318 lines
36 KiB
XML
318 lines
36 KiB
XML
<document id="18B65EA61D331B6D9D03DBBAB06C879F" ID-CLB-Dataset="6433" ID-DOI="10.11646/zootaxa.4969.1.12" ID-GBIF-Dataset="d5160a0e-64cf-4431-bedb-89e429fb6437" ID-ISSN="1175-5326" ID-Zenodo-Dep="4745846" ID-ZooBank="A7F42721-D9FE-4CAA-A746-F89453069B84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620640885300" checkinUser="plazi" docAuthor="Mai, Thai Van & Nguyen, Lien Thi Phuong" docDate="2021" docId="03BB1E28FFB8A10F87ADFD37CCF7FE6C" docLanguage="en" docName="zootaxa.4969.1.12.pdf" docOrigin="Zootaxa 4969 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeleumenes van der Vecht 1963" docType="key" docVersion="7" lastPageNumber="196" masterDocId="FF826650FFBCA10A873AFFB8C96BFFD8" masterDocTitle="New findings on the taxonomy of the genus Coeleumenes van der Vecht (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species" masterLastPageNumber="196" masterPageNumber="191" pageNumber="195" updateTime="1698938491325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="68BBD52CD1C958CD9469FABE03A20DD8">
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<mods:title id="84A6FA61BFF2F2CFEAE7897E714778B0">New findings on the taxonomy of the genus Coeleumenes van der Vecht (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species</mods:title>
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<mods:name id="6146A907EDCF068D4211DEBC764BEEB8" type="personal">
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<mods:namePart id="608563BC4A15958306F649E6621DFDF0">Mai, Thai Van</mods:namePart>
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<mods:nameIdentifier id="5240E6B1C790349D6458E1245FFB3D8B" type="ORCID">0000-0002-3609-4286</mods:nameIdentifier>
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<mods:affiliation id="F0B1FB629EC751D970587B59E3D97435">Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam. & Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam. & https: // orcid. org / 0000 - 0002 - 3609 - 4286</mods:affiliation>
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<mods:roleTerm id="A4A214620513E97B74923AF8628D1267">Author</mods:roleTerm>
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<mods:namePart id="68EFA52A6069AC5F3BCBEF2B748E3402">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:affiliation id="0C7FE245D84B7448805217C5A8E2D7BF">Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam. & Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="0E882D44334C8103844DB5360C0C887E">Zootaxa</mods:title>
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<mods:part id="188D621CA91BDFCF0B0F3FDA425EEC55">
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<mods:date id="57D6EF6F3126687FD006068EDFCBD79E">2021</mods:date>
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<mods:number id="D08ADB5EE8A54B4386384262DD5606C3">2021-05-10</mods:number>
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<mods:number id="D179AA20FD0CB377F7709F96E978A1DE">4969</mods:number>
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<mods:identifier id="162E2B524C1BEC90ECAB07627021E1BE" type="DOI">10.11646/zootaxa.4969.1.12</mods:identifier>
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<treatment id="03BB1E28FFB8A10F87ADFD37CCF7FE6C" ID-DOI="http://doi.org/10.5281/zenodo.4813393" ID-GBIF-Taxon="182398558" ID-Zenodo-Dep="4813393" LSID="urn:lsid:plazi:treatment:03BB1E28FFB8A10F87ADFD37CCF7FE6C" httpUri="http://treatment.plazi.org/id/03BB1E28FFB8A10F87ADFD37CCF7FE6C" lastPageId="5" lastPageNumber="196" pageId="4" pageNumber="195">
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<subSubSection id="C308FCB5FFB8A10E87ADFD37CD20FD70" box="[151,1099,654,681]" pageId="4" pageNumber="195" type="nomenclature">
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<paragraph id="8BADAF3EFFB8A10E87ADFD37CD20FD70" blockId="4.[151,1099,654,681]" box="[151,1099,654,681]" pageId="4" pageNumber="195">
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<heading id="D0E51852FFB8A10E87ADFD37CD20FD70" bold="true" box="[151,1099,654,681]" fontSize="11" level="1" pageId="4" pageNumber="195" reason="1">
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<emphasis id="B966732CFFB8A10E87ADFD37CD20FD70" bold="true" box="[151,1099,654,681]" pageId="4" pageNumber="195">
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Key to all species of the genus
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<taxonomicName id="4C12D4BDFFB8A10E8528FD36CBD8FD71" ID-CoL="3RTM" ID-ENA="2908072" authorityName="van der Vecht" authorityYear="1963" box="[530,691,654,681]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="genus">
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<emphasis id="B966732CFFB8A10E8528FD36CBD8FD71" bold="true" box="[530,691,654,681]" italics="true" pageId="4" pageNumber="195">Coeleumenes</emphasis>
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</taxonomicName>
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occurring in the Oriental region
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C308FCB5FFB8A10E87ADFD6FCD6FFC7D" pageId="4" pageNumber="195" type="description">
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<paragraph id="8BADAF3EFFB8A10E87ADFD6FCDF4FCE1" blockId="4.[151,1436,727,933]" pageId="4" pageNumber="195">
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The characters are applicable to both sexes unless the sex is specified. The characters are taken from descriptions of the species following
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<bibRefCitation id="EF83D2CFFFB8A10E86B7FD43CB4CFCCD" author="Sonan, J." box="[397,551,763,789]" pageId="4" pageNumber="195" pagination="131 - 140" refId="ref4912" refString="Sonan, J. (1939) Description of eight new species of Eumenidae in Formosa (Hymenoptera). Transactions of the Natural History Society of Formosa, 29, 131 - 140." type="journal article" year="1939">Sonan (1939)</bibRefCitation>
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,
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<bibRefCitation id="EF83D2CFFFB8A10E8509FD43CA43FCCD" author="Giordani Soika, A." box="[563,808,762,789]" pageId="4" pageNumber="195" pagination="1 - 38" refId="ref4648" refString="Giordani Soika, A. (1935) Richerche sistematiche sugli Eumenes e Pareumenes dell' - Archipelago Malese e della Nova Guinea. Annali di Museo Civico di Storia Naturale di Genova, 57, 1 - 38." type="journal article" year="1935">Giordani Soika (1935</bibRefCitation>
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,
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<bibRefCitation id="EF83D2CFFFB8A10E840EFD43CA04FCCD" author="Giordani Soika, A." box="[820,879,763,789]" pageId="4" pageNumber="195" pagination="130 - 279" refId="ref4687" refString="Giordani Soika, A. (1941) Studi sui Vespidi Solitari. Bollettino della Societa Veneziana Storia Naturale, 2, 130 - 279." type="journal article" year="1941">1941</bibRefCitation>
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), and van der
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<bibRefCitation id="EF83D2CFFFB8A10E8334FD43CDF4FCCE" author="Vecht, J. van der" box="[1038,1183,763,790]" pageId="4" pageNumber="195" pagination="3 - 113" refId="ref4947" refString="Vecht, J. van der (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen, Leiden, 60, 3 - 113." type="journal article" year="1963">Vecht (1963)</bibRefCitation>
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except
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<taxonomicName id="4C12D4BDFFB8A10E83CDFD43C83CFCE1" authority="(Bingham, 1897)" baseAuthorityName="Bingham" baseAuthorityYear="1897" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="burmanicus">
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<emphasis id="B966732CFFB8A10E83CDFD43CCF7FCCD" box="[1271,1436,763,789]" italics="true" pageId="4" pageNumber="195">C. burmanicus</emphasis>
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(
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<bibRefCitation id="EF83D2CFFFB8A10E87A5FCA7C825FCE1" author="Bingham, C. T." box="[159,334,799,825]" pageId="4" pageNumber="195" refId="ref4456" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738" type="book" year="1897">Bingham, 1897</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C12D4BDFFB8A10E86B7FCA7CBFDFCE1" authority="Nguyen, 2016" authorityName="Nguyen" authorityYear="2016" box="[397,662,799,825]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="flavus">
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<emphasis id="B966732CFFB8A10E86B7FCA7C89AFCE1" box="[397,497,799,825]" italics="true" pageId="4" pageNumber="195">C. flavus</emphasis>
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<bibRefCitation id="EF83D2CFFFB8A10E86CDFCA7CBFDFCE1" author="Nguyen, L. T. P." box="[503,662,799,825]" pageId="4" pageNumber="195" pagination="175 - 180" refId="ref4713" refString="Nguyen, L. T. P. (2016) Contribution to the taxonomy of the genus Coeleumenes van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4121 (2), 175 - 180. https: // doi. org / 10.11646 / zootaxa. 4121.2.7" type="journal article" year="2016">Nguyen, 2016</bibRefCitation>
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</taxonomicName>
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from
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<collectingCountry id="F305EFAEFFB8A10E85E0FCA7CA57FCE1" box="[730,828,799,825]" name="Vietnam" pageId="4" pageNumber="195">Vietnam</collectingCountry>
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, which were examined directly.
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</paragraph>
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<paragraph id="8BADAF3EFFB8A10E87FDFCFBCD6FFC7D" blockId="4.[151,1436,727,933]" pageId="4" pageNumber="195">
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We note that the description of
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<taxonomicName id="4C12D4BDFFB8A10E850CFCFBCA11FC85" authority="(Sonan, 1939)" baseAuthorityName="Sonan" baseAuthorityYear="1939" box="[566,890,835,861]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="thoracicus">
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<emphasis id="B966732CFFB8A10E850CFCFBCBA4FC85" box="[566,719,835,861]" italics="true" pageId="4" pageNumber="195">C. thoracicus</emphasis>
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(
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<bibRefCitation id="EF83D2CFFFB8A10E85DBFCFBCA19FC85" author="Sonan, J." box="[737,882,835,861]" pageId="4" pageNumber="195" pagination="131 - 140" refId="ref4912" refString="Sonan, J. (1939) Description of eight new species of Eumenidae in Formosa (Hymenoptera). Transactions of the Natural History Society of Formosa, 29, 131 - 140." type="journal article" year="1939">Sonan, 1939</bibRefCitation>
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)
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</taxonomicName>
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and its figure is very similar to
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<taxonomicName id="4C12D4BDFFB8A10E83CFFCFBC83DFC59" authority="(Bingham, 1897)" baseAuthorityName="Bingham" baseAuthorityYear="1897" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="burmanicus">
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<emphasis id="B966732CFFB8A10E83CFFCFBCCF7FC85" box="[1269,1436,835,861]" italics="true" pageId="4" pageNumber="195">C. burmanicus</emphasis>
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(
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<bibRefCitation id="EF83D2CFFFB8A10E87A5FCDFC826FC59" author="Bingham, C. T." box="[159,333,871,897]" pageId="4" pageNumber="195" refId="ref4456" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738" type="book" year="1897">Bingham, 1897</bibRefCitation>
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)
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</taxonomicName>
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(only slightly different were the punctures on the dorsal face of the propodeum), but the
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<typeStatus id="54A9119CFFB8A10E8214FCDFCC35FC59" box="[1326,1374,871,897]" pageId="4" pageNumber="195">type</typeStatus>
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of
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<taxonomicName id="4C12D4BDFFB8A10E82B9FCDFC866FC7D" baseAuthorityName="Sonan" baseAuthorityYear="1939" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="thoracicus">
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<emphasis id="B966732CFFB8A10E82B9FCDFC866FC7D" italics="true" pageId="4" pageNumber="195">C. thoracicus</emphasis>
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</taxonomicName>
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has not yet been checked, so the status of this species is left as it is.
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</paragraph>
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</subSubSection>
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<subSubSection id="C308FCB5FFB8A10F87ADFC68CCF7FE6C" lastPageId="5" lastPageNumber="196" pageId="4" pageNumber="195" type="key">
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<keyStep id="30E6B79BFFB8A10E87ADFC68CCF7FBAF" pageId="4" pageNumber="195">
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<paragraph id="8BADAF3EFFB8A10E87ADFC68CCF7FBC7" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
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<keyLead id="30E30C0BFFB8A10E87ADFC68CCF7FBC7" pageId="4" pageNumber="195">1. Head and mesosoma with coarse punctures, border between punctures raised and forming reticulation. Propodeum without apical teeth at lateral sides, without translucent acarinarium. Sternum I with strong striae. Body mostly black, with few yellow markings............................................................................................ 2</keyLead>
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</paragraph>
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<paragraph id="8BADAF3EFFB8A10E87ADFB90CCF7FBAF" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
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<keyLead id="30E30C0BFFB8A10E87ADFB90CCF7FBAF" pageId="4" pageNumber="195">- Head and mesosoma with less coarse punctures, border between punctures smooth. Propodeum with more or less developed apical teeth at lateral sides, with somewhat swollen translucent acarinarium. Sternum I with weaker striae. Body with more extensive yellow markings.............................................................................. 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E6B79BFFB8A10E87ADFB38CCF6FB08" pageId="4" pageNumber="195">
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<paragraph id="8BADAF3EFFB8A10E87ADFB38CCF6FB6B" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
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<keyLead id="30E30C0BFFB8A10E87ADFB38CCF6FB6B" pageId="4" pageNumber="195">
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2. Dorsal face of propodeum with very dense coarse punctures, border between punctures raised and forming reticulation..............................................................................
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<taxonomicName id="4C12D4BDFFB8A10E8360FB25CCF6FB6B" ID-CoL="5ZH5C" authority="(Bingham, 1897)" baseAuthorityName="Bingham" baseAuthorityYear="1897" box="[1114,1437,1180,1203]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="burmanicus">
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<emphasis id="B966732CFFB8A10E8360FB25CCF6FB6B" bold="true" box="[1114,1437,1180,1203]" pageId="4" pageNumber="195">
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<emphasis id="B966732CFFB8A10E8360FB25CD86FB6A" bold="true" box="[1114,1261,1180,1202]" italics="true" pageId="4" pageNumber="195">C. burmanicus</emphasis>
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(
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<bibRefCitation id="EF83D2CFFFB8A10E83C3FB25CCFEFB6B" author="Bingham, C. T." box="[1273,1429,1181,1203]" pageId="4" pageNumber="195" refId="ref4456" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738" type="book" year="1897">Bingham, 1897</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BADAF3EFFB8A10E87ADFB03CCF6FB08" blockId="4.[151,1437,976,2024]" box="[151,1437,1210,1232]" pageId="4" pageNumber="195">
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<keyLead id="30E30C0BFFB8A10E87ADFB03CCF6FB08" box="[151,1437,1210,1232]" pageId="4" pageNumber="195">
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- Dorsal face of propodeum with dense minute punctures................................
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<taxonomicName id="4C12D4BDFFB8A10E83BCFB02CCF6FB08" ID-CoL="5ZGTQ" authority="(Sonan, 1939)" baseAuthorityName="Sonan" baseAuthorityYear="1939" box="[1158,1437,1210,1232]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="thoracicus">
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<emphasis id="B966732CFFB8A10E83BCFB02CCF6FB08" bold="true" box="[1158,1437,1210,1232]" pageId="4" pageNumber="195">
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<emphasis id="B966732CFFB8A10E83BCFB02CC61FB08" bold="true" box="[1158,1290,1210,1232]" italics="true" pageId="4" pageNumber="195">C. thoracicus</emphasis>
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(
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<bibRefCitation id="EF83D2CFFFB8A10E822DFB02CCFEFB08" author="Sonan, J." box="[1303,1429,1210,1232]" pageId="4" pageNumber="195" pagination="131 - 140" refId="ref4912" refString="Sonan, J. (1939) Description of eight new species of Eumenidae in Formosa (Hymenoptera). Transactions of the Natural History Society of Formosa, 29, 131 - 140." type="journal article" year="1939">Sonan, 1939</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E6B79BFFB8A10E87ADFB60CCF7FAD2" pageId="4" pageNumber="195">
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<paragraph id="8BADAF3EFFB8A10E87ADFB60CCF7FB35" blockId="4.[151,1437,976,2024]" box="[151,1436,1240,1262]" pageId="4" pageNumber="195">
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<keyLead id="30E30C0BFFB8A10E87ADFB60CCF7FB35" box="[151,1436,1240,1262]" pageId="4" pageNumber="195">3. Tergum II without, or with narrow hardly raised, apical lamella...............................................….4</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADFB4DCCF7FAD2" blockId="4.[151,1437,976,2024]" box="[151,1436,1269,1291]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADFB4DCCF7FAD2" box="[151,1436,1269,1291]" pageId="4" pageNumber="195">- Tergum II with thicker apical lamella, slightly or strongly raised................................................ 7</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10E87ADFAAACCF6FA9E" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADFAAACCF7FAF0" blockId="4.[151,1437,976,2024]" box="[151,1436,1298,1320]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADFAAACCF7FAF0" box="[151,1436,1298,1320]" pageId="4" pageNumber="195">4. Tergum I without produced node laterally...................................................................5</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADFA88CCF6FA9E" blockId="4.[151,1437,976,2024]" box="[151,1437,1328,1350]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADFA88CCF6FA9E" box="[151,1437,1328,1350]" pageId="4" pageNumber="195">- Tergum I with produced node laterally......................................................................6</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10E87ADFAF5CCF6F9CB" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADFAF5CCF7FA62" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADFAF5CCF7FA62" pageId="4" pageNumber="195">
|
|
5. Clypeus shallowly emarginate anteriorly, with apical teeth blunt. Lateral surface of propodeum and metasomal petiole with fine punctures. Tergum II with apical margin lamella wider. Female clypeus with a broad yellow band at base; mesoscutum with four short yellow lines; tergum I black with a narrow yellow apical band; legs black with yellow markings.......................................................................................
|
|
<taxonomicName id="4C12D4BDFFB8A10E8371FA1DCCF7FA62" ID-CoL="5ZH5Q" authority="van der Vecht, 1963" authorityName="van der Vecht" authorityYear="1963" box="[1099,1436,1445,1466]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="timorensis">
|
|
<emphasis id="B966732CFFB8A10E8371FA1DCCF7FA62" bold="true" box="[1099,1436,1445,1466]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E8371FA1DCDA5FA62" bold="true" box="[1099,1230,1445,1466]" italics="true" pageId="4" pageNumber="195">C. timorensis</emphasis>
|
|
van der
|
|
<bibRefCitation id="EF83D2CFFFB8A10E821CFA1DCCF7FA62" author="Vecht, J. van der" box="[1318,1436,1445,1466]" pageId="4" pageNumber="195" pagination="3 - 113" refId="ref4947" refString="Vecht, J. van der (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen, Leiden, 60, 3 - 113." type="journal article" year="1963">Vecht, 1963</bibRefCitation>
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADFA7BCCF6F9CB" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADFA7BCCF6F9CB" pageId="4" pageNumber="195">
|
|
- Clypeus deeper emarginate anteriorly, with apical teeth sharper. Lateral surface of propodeum and metasomal petiole with stronger punctures. Tergum II with apical margin lamella narrower. Female clypeus with four yellow spots; mesoscutum with two yellow short lines; legs and basal half of tergum I ferruginous.................
|
|
<taxonomicName id="4C12D4BDFFB8A10E8311FA45CCF6F9CB" ID-CoL="5ZGTN" authority="(Giordani Soika, 1935)" baseAuthorityName="Giordani Soika" baseAuthorityYear="1935" box="[1067,1437,1532,1555]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="multicolor">
|
|
<emphasis id="B966732CFFB8A10E8311FA45CCF6F9CB" bold="true" box="[1067,1437,1532,1555]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E8311FA45CDC5F9CA" bold="true" box="[1067,1198,1532,1554]" italics="true" pageId="4" pageNumber="195">C. multicolor</emphasis>
|
|
(
|
|
<bibRefCitation id="EF83D2CFFFB8A10E8380FA45CCFEF9CB" author="Giordani Soika, A." box="[1210,1429,1533,1555]" pageId="4" pageNumber="195" pagination="1 - 38" refId="ref4648" refString="Giordani Soika, A. (1935) Richerche sistematiche sugli Eumenes e Pareumenes dell' - Archipelago Malese e della Nova Guinea. Annali di Museo Civico di Storia Naturale di Genova, 57, 1 - 38." type="journal article" year="1935">Giordani Soika, 1935</bibRefCitation>
|
|
)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10E87ADF9A2CCF6F91B" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF9A2CCF6F9B3" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF9A2CCF6F9B3" pageId="4" pageNumber="195">
|
|
6. Tergum I narrow at base and abrupt expended to apical margin, its width half as wide as tergum II. Mandible black with apex red, spot at base yellow; pronotum entirely black; legs black with tarsi slightly ferruginous; wings brown with purple reflection...................................................................
|
|
<taxonomicName id="4C12D4BDFFB8A10E8364F9EDCCF6F9B3" ID-CoL="5ZH5D" authority="(de Saussure, 1855)" baseAuthorityName="de Saussure" baseAuthorityYear="1855" box="[1118,1437,1620,1643]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="indianus">
|
|
<emphasis id="B966732CFFB8A10E8364F9EDCCF6F9B3" bold="true" box="[1118,1437,1620,1643]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E8364F9EDCDBAF9B2" bold="true" box="[1118,1233,1620,1642]" italics="true" pageId="4" pageNumber="195">C. indianus</emphasis>
|
|
(de Saussure, 1855)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF9CBCCF6F91B" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF9CBCCF6F91B" pageId="4" pageNumber="195">
|
|
- Tergum I gradually expanded from base to apical margin, its width less than half as wide as tergum II. Mandible entirely black; pronotum black with yellow line at anterior margin, interrupted in the middle; legs testaceous red, with black on femora; wings fusco-hyaline.............................................................
|
|
<taxonomicName id="4C12D4BDFFB8A10E8377F915CCF6F91B" authority="(Wickwar, 1908)" baseAuthorityName="Wickwar" baseAuthorityYear="1908" box="[1101,1437,1708,1731]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="rufipetiolatus">
|
|
<emphasis id="B966732CFFB8A10E8377F915CCF6F91B" bold="true" box="[1101,1437,1708,1731]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E8377F915CD85F91A" bold="true" box="[1101,1262,1708,1730]" italics="true" pageId="4" pageNumber="195">C. rufipetiolatus</emphasis>
|
|
(
|
|
<bibRefCitation id="EF83D2CFFFB8A10E83C1F915CCFEF91B" author="Wickwar, O. S." box="[1275,1429,1709,1731]" pageId="4" pageNumber="195" pagination="114 - 124" refId="ref4979" refString="Wickwar, O. S. (1908) Hymenoptera new to Ceylon, with descriptions of new species. Spolia Zeylanica, 5, 114 - 124." type="journal article" year="1908">Wickwar, 1908</bibRefCitation>
|
|
)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10E87ADF973CCF7F925" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF973CCF7F907" blockId="4.[151,1437,976,2024]" box="[151,1436,1738,1760]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF973CCF7F907" box="[151,1436,1738,1760]" pageId="4" pageNumber="195">7. Tergum II with apical margin slightly raised................................................................ 8</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF950CCF7F925" blockId="4.[151,1437,976,2024]" box="[151,1436,1768,1790]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF950CCF7F925" box="[151,1436,1768,1790]" pageId="4" pageNumber="195">- Tergum II with apical margin strongly raised............................................................... 10</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10E87ADF8BDCCF7F8AA" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF8BDCCF6F8E0" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF8BDCCF6F8E0" pageId="4" pageNumber="195">
|
|
8. Tergum II with apical margin wider. The puncturation stronger. Metasomal petiole relatively longer. Terga I-II with more or less narrow yellow apical bands..........................................................
|
|
<taxonomicName id="4C12D4BDFFB8A10E838AF89ACCF6F8E0" ID-CoL="5ZH62" authority="(Smith, 1859)" baseAuthorityName="Smith" baseAuthorityYear="1859" box="[1200,1437,1826,1848]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="vindex">
|
|
<emphasis id="B966732CFFB8A10E838AF89ACCF6F8E0" bold="true" box="[1200,1437,1826,1848]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E838AF89ACC66F8E0" bold="true" box="[1200,1293,1826,1848]" italics="true" pageId="4" pageNumber="195">C. vindex</emphasis>
|
|
(
|
|
<bibRefCitation id="EF83D2CFFFB8A10E8220F89ACCFEF8E0" author="Smith, F." box="[1306,1429,1826,1848]" pageId="4" pageNumber="195" pagination="4 - 27" refId="ref4851" refString="Smith, F. (1859) Catalogue of Hymenopterous insects collected at Celebes by Mr. A. R. Wallace. Journal of the Proceedings of the Linnean Society of London, Zoology, 3, 4 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1858. tb 02506. x" type="journal article" year="1859">Smith, 1859</bibRefCitation>
|
|
)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF8F8CCF7F8AA" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF8F8CCF7F8AA" pageId="4" pageNumber="195">- Tergum II with apical margin narrower. The puncturation much finer. Metasomal petiole relatively shorter. Terga I-III with wide yellow apical bands................................................................................... 9</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB8A10F87ADF8C2CCF7FF12" lastPageId="5" lastPageNumber="196" pageId="4" pageNumber="195">
|
|
<paragraph id="8BADAF3EFFB8A10E87ADF8C2CCF6F813" blockId="4.[151,1437,976,2024]" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10E87ADF8C2CCF6F813" pageId="4" pageNumber="195">
|
|
9. Female clypeus with apical margin deeper emarginate. Puncturation of propodeum and metasomal petiole less coarse and less dense. Propodeum with distinct translucent area at apex. Mandible black, antennal scape reddish yellow in front, legs largely red; colour of yellow markings darker...........................................
|
|
<taxonomicName id="4C12D4BDFFB8A10E8367F80DCCF6F813" ID-CoL="5ZH5P" authority="(Dalla Torre, 1889)" baseAuthorityName="Dalla Torre" baseAuthorityYear="1889" box="[1117,1437,1972,1995]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="195" phylum="Arthropoda" rank="species" species="secundus">
|
|
<emphasis id="B966732CFFB8A10E8367F80DCCF6F813" bold="true" box="[1117,1437,1972,1995]" pageId="4" pageNumber="195">
|
|
<emphasis id="B966732CFFB8A10E8367F80DCDBEF812" bold="true" box="[1117,1237,1972,1994]" italics="true" pageId="4" pageNumber="195">C. secundus</emphasis>
|
|
(
|
|
<bibRefCitation id="EF83D2CFFFB8A10E83DBF80DCCFEF813" author="Dalla Torre, K. W." box="[1249,1429,1973,1995]" pageId="4" pageNumber="195" pagination="124 - 125" refId="ref4608" refString="Dalla Torre, K. W. (1889) Hymenopterologische Notizen. Wiener Entomologische Zeitung, 8 (3), 124 - 125. https: // doi. org / 10.5962 / bhl. part. 20036" type="journal article" year="1889">Dalla Torre, 1889</bibRefCitation>
|
|
)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB8A10F87ADF86BCCF7FF12" blockId="4.[151,1437,976,2024]" lastBlockId="5.[151,1437,151,437]" lastPageId="5" lastPageNumber="196" pageId="4" pageNumber="195">
|
|
<keyLead id="30E30C0BFFB8A10F87ADF86BCCF7FF12" lastPageId="5" lastPageNumber="196" pageId="4" pageNumber="195">
|
|
- Female clypeus with apical margin shallowly emarginate. Puncturation of propodeum and metasomal petiole coarser and denser. Propodeum without distinct translucent area at apex. Mandible largely, antennal scape and legs largely bright red; colour of yellow markings distinctly paler..............................................
|
|
<taxonomicName id="4C12D4BDFFB9A10F835BFF0DCCF7FF12" ID-CoL="5ZH5N" authority="van der Vecht, 1963" authorityName="van der Vecht" authorityYear="1963" box="[1121,1436,180,202]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="196" phylum="Arthropoda" rank="species" species="ruficrus">
|
|
<emphasis id="B966732CFFB9A10F835BFF0DCCF7FF12" bold="true" box="[1121,1436,180,202]" pageId="5" pageNumber="196">
|
|
<emphasis id="B966732CFFB9A10F835BFF0DCDA6FF12" bold="true" box="[1121,1229,180,202]" italics="true" pageId="5" pageNumber="196">C. ruficrus</emphasis>
|
|
van der
|
|
<bibRefCitation id="EF83D2CFFFB9A10F821CFF0DCCF7FF12" author="Vecht, J. van der" box="[1318,1436,181,202]" pageId="5" pageNumber="196" pagination="3 - 113" refId="ref4947" refString="Vecht, J. van der (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen, Leiden, 60, 3 - 113." type="journal article" year="1963">Vecht, 1963</bibRefCitation>
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB9A10F87ADFF6ACCF7FEDD" pageId="5" pageNumber="196">
|
|
<paragraph id="8BADAF3EFFB9A10F87ADFF6ACCF6FF30" blockId="5.[151,1437,151,437]" box="[151,1437,210,232]" pageId="5" pageNumber="196">
|
|
<keyLead id="30E30C0BFFB9A10F87ADFF6ACCF6FF30" box="[151,1437,210,232]" pageId="5" pageNumber="196">
|
|
10. Tergum I without produced node laterally..........................................
|
|
<taxonomicName id="4C12D4BDFFB9A10F8357FF6ACCF6FF30" authority="(Bingham, 1897)" baseAuthorityName="Bingham" baseAuthorityYear="1897" box="[1133,1437,210,232]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="196" phylum="Arthropoda" rank="species" species="imparidus">
|
|
<emphasis id="B966732CFFB9A10F8357FF6ACCF6FF30" bold="true" box="[1133,1437,210,232]" pageId="5" pageNumber="196">
|
|
<emphasis id="B966732CFFB9A10F8357FF6ACD86FF30" bold="true" box="[1133,1261,210,232]" italics="true" pageId="5" pageNumber="196">C. imparidus</emphasis>
|
|
(
|
|
<bibRefCitation id="EF83D2CFFFB9A10F83C3FF6ACCFEFF30" author="Bingham, C. T." box="[1273,1429,210,232]" pageId="5" pageNumber="196" refId="ref4456" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738" type="book" year="1897">Bingham, 1897</bibRefCitation>
|
|
)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB9A10F87ADFF48CCF7FEDD" blockId="5.[151,1437,151,437]" box="[151,1436,239,261]" pageId="5" pageNumber="196">
|
|
<keyLead id="30E30C0BFFB9A10F87ADFF48CCF7FEDD" box="[151,1436,239,261]" pageId="5" pageNumber="196">- Tergum I with produced node laterally.................................................................... 11</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30E6B79BFFB9A10F87ADFEB5CCF7FE6C" pageId="5" pageNumber="196">
|
|
<paragraph id="8BADAF3EFFB9A10F87ADFEB5CCF7FE85" blockId="5.[151,1437,151,437]" pageId="5" pageNumber="196">
|
|
<keyLead id="30E30C0BFFB9A10F87ADFEB5CCF7FE85" pageId="5" pageNumber="196">
|
|
11. Body with find punctures. Propodeum with translucent, bluntly square expansion laterally. Tergum I with weak produced node laterally, slightly greater than 2.5 times as long as wide in dorsal view. Sternum II gradually convex in lateral view. Body black with less yellow marks...............................................................
|
|
<emphasis id="B966732CFFB9A10F838DFEFFCCF7FE85" bold="true" box="[1207,1436,327,349]" pageId="5" pageNumber="196">
|
|
<taxonomicName id="4C12D4BDFFB9A10F838DFEFFCC21FE85" authority="Mai & Nguyen, 2021" authorityName="Mai & Nguyen" authorityYear="2021" box="[1207,1354,327,349]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="196" phylum="Arthropoda" rank="species" species="chumomray" status="sp. nov.">
|
|
<emphasis id="B966732CFFB9A10F838DFEFFCC21FE85" bold="true" box="[1207,1354,327,349]" italics="true" pageId="5" pageNumber="196">C. chumomray</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A255CE57FFB9A10F826AFEF0CCF7FE85" box="[1360,1436,328,349]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BADAF3EFFB9A10F87ADFEDDCCF7FE6C" blockId="5.[151,1437,151,437]" pageId="5" pageNumber="196">
|
|
<keyLead id="30E30C0BFFB9A10F87ADFEDDCCF7FE6C" pageId="5" pageNumber="196">
|
|
- Body with well-defined punctures. Propodeum with translucent, bluntly round expansion laterally. Tergum I with strong produced node laterally, about twice as long as wide in dorsal view. Sternum II strongly convex in lateral view. Body black with extensive yellow marks..........................................................
|
|
<taxonomicName id="4C12D4BDFFB9A10F838EFE27CCF7FE6C" authority="Nguyen, 2016" authorityName="Nguyen" authorityYear="2016" box="[1204,1436,415,437]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="196" phylum="Arthropoda" rank="species" species="flavus">
|
|
<emphasis id="B966732CFFB9A10F838EFE27CCF7FE6C" bold="true" box="[1204,1436,415,437]" pageId="5" pageNumber="196">
|
|
<emphasis id="B966732CFFB9A10F838EFE27CC67FE6D" bold="true" box="[1204,1292,415,437]" italics="true" pageId="5" pageNumber="196">C. flavus</emphasis>
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<bibRefCitation id="EF83D2CFFFB9A10F8228FE18CCF7FE6C" author="Nguyen, L. T. P." box="[1298,1436,415,437]" pageId="5" pageNumber="196" pagination="175 - 180" refId="ref4713" refString="Nguyen, L. T. P. (2016) Contribution to the taxonomy of the genus Coeleumenes van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4121 (2), 175 - 180. https: // doi. org / 10.11646 / zootaxa. 4121.2.7" type="journal article" year="2016">Nguyen, 2016</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document> |