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<document id="B2C8F86DA748834F1289754BFA5CD530" ID-DOI="10.11646/zootaxa.5514.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849780" ID-ZooBank="E047925A-2056-444B-9467-122F66BE9BE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727441596547" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03BB87AA0B576364FF082642FB1DFE46" docLanguage="en" docName="zootaxa.5514.3.2.pdf" docOrigin="Zootaxa 5514 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Tetramelus) mediorufus Jałoszyński, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="254" masterDocId="FF82FFD20B436372FF9F2221FFBAFF97" masterDocTitle="Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="256" masterPageNumber="232" pageNumber="252" updateTime="1727443808472" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75468C027C21E1FF4E3FDB089D67C1FE">Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="9B77A0BA1D61674CC42DA1FA7061722D">2024</mods:date>
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<treatment id="03BB87AA0B576364FF082642FB1DFE46" ID-DOI="http://doi.org/10.5281/zenodo.13849916" ID-Zenodo-Dep="13849916" LSID="urn:lsid:plazi:treatment:03BB87AA0B576364FF082642FB1DFE46" httpUri="http://treatment.plazi.org/id/03BB87AA0B576364FF082642FB1DFE46" lastPageId="22" lastPageNumber="254" pageId="20" pageNumber="252">
<subSubSection id="C30865370B576366FF082642FD11FBEA" box="[151,683,1122,1149]" pageId="20" pageNumber="252" type="nomenclature">
<paragraph id="8BAD36BC0B576366FF082642FD11FBEA" blockId="20.[151,683,1122,1185]" box="[151,683,1122,1149]" pageId="20" pageNumber="252">
<heading id="D0E581D00B576366FF082642FD11FBEA" bold="true" box="[151,683,1122,1149]" fontSize="11" level="1" pageId="20" pageNumber="252" reason="1">
<emphasis id="B966EAAE0B576366FF082642FD11FBEA" bold="true" box="[151,683,1122,1149]" pageId="20" pageNumber="252">
<taxonomicName id="4C124D3F0B576366FF082642FDFCFBEA" authorityName="Jałoszyński" authorityYear="2024" box="[151,582,1122,1149]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="252" phylum="Arthropoda" rank="species" species="mediorufus" status="sp. nov." subGenus="Tetramelus">
<emphasis id="B966EAAE0B576366FF082642FEA9FBEA" bold="true" box="[151,275,1123,1149]" italics="true" pageId="20" pageNumber="252">Euconnus</emphasis>
(
<emphasis id="B966EAAE0B576366FEBB2643FE10FBEA" bold="true" box="[292,426,1122,1149]" italics="true" pageId="20" pageNumber="252">Tetramelus</emphasis>
)
<emphasis id="B966EAAE0B576366FE242643FDFCFBEA" bold="true" box="[443,582,1122,1149]" italics="true" pageId="20" pageNumber="252">mediorufus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557D50B576366FDD22642FD11FBEA" box="[589,683,1123,1149]" pageId="20" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C30865370B576366FF0826A6FE1FFB36" box="[151,421,1159,1185]" pageId="20" pageNumber="252" type="description">
<paragraph id="8BAD36BC0B576366FF0826A6FE1FFB36" blockId="20.[151,683,1122,1185]" box="[151,421,1159,1185]" pageId="20" pageNumber="252">
(
<figureCitation id="13292A390B576366FF0026A6FF4DFB36" box="[159,247,1159,1185]" captionStart="FIGURES 1320" captionStartId="7.[151,264,1725,1749]" captionTargetBox="[157,1430,187,1695]" captionTargetId="figure-19@7.[151,1436,181,1701]" captionTargetPageId="7" captionText="FIGURES 1320. Habitus in dorsal view, males. Euconnus (Tetramelus) punctator sp. nov. (13), E. (Tetramelus) aurilobatus sp. nov. (14), E. (Tetramelus) magnirufus sp. nov. (15), E. (Tetramelus) tetraspinosus sp. nov. (16), E. (Tetramelus) gansuminor sp. nov. (17), E. (Tetramelus) microrufus sp. nov. (18), E. (Tetramelus) sinespinosus sp. nov. (19), and E. (Tetramelus) mediorufus sp. nov. (20)." figureDoi="http://doi.org/10.5281/zenodo.13849790" httpUri="https://zenodo.org/record/13849790/files/figure.png" pageId="20" pageNumber="252">Figs 20</figureCitation>
,
<figureCitation id="13292A390B576366FE9D26A6FEF1FB36" box="[258,331,1159,1185]" captionStart="FIGURES 2130" captionStartId="8.[151,264,1444,1468]" captionTargetBox="[156,1435,316,1415]" captionTargetId="figure-72@8.[151,1435,309,1421]" captionTargetPageId="8" captionText="FIGURES 2130. Male protrochanter (24) and metatrochanter (2123, 2530). Euconnus (Tetramelus) punctator sp. nov. (21), E. (Tetramelus) aurilobatus sp. nov. (22), E. (Tetramelus) magnirufus sp. nov. (23), E. (Tetramelus) tetraspinosus sp. nov. (24, 25), E. (Tetramelus) gansuminor sp. nov. (26), E. (Tetramelus) microrufus sp. nov. (27), E. (Tetramelus) sinespinosus sp. nov. (28), and E. (Tetramelus) mediorufus sp. nov., holotype (29) and paratype (30)." figureDoi="http://doi.org/10.5281/zenodo.13849794" httpUri="https://zenodo.org/record/13849794/files/figure.png" pageId="20" pageNumber="252">2930</figureCitation>
,
<figureCitation id="13292A390B576366FEC926A6FE24FB36" box="[342,414,1159,1185]" captionStart="FIGURES 8996" captionStartId="21.[151,264,1949,1973]" captionTargetBox="[158,1429,820,1919]" captionTargetId="figure-258@21.[151,1436,813,1926]" captionTargetPageId="21" captionText="FIGURES 8996. Euconnus (Tetramelus) mediorufus sp. nov., holotype (8992) and paratype (9396). Aedeagus in ventral (89, 91, 93, 95) and lateral (90, 92, 94, 96) views." figureDoi="http://doi.org/10.5281/zenodo.13849812" httpUri="https://zenodo.org/record/13849812/files/figure.png" pageId="20" pageNumber="252">8996</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30865370B576366FF0826EFFE82FA0A" pageId="20" pageNumber="252" type="materials_examined">
<paragraph id="8BAD36BC0B576366FF0826EFFE82FA0A" blockId="20.[151,1437,1230,2013]" pageId="20" pageNumber="252">
<emphasis id="B966EAAE0B576366FF0826EFFEE4FB7F" bold="true" box="[151,350,1230,1256]" pageId="20" pageNumber="252">Material studied</emphasis>
.
<materialsCitation id="3B7A3CE10B576366FEF326EEFB4CFAC2" collectingDate="2005-06-05" collectionCode="MNHB" collectorName="M. Schulke" country="China" county="Zhongdian Co." elevation="3895" latitude="27.618334" location="Xue Shan" longLatPrecision="123" longitude="99.64167" municipality="Diqing" pageId="20" pageNumber="252" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B966EAAE0B576366FEF326EEFD5EFB7F" bold="true" box="[364,740,1230,1260]" pageId="20" pageNumber="252">
<typeStatus id="54A9881E0B576366FEF326EEFE67FB7E" box="[364,477,1231,1257]" pageId="20" pageNumber="252" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D14FD350B576366FE7526F1FE45FB7B" box="[490,511,1232,1260]" count="1" pageId="20" pageNumber="252" type="male"></specimenCount>
(
<collectingCountry id="F305762C0B576366FD9026EFFDC8FB7F" box="[527,626,1230,1256]" name="China" pageId="20" pageNumber="252">CHINA</collectingCountry>
:
<collectingRegion id="49D6F85E0B576366FDE226EFFD67FB7F" box="[637,733,1230,1256]" country="China" name="Yunnan" pageId="20" pageNumber="252">Yunnan</collectingRegion>
)
</emphasis>
: “CHINA: N-Yunnan [C2005-05] /
<collectingMunicipality id="6BC9ACC60B576366FBE326EEFB70FB7E" box="[1148,1226,1231,1257]" pageId="20" pageNumber="252">Diqing</collectingMunicipality>
<collectingRegion id="49D6F85E0B576366FB4E26EEFAB5FB7E" box="[1233,1295,1231,1257]" country="China" name="Xizang" pageId="20" pageNumber="252">Tibet</collectingRegion>
. Aut. Pref., /
<collectingCounty id="62CC4E300B576366FF0826D2FEFBFA9A" box="[151,321,1267,1293]" pageId="20" pageNumber="252">Zhongdian Co.</collectingCounty>
,
<location id="8ECD60670B576366FED326D2FE00FA9A" LSID="urn:lsid:plazi:treatment:03BB87AA0B576364FF082642FB1DFE46:8ECD60670B576366FED326D2FE00FA9A" box="[332,442,1267,1293]" country="China" county="Zhongdian Co." latitude="27.618334" longLatPrecision="123" longitude="99.64167" municipality="Diqing" name="Xue Shan" pageId="20" pageNumber="252" stateProvince="Yunnan">Xue Shan</location>
near / lake
<locationDeviation id="9A1F2F9E0B576366FDA426D2FCA0FA9A" box="[571,794,1267,1293]" location="Zhongdian" pageId="20" pageNumber="252">
<quantity id="4CEA9B590B576366FDA426D2FD3BFA9A" box="[571,641,1267,1293]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="252" unit="km" value="23.0">23 km</quantity>
S Zhongdian
</locationDeviation>
,
<quantity id="4CEA9B590B576366FCBA26D3FCC7FA9A" box="[805,893,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.895" pageId="20" pageNumber="252" unit="m" value="3895.0">
<elevation id="003FD18F0B576366FCBA26D3FCC7FA9A" box="[805,893,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.895" pageId="20" pageNumber="252" unit="m" value="3895.0">3895 m</elevation>
</quantity>
”, “
<geoCoordinate id="EE26507B0B576366FC0126D2FBA7FA9A" box="[926,1053,1267,1293]" degrees="27" direction="north" minutes="37.1" orientation="latitude" pageId="20" pageNumber="252" precision="92" value="27.618334">27°37.1 N</geoCoordinate>
,
<geoCoordinate id="EE26507B0B576366FBB726D3FB20FA9A" box="[1064,1178,1266,1293]" degrees="99" direction="east" minutes="38.5" orientation="longitude" pageId="20" pageNumber="252" precision="92" value="99.64167">99°38.5 E</geoCoordinate>
, devast. / mixed forest, meadows, lake / border, leaf litter, dead wood, / sifted,
<date id="FFAC107C0B576366FD6B2737FCD3FAA6" box="[756,873,1302,1329]" pageId="20" pageNumber="252" value="2005-06-05">
<collectingDate id="EFE8E9940B576366FD6B2737FCD3FAA6" box="[756,873,1302,1329]" pageId="20" pageNumber="252" value="2005-06-05">5.VI.2005</collectingDate>
</date>
, / leg.
<collectorName id="26E7536A0B576366FC2F2736FB88FAA6" box="[944,1074,1303,1329]" pageId="20" pageNumber="252">M. Schülke</collectorName>
” [white, printed], “
<taxonomicName id="4C124D3F0B576366FA972736FA26FAA6" box="[1288,1436,1303,1329]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B576366FA972736FA26FAA6" box="[1288,1436,1303,1329]" italics="true" pageId="20" pageNumber="252">EUCONNUS</emphasis>
</taxonomicName>
/ (
<taxonomicName id="4C124D3F0B576366FF2F271AFEE7FAC2" box="[176,349,1339,1365]" pageId="20" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B576366FF2F271AFEE7FAC2" box="[176,349,1339,1365]" italics="true" pageId="20" pageNumber="252">TETRAMELUS</emphasis>
</taxonomicName>
) /
<taxonomicName id="4C124D3F0B576366FEE1271AFE46FAC2" authorityName="Jałoszyński" authorityYear="2024" box="[382,508,1339,1365]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="252" phylum="Arthropoda" rank="species" species="mediorufus">
<emphasis id="B966EAAE0B576366FEE1271AFE46FAC2" box="[382,508,1339,1365]" italics="true" pageId="20" pageNumber="252">mediorufus</emphasis>
</taxonomicName>
m. / P.
<bibRefCitation id="EF834B4D0B576366FDCB271AFCA7FAC2" author="Jaloszynski, P." box="[596,797,1339,1365]" pageId="20" pageNumber="252" pagination="549 - 570" refId="ref15479" refString="Jaloszynski, P. (2024) Palposcydmus gen. n. from India, with a preliminary key to world genera of Stenichnini (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5428 (4), 549 - 570. https: // doi. org / 10.11646 / zootaxa. 5428.4.5" type="journal article" year="2024">Jałoszyński, 2024</bibRefCitation>
/
<typeStatus id="54A9881E0B576366FCAB271AFC67FAC2" box="[820,989,1339,1365]" pageId="20" pageNumber="252" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="ED03AE790B576366FB0B271AFB54FAC2" box="[1172,1262,1339,1365]" pageId="20" pageNumber="252">MNHB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7A3CE10B576366FA9D271AFE89FA0A" collectingDate="2012-06-29" collectionCode="V" country="China" elevation="3362" location="Yunnan" pageId="20" pageNumber="252" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B966EAAE0B576366FA9D271AFA2FFACF" bold="true" box="[1282,1429,1339,1368]" pageId="20" pageNumber="252">
<typeStatus id="54A9881E0B576366FA9D271AFAC9FAC2" box="[1282,1395,1339,1365]" pageId="20" pageNumber="252" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D14FD350B576366FA1F271DFA2FFACF" box="[1408,1429,1340,1368]" count="1" pageId="20" pageNumber="252" type="male"></specimenCount>
</emphasis>
, “CHINA,
<collectingRegion id="49D6F85E0B576366FE96277EFED8FAEE" box="[265,354,1375,1401]" country="China" name="Yunnan" pageId="20" pageNumber="252">Yunnan</collectingRegion>
, / Haba Shan, N27°2157 / E100°0655, /
<date id="FFAC107C0B576366FCC9277EFC6AFAEE" box="[854,976,1375,1401]" pageId="20" pageNumber="252" value="2012-06-29">
<collectingDate id="EFE8E9940B576366FCC9277EFC6AFAEE" box="[854,976,1375,1401]" pageId="20" pageNumber="252" value="2012-06-29">29.vi.2012</collectingDate>
</date>
,
<quantity id="4CEA9B590B576366FC44277EFB96FAEE" box="[987,1068,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="3.3619999999999997" pageId="20" pageNumber="252" unit="m" value="3362.0">
<elevation id="003FD18F0B576366FC44277EFB96FAEE" box="[987,1068,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="3.3619999999999997" pageId="20" pageNumber="252" unit="m" value="3362.0">3362m</elevation>
</quantity>
, / sift36
<collectionCode id="ED03AE790B576366FB15277EFB24FAEE" box="[1162,1182,1375,1401]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="252" type="Museum">V</collectionCode>
. Grebennikov” [white, printed] (cPJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30865370B576366FF582787FE1EF9E2" pageId="20" pageNumber="252" type="diagnosis">
<paragraph id="8BAD36BC0B576366FF582787FE1EF9E2" blockId="20.[151,1437,1230,2013]" pageId="20" pageNumber="252">
<emphasis id="B966EAAE0B576366FF582787FEF8FA57" bold="true" box="[199,322,1446,1472]" pageId="20" pageNumber="252">Diagnosis.</emphasis>
Body rufous brown; male metatrochanter distally produced into long subtriangular spine, ventral margin of trochanter evenly concave in distal half; aedeagus in ventral view slightly less than twice as long as wide, its ventral plate much wider than long, with lateral margins strongly sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging to narrowly rounded apex, endophallus in ventral view with one short elongate sclerite surrounded by endophallic folds covered densely with cuticular denticles.
</paragraph>
</subSubSection>
<subSubSection id="C30865370B576367FF58245FFE0BFC82" lastPageId="21" lastPageNumber="253" pageId="20" pageNumber="252" type="description">
<paragraph id="8BAD36BC0B576366FF58245FFB71F92A" blockId="20.[151,1437,1230,2013]" pageId="20" pageNumber="252">
<emphasis id="B966EAAE0B576366FF58245FFEE3F90F" bold="true" box="[199,345,1662,1688]" pageId="20" pageNumber="252">Description.</emphasis>
Body of male (
<figureCitation id="13292A390B576366FD88245EFDD7F90E" box="[535,621,1663,1689]" captionStart="FIGURES 1320" captionStartId="7.[151,264,1725,1749]" captionTargetBox="[157,1430,187,1695]" captionTargetId="figure-19@7.[151,1436,181,1701]" captionTargetPageId="7" captionText="FIGURES 1320. Habitus in dorsal view, males. Euconnus (Tetramelus) punctator sp. nov. (13), E. (Tetramelus) aurilobatus sp. nov. (14), E. (Tetramelus) magnirufus sp. nov. (15), E. (Tetramelus) tetraspinosus sp. nov. (16), E. (Tetramelus) gansuminor sp. nov. (17), E. (Tetramelus) microrufus sp. nov. (18), E. (Tetramelus) sinespinosus sp. nov. (19), and E. (Tetramelus) mediorufus sp. nov. (20)." figureDoi="http://doi.org/10.5281/zenodo.13849790" httpUri="https://zenodo.org/record/13849790/files/figure.png" pageId="20" pageNumber="252">Fig. 20</figureCitation>
) elongate and slender, moderately convex, with long appendages, BL
<quantity id="4CEA9B590B576366FF082482FE87F92A" box="[151,317,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.0549999999999997" metricValueMax="2.13" metricValueMin="1.98" pageId="20" pageNumber="252" unit="mm" value="2.0549999999999997" valueMax="2.13" valueMin="1.98">1.982.13 mm</quantity>
; cuticle glossy, pigmentation rufous brown, vestiture of setae lighter than cuticle.
</paragraph>
<paragraph id="8BAD36BC0B576366FF5824E6FE8CF84A" blockId="20.[151,1437,1230,2013]" pageId="20" pageNumber="252">
Head in dorsal view round, broadest across eyes, HL
<quantity id="4CEA9B590B576366FCBB24E7FC70F976" box="[804,970,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="20" pageNumber="252" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.330.35 mm</quantity>
, HW
<quantity id="4CEA9B590B576366FB9324E7FB09F976" box="[1036,1203,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="20" pageNumber="252" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.330.35 mm</quantity>
; in lateral view each temple about 3.5 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, indistinctly shorter than half BL, AnL
<quantity id="4CEA9B590B576366FDD1255AFD4CF802" box="[590,758,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" metricValueMax="9.0" metricValueMin="8.8" pageId="20" pageNumber="252" unit="mm" value="0.89" valueMax="0.9" valueMin="0.88">0.880.90 mm</quantity>
; antennomeres 15 elongate (35 indistinctly so), 68 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 910 combined, about 1.5 times as long as broad.
</paragraph>
<paragraph id="8BAD36BC0B566367FF5822B6FAC8FE8A" blockId="21.[151,1437,150,789]" pageId="21" pageNumber="253">
Pronotum broadest anterior to middle, PL
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, PW
<quantity id="4CEA9B590B566367FC1322B7FB89FF26" box="[908,1075,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="21" pageNumber="253" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.430.45 mm</quantity>
; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small but distinct pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.
</paragraph>
<paragraph id="8BAD36BC0B566367FF582306FC1BFE1E" blockId="21.[151,1437,150,789]" pageId="21" pageNumber="253">
Elytra oval, broadest slightly in front of middle, EL
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, EW
<quantity id="4CEA9B590B566367FC4E2306FB8FFED6" box="[977,1077,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="253" unit="mm" value="0.8">0.80 mm</quantity>
, EI 1.471.56; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.
</paragraph>
<paragraph id="8BAD36BC0B566367FF5823B2FD76FE46" blockId="21.[151,1437,150,789]" pageId="21" pageNumber="253">
Legs long and slender, metatrochanter (
<figureCitation id="13292A390B566367FD1023B2FCAEFE3A" box="[655,788,403,429]" captionStart="FIGURES 2130" captionStartId="8.[151,264,1444,1468]" captionTargetBox="[156,1435,316,1415]" captionTargetId="figure-72@8.[151,1435,309,1421]" captionTargetPageId="8" captionText="FIGURES 2130. Male protrochanter (24) and metatrochanter (2123, 2530). Euconnus (Tetramelus) punctator sp. nov. (21), E. (Tetramelus) aurilobatus sp. nov. (22), E. (Tetramelus) magnirufus sp. nov. (23), E. (Tetramelus) tetraspinosus sp. nov. (24, 25), E. (Tetramelus) gansuminor sp. nov. (26), E. (Tetramelus) microrufus sp. nov. (27), E. (Tetramelus) sinespinosus sp. nov. (28), and E. (Tetramelus) mediorufus sp. nov., holotype (29) and paratype (30)." figureDoi="http://doi.org/10.5281/zenodo.13849794" httpUri="https://zenodo.org/record/13849794/files/figure.png" pageId="21" pageNumber="253">Figs 29, 30</figureCitation>
) distally produced into long subtriangular spine, ventral margin of trochanter evenly concave in distal half.
</paragraph>
<paragraph id="8BAD36BC0B566367FF5823FAFF5FFD8E" blockId="21.[151,1437,150,789]" pageId="21" pageNumber="253">Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</paragraph>
<paragraph id="8BAD36BC0B566367FF582002FDDCFD66" blockId="21.[151,1437,150,789]" pageId="21" pageNumber="253">
Aedeagus (
<figureCitation id="13292A390B566367FED92002FE72FDAA" box="[326,456,547,573]" captionStart="FIGURES 8996" captionStartId="21.[151,264,1949,1973]" captionTargetBox="[158,1429,820,1919]" captionTargetId="figure-258@21.[151,1436,813,1926]" captionTargetPageId="21" captionText="FIGURES 8996. Euconnus (Tetramelus) mediorufus sp. nov., holotype (8992) and paratype (9396). Aedeagus in ventral (89, 91, 93, 95) and lateral (90, 92, 94, 96) views." figureDoi="http://doi.org/10.5281/zenodo.13849812" httpUri="https://zenodo.org/record/13849812/files/figure.png" pageId="21" pageNumber="253">Figs 8996</figureCitation>
) stout with slender apical region, AeL
<quantity id="4CEA9B590B566367FC1C2003FC51FDAA" box="[899,1003,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="21" pageNumber="253" unit="mm" value="0.55">0.55 mm</quantity>
; median lobe in ventral view broadest distally to middle or near middle, distally rapidly narrowing, ventral plate much wider than long, with lateral margins strongly sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging to narrowly rounded apex, endophallus in ventral view with one short elongate sclerite surrounded by endophallic folds covered densely with cuticular denticles. Dorsal wall with short setae; parameres slender, each with 3 long subapical setae.
</paragraph>
<paragraph id="8BAD36BC0B566367FF5820DAFE0BFC82" blockId="21.[151,1437,150,789]" box="[199,433,763,789]" pageId="21" pageNumber="253">
<emphasis id="B966EAAE0B566367FF5820DAFEA4FC82" bold="true" box="[199,286,763,789]" pageId="21" pageNumber="253">Female</emphasis>
. Not known.
</paragraph>
</subSubSection>
<caption id="DF6D66340B566367FF0825BCFD35F84E" ID-DOI="http://doi.org/10.5281/zenodo.13849812" ID-Zenodo-Dep="13849812" httpUri="https://zenodo.org/record/13849812/files/figure.png" pageId="21" pageNumber="253" startId="21.[151,264,1949,1973]" targetBox="[158,1429,820,1919]" targetPageId="21" targetType="figure">
<paragraph id="8BAD36BC0B566367FF0825BCFD35F84E" blockId="21.[151,1436,1949,2010]" pageId="21" pageNumber="253">
<emphasis id="B966EAAE0B566367FF0825BCFEECF822" bold="true" box="[151,342,1949,1973]" pageId="21" pageNumber="253">FIGURES 8996.</emphasis>
<taxonomicName id="4C124D3F0B566367FEC225BFFD7EF822" authorityName="Jałoszyński" authorityYear="2024" box="[349,708,1949,1974]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="253" phylum="Arthropoda" rank="species" species="mediorufus" status="sp. nov." subGenus="Tetramelus">
<emphasis id="B966EAAE0B566367FEC225BFFE7BF822" box="[349,449,1950,1973]" italics="true" pageId="21" pageNumber="253">Euconnus</emphasis>
(
<emphasis id="B966EAAE0B566367FE4D25BCFDF8F822" box="[466,578,1949,1973]" italics="true" pageId="21" pageNumber="253">Tetramelus</emphasis>
)
<emphasis id="B966EAAE0B566367FDCD25BCFD7EF822" box="[594,708,1949,1973]" italics="true" pageId="21" pageNumber="253">mediorufus</emphasis>
</taxonomicName>
<emphasis id="B966EAAE0B566367FD5325BFFCA4F821" bold="true" box="[716,798,1950,1974]" pageId="21" pageNumber="253">
<taxonomicNameLabel id="A25557D50B566367FD5325BFFCA4F821" box="[716,798,1950,1974]" pageId="21" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (
<emphasis id="B966EAAE0B566367FC0925BCFC6FF822" bold="true" box="[918,981,1949,1973]" pageId="21" pageNumber="253">8992</emphasis>
) and paratype (
<emphasis id="B966EAAE0B566367FBE525BCFB03F822" bold="true" box="[1146,1209,1949,1973]" pageId="21" pageNumber="253">9396</emphasis>
). Aedeagus in ventral (
<emphasis id="B966EAAE0B566367FF3F25E0FE91F84E" bold="true" box="[160,299,1985,2009]" pageId="21" pageNumber="253">89, 91, 93, 95</emphasis>
) and lateral (
<emphasis id="B966EAAE0B566367FE2C25E0FD84F84E" bold="true" box="[435,574,1985,2009]" pageId="21" pageNumber="253">90, 92, 94, 96</emphasis>
) views.
</paragraph>
</caption>
<subSubSection id="C30865370B556364FF5822B7FBB7FF26" box="[199,1037,150,177]" pageId="22" pageNumber="254" type="distribution">
<paragraph id="8BAD36BC0B556364FF5822B7FBB7FF26" blockId="22.[151,1437,150,465]" box="[199,1037,150,177]" pageId="22" pageNumber="254">
<emphasis id="B966EAAE0B556364FF5822B7FEE3FF27" bold="true" box="[199,345,150,176]" pageId="22" pageNumber="254">Distribution</emphasis>
. WS
<collectingCountry id="F305762C0B556364FE0722B6FE65FF26" box="[408,479,151,177]" name="China" pageId="22" pageNumber="254">China</collectingCountry>
:
<collectingRegion id="49D6F85E0B556364FE7622B6FDF8FF26" box="[489,578,151,177]" country="China" name="Yunnan" pageId="22" pageNumber="254">Yunnan</collectingRegion>
, Diqing Tibetan Autonomous Prefecture.
</paragraph>
</subSubSection>
<subSubSection id="C30865370B556364FF58229AFA2EFF42" box="[199,1428,187,213]" pageId="22" pageNumber="254" type="etymology">
<paragraph id="8BAD36BC0B556364FF58229AFA2EFF42" blockId="22.[151,1437,150,465]" box="[199,1428,187,213]" pageId="22" pageNumber="254">
<emphasis id="B966EAAE0B556364FF58229AFEF4FF42" bold="true" box="[199,334,187,213]" pageId="22" pageNumber="254">Etymology.</emphasis>
The adjective
<taxonomicName id="4C124D3F0B556364FE6C229AFDCBFF42" authorityName="Jałoszyński" authorityYear="2024" box="[499,625,187,213]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="mediorufus">
<emphasis id="B966EAAE0B556364FE6C229AFDCBFF42" box="[499,625,187,213]" italics="true" pageId="22" pageNumber="254">mediorufus</emphasis>
</taxonomicName>
refers to the medium body size and to the reddish-brown pigmentation.
</paragraph>
</subSubSection>
<subSubSection id="C30865370B556364FF5822FEFB1DFE46" pageId="22" pageNumber="254" type="discussion">
<paragraph id="8BAD36BC0B556364FF5822FEFB98FE1E" blockId="22.[151,1437,150,465]" pageId="22" pageNumber="254">
<emphasis id="B966EAAE0B556364FF5822FEFE81FF6E" bold="true" box="[199,315,223,249]" pageId="22" pageNumber="254">Remarks.</emphasis>
Males of
<taxonomicName id="4C124D3F0B556364FE3522FEFD27FF6E" authorityName="Jałoszyński" authorityYear="2024" box="[426,669,223,249]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="mediorufus">
<emphasis id="B966EAAE0B556364FE3522FEFD27FF6E" box="[426,669,223,249]" italics="true" pageId="22" pageNumber="254">Euconnus mediorufus</emphasis>
</taxonomicName>
have the metatrochanters modified similarly to those in
<taxonomicName id="4C124D3F0B556364FA8E22FEFA26FF6F" authorityName="Jałoszyński" authorityYear="2024" box="[1297,1436,223,248]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="punctator">
<emphasis id="B966EAAE0B556364FA8E22FEFA26FF6F" box="[1297,1436,223,248]" italics="true" pageId="22" pageNumber="254">E. punctator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D3F0B556364FF572322FED3FE8A" authorityName="Jałoszyński" authorityYear="2024" box="[200,361,259,285]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="magnirufus">
<emphasis id="B966EAAE0B556364FF572322FED3FE8A" box="[200,361,259,285]" italics="true" pageId="22" pageNumber="254">E. magnirufus</emphasis>
</taxonomicName>
, with a relatively long subtriangular distal spine and the ventral margin arcuate, not bent. The three species share also a similar endophallic structure, with membranous folds densely covered with numerous cuticular denticles. The dorsal plate in
<taxonomicName id="4C124D3F0B556364FDDB236AFD5AFEF2" authorityName="Jałoszyński" authorityYear="2024" box="[580,736,331,357]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="mediorufus">
<emphasis id="B966EAAE0B556364FDDB236AFD5AFEF2" box="[580,736,331,357]" italics="true" pageId="22" pageNumber="254">E. mediorufus</emphasis>
</taxonomicName>
is conspicuously elongate and narrow, whereas in
<taxonomicName id="4C124D3F0B556364FA8E236AFA26FEF3" authorityName="Jałoszyński" authorityYear="2024" box="[1297,1436,331,356]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="punctator">
<emphasis id="B966EAAE0B556364FA8E236AFA26FEF3" box="[1297,1436,331,356]" italics="true" pageId="22" pageNumber="254">E. punctator</emphasis>
</taxonomicName>
it is elongate but broad, and in
<taxonomicName id="4C124D3F0B556364FE6D234EFD2BFE1E" authorityName="Jałoszyński" authorityYear="2024" box="[498,657,367,393]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="254" phylum="Arthropoda" rank="species" species="magnirufus">
<emphasis id="B966EAAE0B556364FE6D234EFD2BFE1E" box="[498,657,367,393]" italics="true" pageId="22" pageNumber="254">E. magnirufus</emphasis>
</taxonomicName>
the plate is nearly as long as broad.
</paragraph>
<paragraph id="8BAD36BC0B556364FF5823B2FB1DFE46" blockId="22.[151,1437,150,465]" pageId="22" pageNumber="254">
The two studied males differ slightly in metatrochanters (
<figureCitation id="13292A390B556364FCFD23B2FBAEFE3A" box="[866,1044,403,429]" captionStart="FIGURES 2130" captionStartId="8.[151,264,1444,1468]" captionTargetBox="[156,1435,316,1415]" captionTargetId="figure-72@8.[151,1435,309,1421]" captionTargetPageId="8" captionText="FIGURES 2130. Male protrochanter (24) and metatrochanter (2123, 2530). Euconnus (Tetramelus) punctator sp. nov. (21), E. (Tetramelus) aurilobatus sp. nov. (22), E. (Tetramelus) magnirufus sp. nov. (23), E. (Tetramelus) tetraspinosus sp. nov. (24, 25), E. (Tetramelus) gansuminor sp. nov. (26), E. (Tetramelus) microrufus sp. nov. (27), E. (Tetramelus) sinespinosus sp. nov. (28), and E. (Tetramelus) mediorufus sp. nov., holotype (29) and paratype (30)." figureDoi="http://doi.org/10.5281/zenodo.13849794" httpUri="https://zenodo.org/record/13849794/files/figure.png" pageId="22" pageNumber="254">Figs 29 and 30</figureCitation>
) and in aedeagi (
<figureCitation id="13292A390B556364FB7E23B2FADFFE3A" box="[1249,1381,403,429]" captionStart="FIGURES 8996" captionStartId="21.[151,264,1949,1973]" captionTargetBox="[158,1429,820,1919]" captionTargetId="figure-258@21.[151,1436,813,1926]" captionTargetPageId="21" captionText="FIGURES 8996. Euconnus (Tetramelus) mediorufus sp. nov., holotype (8992) and paratype (9396). Aedeagus in ventral (89, 91, 93, 95) and lateral (90, 92, 94, 96) views." figureDoi="http://doi.org/10.5281/zenodo.13849812" httpUri="https://zenodo.org/record/13849812/files/figure.png" pageId="22" pageNumber="254">Figs 8992</figureCitation>
, and 9396). However, these differences are too small to justify establishing two separate species.
</paragraph>
</subSubSection>
</treatment>
</document>