFFC7AE23FD1A91ECFCD1FD07" box="[707,852,719,745]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="17" pageNumber="18" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
; female 
<quantity id="4CEA9B4DFFC7AE23FC6391ECFBC9FD07" box="[954,1100,719,745]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="17" pageNumber="18" unit="mm" value="2.9" valueMax="3.2" valueMin="2.6">2.6–3.2 mm</quantity>
. Color light brown (in alcohol), with distinct transverse bar of whitish setae along cord.
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FEE1903CFB87FA0F" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">Male genitalia. Sternum VI process prominent, slightly curved, longer than wide, apex subacute. Tergum VIII posterior margin with row of elongate setae; sternum VIII strongly produced ventrally, distinctly bifurcate apically, branches acute apically, narrow, digitate, very elongate and strongly upturned. Segment IX with anterior margin broadly rounded; sternum IX medially produced posteriorly, in ventral view, narrow basally, broad apically, apically emarginate; posterolateral margin of segment IX without processes. Preanal appendages small, round. Tergum X divided mesally to form sclerotized lateral parts with basal and apical sections; basal section subquadrate, about as long as wide, apical section elongate, narrow, with rounded basodorsal protuberance. Phallobase dorsally with large, semicircular, laterally compressed apodeme; ventrally with short, rod­like, articulated appendages with apical setae, appendages fitting into sclerotized pockets on ventral margin of phallobase; posteroventral margin of phallobase with very short, paired, upturned, apically acute basal processes and also dorsolateral processes; dorsolateral processes prominent, slender, acute, angularly upturned near base; paramere very short, membranous, apparently vestigial; phallicata short, narrow and angularly bent near middle, with large, paired upright horn­like dorsolateral processes and very elongate, sinuous, narrow, apically acute ventral processes; endophallus membranous, enlarged when everted; phallotremal sclerite lightly sclerotized, indistinct.</paragraph>
<paragraph id="8BAD36A8FFC7AE23FEE196CDFD87F9B7" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">
<materialsCitation id="3B7A3CF5FFC7AE23FEE196CDFD87F9B7" ID-GBIF-Occurrence="2236573137" collectingDate="1986-03-12" collectionCode="UMSP" collectorName="S Brasilia &amp; Holzenthal &amp; Fasth" country="Costa Rica" elevation="390" latitude="10.971944" location="Rio Pizote" longLatPrecision="21" longitude="-85.345" pageId="17" pageNumber="18" specimenCode="UMSP000019033" specimenCount="1" stateProvince="Alajuela" typeStatus="holotype">
<emphasis id="B966EABAFFC7AE23FEE196CDFE76F9E6" bold="true" box="[312,499,1518,1544]" pageId="17" pageNumber="18">
<typeStatus id="54A9880AFFC7AE23FEE196CDFE2FF9E6" box="[312,426,1518,1544]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
male
</emphasis>
: 
<emphasis id="B966EABAFFC7AE23FDD196CDFD41F9E6" bold="true" box="[520,708,1518,1544]" pageId="17" pageNumber="18">
<collectingCountry id="F3057638FFC7AE23FDD196CDFD41F9E6" box="[520,708,1518,1544]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFC7AE23FD0096CDFCC6F9E6" bold="true" box="[729,835,1518,1544]" pageId="17" pageNumber="18">
<collectingRegion id="49D6F84AFFC7AE23FD0096CDFCC6F9E6" box="[729,835,1518,1544]" country="Costa Rica" name="Alajuela" pageId="17" pageNumber="18">Alajuela</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFC7AE23FC8196CCFC59F9E7" LSID="urn:lsid:plazi:treatment:03BB87BEFFDAAE23FED19677FB29F89F:8ECD6073FFC7AE23FC8196CCFC59F9E7" box="[856,988,1519,1545]" country="Costa Rica" latitude="10.971944" longLatPrecision="21" longitude="-85.345" name="Rio Pizote" pageId="17" pageNumber="18" stateProvince="Alajuela">Río Pizote</location>
, ca. 
<quantity id="4CEA9B4DFFC7AE23FBC596CCFBD8F9E7" box="[1052,1117,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="5.0">5 km</quantity>
(air) 
<collectorName id="26E7537EFFC7AE23FB7296CCFAA3F9E7" box="[1195,1318,1519,1545]" pageId="17" pageNumber="18">S Brasilia</collectorName>
, 
<geoCoordinate id="EE26506FFFC7AE23FED19534FE1FF9DF" box="[264,410,1559,1585]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="10.971944">10°58’19”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFC7AE23FE7C9534FDCFF9DF" box="[421,586,1559,1585]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-85.345">085°20’42”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFC7AE23FDAD9534FD3AF9DF" box="[628,703,1559,1585]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="17" pageNumber="18" unit="m" value="390.0">
<elevation id="003FD19BFFC7AE23FDAD9534FD3AF9DF" box="[628,703,1559,1585]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="17" pageNumber="18" unit="m" value="390.0">390 m</elevation>
</quantity>
, 
<date id="FFAC1068FFC7AE23FD139534FCC9F9DF" box="[714,844,1559,1585]" pageId="17" pageNumber="18" value="1986-03-12">
<collectingDate id="EFE8E980FFC7AE23FD139534FCC9F9DF" box="[714,844,1559,1585]" pageId="17" pageNumber="18" value="1986-03-12">12.iii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFC7AE23FC819534FC5DF9DF" box="[856,984,1559,1585]" pageId="17" pageNumber="18">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFC7AE23FC249534FBB8F9DF" box="[1021,1085,1559,1585]" pageId="17" pageNumber="18">Fasth</collectorName>
(
<specimenCode id="DBB49ED3FFC7AE23FB929534FAA6F9DF" box="[1099,1315,1559,1585]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="17" pageNumber="18" type="University or college">UMSP000019033</specimenCode>
) (in alcohol) (
<collectionCode id="ED03AE6DFFC7AE23FE46951CFE70F9B7" box="[415,501,1599,1625]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="17" pageNumber="18" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FEE19545FC42F8CF" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">
<materialsCitation id="3B7A3CF5FFC7AE23FEE19545FAA0F947" ID-GBIF-Occurrence="2236573118" collectionCode="UMSP" collectorName="Holzenthal &amp; Fasth" country="Costa Rica" elevation="470" latitude="10.948055" location="Rio Pizote" longLatPrecision="21" longitude="-85.29111" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<emphasis id="B966EABAFFC7AE23FEE19545FE30F96E" bold="true" box="[312,437,1638,1664]" pageId="17" pageNumber="18">
<typeStatus id="54A9880AFFC7AE23FEE19545FE30F96E" box="[312,437,1638,1664]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<emphasis id="B966EABAFFC7AE23FE1C9545FDFFF96E" bold="true" box="[453,634,1638,1664]" pageId="17" pageNumber="18">
<collectingCountry id="F3057638FFC7AE23FE1C9545FDFFF96E" box="[453,634,1638,1664]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFC7AE23FD539545FD71F96E" bold="true" box="[650,756,1638,1664]" pageId="17" pageNumber="18">
<collectingRegion id="49D6F84AFFC7AE23FD539545FD71F96E" box="[650,756,1638,1664]" country="Costa Rica" name="Alajuela" pageId="17" pageNumber="18">Alajuela</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFC7AE23FCDD9544FC07F96F" LSID="urn:lsid:plazi:treatment:03BB87BEFFDAAE23FED19677FB29F89F:8ECD6073FFC7AE23FCDD9544FC07F96F" box="[772,898,1639,1665]" country="Costa Rica" latitude="10.948055" longLatPrecision="21" longitude="-85.29111" name="Rio Pizote" pageId="17" pageNumber="18" stateProvince="Alajuela">Río Pizote</location>
, ca. 
<locationDeviation id="9A1F2F8AFFC7AE23FC6E9544FB0DF96F" box="[951,1160,1639,1665]" location="Dos Rios" pageId="17" pageNumber="18">
<quantity id="4CEA9B4DFFC7AE23FC6E9544FC71F96F" box="[951,1012,1639,1665]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="5.0">5 km</quantity>
N Dos Ríos
</locationDeviation>
, 
<geoCoordinate id="EE26506FFFC7AE23FB4D9544FAA3F96F" box="[1172,1318,1639,1665]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="10.948055">10°56’53”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFC7AE23FED195ACFE29F947" box="[264,428,1679,1705]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-85.29111">085°17’28”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFC7AE23FE2B95ACFDCEF947" box="[498,587,1679,1705]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="17" pageNumber="18" unit="m" value="470.0">
<elevation id="003FD19BFFC7AE23FE2B95ACFDCEF947" box="[498,587,1679,1705]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="17" pageNumber="18" unit="m" value="470.0">470 m</elevation>
</quantity>
, 
<collectorName id="26E7537EFFC7AE23FDBB95ACFD67F947" box="[610,738,1679,1705]" pageId="17" pageNumber="18">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFC7AE23FCC795ACFCDBF947" box="[798,862,1679,1705]" pageId="17" pageNumber="18">Fasth</collectorName>
— 
<specimenCount id="9D14FD21FFC7AE23FC7995ACFB83F947" box="[928,1030,1679,1705]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFC7AE23FB1F95ACFA99F947" box="[1222,1308,1679,1705]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="17" pageNumber="18" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B7A3CF5FFC7AE23FED19595FE50F8CF" ID-GBIF-Occurrence="2236573146" collectionCode="UMSP" collectorName="Chamorro &amp; Lopez" country="Nicaragua" elevation="63" latitude="11.7027" location="Reserva Silvestre Privada Domitila" longLatPrecision="1" longitude="-85.951965" pageId="17" pageNumber="18" specimenCount="170" specimenCount-female="110" specimenCount-male="60" typeStatus="paratype">
<emphasis id="B966EABAFFC7AE23FED19595FDD7F93E" bold="true" box="[264,594,1718,1744]" pageId="17" pageNumber="18">
<collectingCountry id="F3057638FFC7AE23FED19595FE46F93E" box="[264,451,1718,1744]" name="Nicaragua" pageId="17" pageNumber="18">NICARAGUA</collectingCountry>
: Granada:
</emphasis>
<location id="8ECD6073FFC7AE23FDBB9594FB98F93F" LSID="urn:lsid:plazi:treatment:03BB87BEFFDAAE23FED19677FB29F89F:8ECD6073FFC7AE23FDBB9594FB98F93F" box="[610,1053,1719,1745]" country="Nicaragua" latitude="11.7027" longLatPrecision="1" longitude="-85.951965" name="Reserva Silvestre Privada Domitila" pageId="17" pageNumber="18">Reserva Silvestre Privada Domitila</location>
, río cerca manantial, 
<geoCoordinate id="EE26506FFFC7AE23FED195FCFE27F917" box="[264,418,1759,1785]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="11.7027">11°42.162’N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFC7AE23FE6895FCFDD5F917" box="[433,592,1759,1785]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-85.951965">85°57.118’W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFC7AE23FD5C95FCFD42F917" box="[645,711,1759,1785]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="17" pageNumber="18" unit="m" value="63.0">
<elevation id="003FD19BFFC7AE23FD5C95FCFD42F917" box="[645,711,1759,1785]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="17" pageNumber="18" unit="m" value="63.0">63 m</elevation>
</quantity>
, 
<collectorName id="26E7537EFFC7AE23FD0F95FCFCC8F917" box="[726,845,1759,1785]" pageId="17" pageNumber="18">Chamorro</collectorName>
&amp; 
<collectorName id="26E7537EFFC7AE23FCA195FCFC44F917" box="[888,961,1759,1785]" pageId="17" pageNumber="18">López</collectorName>
— 
<specimenCount id="9D14FD21FFC7AE23FC2A95FCFBE7F917" box="[1011,1122,1759,1785]" pageId="17" pageNumber="18" type="male">60 males</specimenCount>
, 
<specimenCount id="9D14FD21FFC7AE23FBA895FCFA84F917" box="[1137,1281,1759,1785]" pageId="17" pageNumber="18" type="female">110 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFC7AE23FEAE9424FE48F8CF" box="[375,461,1799,1825]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="17" pageNumber="18" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFC7AE23FE3B9424FC46F8CF" ID-GBIF-Occurrence="2236573112" box="[482,963,1799,1825]" collectionCode="NMNH" collectorName="Chamorro &amp; Lopez" country="Grenada" elevation="63" isTailEnumeration="true" latitude="11.7027" location="Reserva Silvestre Privada Domitila" longLatPrecision="1" longitude="-85.951965" pageId="17" pageNumber="18" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D14FD21FFC7AE23FE3B9424FDBAF8CF" box="[482,575,1799,1825]" pageId="17" pageNumber="18" type="male">3 males</specimenCount>
, 
<specimenCount id="9D14FD21FFC7AE23FD929424FD39F8CF" box="[587,700,1799,1825]" pageId="17" pageNumber="18" type="female">3 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFC7AE23FC839424FC3EF8CF" box="[858,955,1799,1825]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36A8FFC7AE23FEE1940CFB29F89F" blockId="17.[264,1325,399,1905]" pageId="17" pageNumber="18">
Etymology. This species is named 
<taxonomicName id="4C124D2BFFC7AE23FD0C940CFCABF8A6" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[725,814,1839,1864]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cristula">
<emphasis id="B966EABAFFC7AE23FD0C940CFCABF8A6" box="[725,814,1839,1864]" italics="true" pageId="17" pageNumber="18">cristula</emphasis>
</taxonomicName>
, a diminutive of the Latin 
<emphasis id="B966EABAFFC7AE23FBAA940CFB30F8A6" box="[1139,1205,1839,1864]" italics="true" pageId="17" pageNumber="18">crista</emphasis>
, meaning “crest,” and referring to the crest­like processes of the phallicata of this species.
</paragraph>
</subSubSection>
</treatment>
</document>