729 lines
102 KiB
XML
729 lines
102 KiB
XML
<document id="D9F4C8D98CD8335BE6B08B441A0C2A66" ID-DOI="10.1080/14772019.2020.1783380" ID-ISSN="1478-0941" ID-PMC="PMC7455076" ID-PubMed="32939187" ID-Zenodo-Dep="10932527" ID-ZooBank="B4C7A979-7972-409B-B489-A6DDD5E35FE5" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1711484709863" checkinUser="felipe" docAuthor="Marram ̀, Giuseppe, Carneval, Giorgio, Claeso, Kerin M., Naylo, Gavin J. P. & Kriwe, Jurgen" docDate="2020" docId="0385D508FFE8FFC190DA6CFEB2962379" docLanguage="en" docName="JSystPalaeontol.18.18.1519-1542.pdf" docOrigin="Journal of Systematic Palaeontology 18 (18)" docSource="http://dx.doi.org/10.1080/14772019.2020.1783380" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.3:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="3" docTitle="Eoplatyrhina bolcensis Marram̀ & Carneval & Claeso & Naylo & Kriwe 2020, comb. nov." docType="treatment" docVersion="3" lastPageNumber="1528" masterDocId="FFBCAD70FFEDFFCB90636F6AB32D253D" masterDocTitle="Revision of the Eocene ‘ Platyrhina’ species from the Bolca Lagerst atte (Italy) reveals the first panray (Batomorphii: Zanobatidae) in the fossil record" masterLastPageNumber="1542" masterPageNumber="1519" pageNumber="1523" updateTime="1712325147553" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="2CA873EBC1CFCEC5D857167AD6963872" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F0E9352609B86711E4E43C147CBD7987">
|
||
<mods:title id="943DFEAFB93ED8048154A60B9B7C0E0C">Revision of the Eocene ‘ Platyrhina’ species from the Bolca Lagerst atte (Italy) reveals the first panray (Batomorphii: Zanobatidae) in the fossil record</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="57443303BCCBD1BE38F0918EA29FDEF7" type="personal">
|
||
<mods:role id="AFB89E440DC582CB94BE2052A74B8DBB">
|
||
<mods:roleTerm id="DAA5226AE61EAC33B9CC21F115A11C60">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DC8CADF89AD751B2D6158C88EF34389D">Marram ̀, Giuseppe</mods:namePart>
|
||
<mods:affiliation id="638AAE0EC647938ACE4AC920BA40E3E3">Universita degli Studi di Torino, Dipartimento di Scienze della Terra, via Valperga Caluso 35, 10125 Torino, Italy; & University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="BDA8ADEDF32CC28900BE4D8A11D6A9C2" type="personal">
|
||
<mods:role id="5BF53F4D49EF6A3F5C2FBAB0DB6EB516">
|
||
<mods:roleTerm id="29119818D991E0817B39DB95828DDBDD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="503B8B70F62AE4AEEFB06E823F7E28AE">Carneval, Giorgio</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F022A509CEFEFC516B25970D5A3C8029" type="personal">
|
||
<mods:role id="9AF13742CE868618FC5234FF8F13BFDF">
|
||
<mods:roleTerm id="736B1D8D8EEDEF482B3B2158369E1B84">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2D02CDFD2DEC130C58327FD897158C70">Claeso, Kerin M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="333C162592FA8E5A6420E8D7D121DEBE" type="personal">
|
||
<mods:role id="C2E68887CB3DBE4C52AB2C652EA74687">
|
||
<mods:roleTerm id="CDCA314CF6BE321B4924D4E436396F1A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="17FEA20362B3952607635C98CA22A520">Naylo, Gavin J. P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1BFEC59554F878B0B9DF3EF0C6411408" type="personal">
|
||
<mods:role id="E91736BCBD07F03CCEC0AB7904180E41">
|
||
<mods:roleTerm id="5DEB049C76F5A6B549B3A5B268DCFF09">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D70AEA5C0D5FEDC7ECB8CA47A554F3DD">Kriwe, Jurgen</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A7E0FFCCEF3B4C186E35C7659BDBA2D3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7A85D280DEFA3A48DF0DD31DF4E2726C" type="host">
|
||
<mods:titleInfo id="0122A7F4F87911E25579101F46A64EA1">
|
||
<mods:title id="39F962920BA2ACAEEBEADE34591C0639">Journal of Systematic Palaeontology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E54D24B731BFD5FF253C893F7B2A82B7">
|
||
<mods:date id="B2AA560A73E57AF7D1E3A6B892BA6C5F">2020</mods:date>
|
||
<mods:detail id="195420CD10F379EAA3FDB6E38D48B322" type="pubDate">
|
||
<mods:number id="C5DED5F1720E0C38BA5C4C2F1AE20481">2020-07-13</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0526E8162491095F8CF3108229CF6DA2" type="volume">
|
||
<mods:number id="9B3D56C440E3AD24CF4D7D726D464831">18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="02FD74837DA61DF60FE2018C55F317CC" type="issue">
|
||
<mods:number id="2948EBA5CF29AC0515779A4347372D0B">18</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5544C6D6DB1CA4AF3DE49F26804BBA64" unit="page">
|
||
<mods:start id="14F90BE2B8EAA1256BEE986B24CC8686">1519</mods:start>
|
||
<mods:end id="B431282E6F3C6A8064037FF80C518EE3">1542</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="F98907D25F7FE8ACD211B9748502BC7B">
|
||
<mods:url id="A3F7316E587E14C894C6630E016A4437">http://dx.doi.org/10.1080/14772019.2020.1783380</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="6956AA01596092F754E55DF6BDE56B57">journal article</mods:classification>
|
||
<mods:identifier id="04A1B577269A6ABDE19892CEC6702FB9" type="DOI">10.1080/14772019.2020.1783380</mods:identifier>
|
||
<mods:identifier id="38F4C1B2B2B4505B44CE837569BAC310" type="ISSN">1478-0941</mods:identifier>
|
||
<mods:identifier id="8C8EEB3404DA14CF051B310312BBDF4C" type="PMC">PMC7455076</mods:identifier>
|
||
<mods:identifier id="E55493BBB7AA1D6594B2B9D5B62FB9BD" type="PubMed">32939187</mods:identifier>
|
||
<mods:identifier id="1E97747862B436FF7F58D8F97DB9C1BE" type="Zenodo-Dep">10932527</mods:identifier>
|
||
<mods:identifier id="26226EA94193B486837E9D631E9166EF" type="ZooBank">B4C7A979-7972-409B-B489-A6DDD5E35FE5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0385D508FFE8FFC190DA6CFEB2962379" ID-DOI="http://doi.org/10.5281/zenodo.10932539" ID-Zenodo-Dep="10932539" LSID="urn:lsid:plazi:treatment:0385D508FFE8FFC190DA6CFEB2962379" httpUri="http://treatment.plazi.org/id/0385D508FFE8FFC190DA6CFEB2962379" lastPageId="10" lastPageNumber="1528" pageId="5" pageNumber="1523">
|
||
<subSubSection id="C3363795FFE8FFCE90DA6CFEB1DE2691" box="[185,755,916,940]" pageId="5" pageNumber="1523" type="nomenclature">
|
||
<paragraph id="8B93641EFFE8FFCE90DA6CFEB1DE2691" blockId="5.[185,755,916,940]" box="[185,755,916,940]" pageId="5" pageNumber="1523">
|
||
<heading id="D0DBD372FFE8FFCE90DA6CFEB1DE2691" box="[185,755,916,940]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="1523" reason="8">
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90A56CFEB15D2691" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[198,624,916,940]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE8FFCE90A56CFEB2E82691" bold="true" box="[198,453,916,940]" italics="true" pageId="5" pageNumber="1523">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE91B66CFFB14A2691" author="Heckel, M. J." box="[469,615,917,940]" pageId="5" pageNumber="1523" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE8FFCE921A6CFFB1DE2691" box="[633,755,917,940]" pageId="5" pageNumber="1523" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3363795FFE8FFCE91C56CA0B1A020C4" pageId="5" pageNumber="1523" type="description">
|
||
<paragraph id="8B93641EFFE8FFCE91C56CA0B12826DC" blockId="5.[422,517,970,994]" box="[422,517,970,994]" pageId="5" pageNumber="1523">
|
||
<figureCitation id="1317789BFFE8FFCE91C56CA0B12826DC" box="[422,517,970,994]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStartId-0="6.[159,226,1315,1336]" captionStartId-1="7.[159,226,782,803]" captionStartId-2="8.[159,226,1157,1178]" captionStartId-3="9.[159,226,951,972]" captionStartId-4="9.[159,226,1661,1682]" captionStartId-5="10.[159,226,872,893]" captionTargetBox-0="[159,1466,189,1297]" captionTargetBox-1="[159,1467,188,765]" captionTargetBox-2="[159,1460,188,1140]" captionTargetBox-3="[159,1467,188,933]" captionTargetBox-4="[160,1466,1145,1643]" captionTargetBox-5="[159,1446,204,854]" captionTargetId-0="figure-269@6.[159,1467,188,1298]" captionTargetId-1="figure-528@7.[159,1467,188,765]" captionTargetId-2="figure-295@8.[159,1467,188,1140]" captionTargetId-3="figure-57@9.[159,1467,188,933]" captionTargetId-4="figure-150@9.[159,1467,1144,1644]" captionTargetId-5="figure-443@10.[159,1467,188,854]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="Figure 2. The holotype of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site in part and counterpart. A, MGP-PD 8873C; B, MGP-PD 8874C. Scale bars = 100mm." captionText-1="Figure 3. Specimens of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C; B, original historical plate of the specimen MGP-PD 26279C, illustrated by Achille de Zigno (1813–1892). Photo: courtesy of Universit̀a degli Studi di Padova; C, specimen MGGC 7449; D, MGGC 7449 under ultraviolet light. Scale bars = 100mm." captionText-2="Figure 4. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGGC 7449, close-up of the head and pectoral girdle under UV light; B, reconstruction. Abbreviations: af, anterior fontanel; ao, antorbital cartilage; cb5, fifth ceratobranchial; hp, horn-like process; hyo, hyomandibula; mc, Meckel’s cartilage; mes, mesopterygium; met, metapterygium; nc, nasal capsule; pq, palatoquadrate; pro, propterygium; ro, rostral cartilage; rp, rostral process; sca, scapulocoracoid; syn, synarcual. Scale bars = 50 mm." captionText-3="Figure 5. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C, close-up of the rostral cartilage; B, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula; C, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin. Abbreviations: af, anterior fontanel; co, coracoid bar of the scapulocoracoid; mes, mesopterygium; met, metapterygium; pro, propterygium; ro, rostral cartilage; scap, scapular process of the scapulocoracoid; ss, suprascapulae; syn, synarcual. Scale bars = 20 mm." captionText-4="Figure 6. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, close-up of the pelvic girdle and fins in MGGC 7449; B, the same area under ultraviolet light; C, detail of one of the claspers. Abbreviations: ax, axial cartilage; cl, clasper; pf, pelvic fin radials; pub, puboischiadic bar; vc, vertebral centra. Scale bars: A, B = 50mm; C = 10 mm." captionText-5="Figure 7. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, dermal denticles from the tail of MGP-PD 26279C (dorsal view); B, dermal denticles from the tail of MGP-PD 26279C (basal view). Scale bar = 400 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10932531" figureDoi-1="http://doi.org/10.5281/zenodo.10932533" figureDoi-2="http://doi.org/10.5281/zenodo.10932537" figureDoi-3="http://doi.org/10.5281/zenodo.10932541" figureDoi-4="http://doi.org/10.5281/zenodo.10932543" figureDoi-5="http://doi.org/10.5281/zenodo.10932545" httpUri-0="https://zenodo.org/record/10932531/files/figure.png" httpUri-1="https://zenodo.org/record/10932533/files/figure.png" httpUri-2="https://zenodo.org/record/10932537/files/figure.png" httpUri-3="https://zenodo.org/record/10932541/files/figure.png" httpUri-4="https://zenodo.org/record/10932543/files/figure.png" httpUri-5="https://zenodo.org/record/10932545/files/figure.png" pageId="5" pageNumber="1523">Figs 2–7</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6B6AB1AE210A" blockId="5.[159,781,1023,1787]" pageId="5" pageNumber="1523">
|
||
1833–1843
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE91456B6AB1A9212A" authority="Agassiz" authorityName="Agassiz" box="[294,644,1024,1047]" class="Elasmobranchii" genus="Narcopterus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcanus">
|
||
<emphasis id="B958B80CFFE8FFCE91456B6AB136212A" box="[294,539,1024,1047]" italics="true" pageId="5" pageNumber="1523">Narcopterus bolcanus</emphasis>
|
||
Agassiz
|
||
</taxonomicName>
|
||
: vol. 1: 44 (
|
||
<emphasis id="B958B80CFFE8FFCE90AB6B4AB24C210A" box="[200,353,1056,1079]" italics="true" pageId="5" pageNumber="1523">nomen nudum</emphasis>
|
||
; no description or figure).
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6B2AB1D7216A" blockId="5.[159,781,1023,1787]" box="[159,762,1088,1112]" pageId="5" pageNumber="1523">
|
||
1833–1843
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE91436B2AB15D216A" authority="Agassiz" authorityName="Agassiz" box="[288,624,1088,1111]" class="Elasmobranchii" genus="Narcopterus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcanus">
|
||
<emphasis id="B958B80CFFE8FFCE91436B2AB123216A" box="[288,526,1088,1111]" italics="true" pageId="5" pageNumber="1523">Narcopterus bolcanus</emphasis>
|
||
Agassiz
|
||
</taxonomicName>
|
||
: vol. 3: 382.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6B0AB1C0214A" blockId="5.[159,781,1023,1787]" box="[159,749,1120,1144]" pageId="5" pageNumber="1523">
|
||
1833–1843
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE91436B0AB15D214A" authority="Agassiz" authorityName="Agassiz" box="[288,624,1120,1143]" class="Elasmobranchii" genus="Narcopterus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcanus">
|
||
<emphasis id="B958B80CFFE8FFCE91436B0AB123214A" box="[288,526,1120,1143]" italics="true" pageId="5" pageNumber="1523">Narcopterus bolcanus</emphasis>
|
||
Agassiz
|
||
</taxonomicName>
|
||
: vol. 4: 38.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6BEAB17621AA" blockId="5.[159,781,1023,1787]" box="[159,603,1152,1175]" pageId="5" pageNumber="1523">
|
||
1835
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6BEAB10021AA" authority="Agassiz" authorityName="Agassiz" box="[221,557,1152,1175]" class="Elasmobranchii" genus="Narcopterus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcanus">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6BEAB2E621AA" box="[221,459,1152,1175]" italics="true" pageId="5" pageNumber="1523">Narcopterus bolcanus</emphasis>
|
||
Agassiz
|
||
</taxonomicName>
|
||
: 14.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6BCAB2FA21E5" blockId="5.[159,781,1023,1787]" pageId="5" pageNumber="1523">
|
||
1851
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BC6BCAB134218A" authority="Heckel" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[223,537,1184,1207]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BC6BCAB293218A" box="[223,446,1184,1207]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
Heckel
|
||
</taxonomicName>
|
||
: 324 (first occurrence of name and description).
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6B8BB14C21C5" blockId="5.[159,781,1023,1787]" box="[159,609,1249,1272]" pageId="5" pageNumber="1523">
|
||
1854
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6B8BB26221C5" authorityName="Muller & Henle" authorityYear="1838" box="[221,335,1249,1272]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6B8BB26221C5" box="[221,335,1249,1272]" italics="true" pageId="5" pageNumber="1523">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
(?)
|
||
<emphasis id="B958B80CFFE8FFCE911D6B8BB2F921C5" box="[382,468,1249,1272]" italics="true" pageId="5" pageNumber="1523">bolcana</emphasis>
|
||
; Pictet: 277.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6A6BB18C2025" blockId="5.[159,781,1023,1787]" box="[159,673,1281,1304]" pageId="5" pageNumber="1523">
|
||
1860
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6A6BB13E2025" authority="Heckel" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[221,531,1281,1304]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6A6BB2962025" box="[221,443,1281,1304]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
Heckel
|
||
</taxonomicName>
|
||
; Molin: 587.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6A4BB1EA2005" blockId="5.[159,781,1023,1787]" box="[159,711,1313,1336]" pageId="5" pageNumber="1523">
|
||
1874
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6A4BB13E2005" authority="Heckel" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[221,531,1313,1336]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6A4BB2962005" box="[221,443,1313,1336]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
Heckel
|
||
</taxonomicName>
|
||
; De Zigno: 177.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6A2BB2202044" blockId="5.[159,781,1023,1787]" pageId="5" pageNumber="1523">
|
||
1894
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BC6A2BB1542065" authority="(Heckel) Molin" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[223,633,1345,1368]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BC6A2BB2ED2065" box="[223,448,1345,1368]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
(Heckel) Molin
|
||
</taxonomicName>
|
||
; Jaekel: 106, fig. 18.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6AE8B19220A4" blockId="5.[159,781,1023,1787]" box="[159,703,1410,1433]" pageId="5" pageNumber="1523">
|
||
1904
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6AE8B10920A4" authority="(Heckel)" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[221,548,1410,1433]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6AE8B29620A4" box="[221,443,1410,1433]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
(Heckel)
|
||
</taxonomicName>
|
||
; Eastman: 27.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6AC8B1E02084" blockId="5.[159,781,1023,1787]" box="[159,717,1442,1465]" pageId="5" pageNumber="1523">
|
||
1905
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6AC8B1092084" authority="(Heckel)" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[221,548,1442,1465]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6AC8B2962084" box="[221,443,1442,1465]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
(Heckel)
|
||
</taxonomicName>
|
||
; Eastman: 351.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6AA8B02A20E4" blockId="5.[159,781,1023,1787]" box="[159,775,1473,1497]" pageId="5" pageNumber="1523">
|
||
1922
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BA6AA8B14820E4" authority="(Agassiz) Heckel" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[217,613,1474,1497]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BA6AA8B28620E4" box="[217,427,1474,1497]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
(Agassiz) Heckel
|
||
</taxonomicName>
|
||
; D’ Erasmo: 12.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6A88B1A020C4" blockId="5.[159,781,1023,1787]" box="[159,653,1506,1529]" pageId="5" pageNumber="1523">
|
||
1980
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6A88B13E20C4" authority="Heckel" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[221,531,1506,1529]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6A88B29620C4" box="[221,443,1506,1529]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
Heckel
|
||
</taxonomicName>
|
||
; Blot: 344.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3363795FFE8FFCE90FC6968B2FE23C6" pageId="5" pageNumber="1523" type="reference_group">
|
||
<paragraph id="8B93641EFFE8FFCE90FC6968B1D42324" blockId="5.[159,781,1023,1787]" box="[159,761,1538,1561]" pageId="5" pageNumber="1523">
|
||
1987
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6968B1672324" authority="Molin, 1860" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[221,586,1538,1561]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6968B2962324" box="[221,443,1538,1561]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE91A76968B1672324" author="Molin, R." box="[452,586,1538,1561]" pageId="5" pageNumber="1523" pagination="582 - 588" refId="ref16443" refString="Molin, R. 1860. Primitiae Musei Archigymnasii patavini. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 40, 582 - 588." type="journal article" year="1860">Molin, 1860</bibRefCitation>
|
||
</taxonomicName>
|
||
; Cappetta: 139.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6948B1C62304" blockId="5.[159,781,1023,1787]" box="[159,747,1570,1593]" pageId="5" pageNumber="1523">
|
||
2004
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6948B2962304" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[221,443,1570,1593]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6948B2962304" box="[221,443,1570,1593]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
</taxonomicName>
|
||
; Carvalho: 78, fig. 12A, C.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6928B1D42364" blockId="5.[159,781,1023,1787]" box="[159,761,1602,1625]" pageId="5" pageNumber="1523">
|
||
2012
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE90BE6928B1672364" authority="Molin, 1860" authorityName="Molin" authorityYear="1860" baseAuthorityName="Heckel" box="[221,586,1602,1625]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE90BE6928B2962364" box="[221,443,1602,1625]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE91A76928B1672364" author="Molin, R." box="[452,586,1602,1625]" pageId="5" pageNumber="1523" pagination="582 - 588" refId="ref16443" refString="Molin, R. 1860. Primitiae Musei Archigymnasii patavini. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 40, 582 - 588." type="journal article" year="1860">Molin, 1860</bibRefCitation>
|
||
</taxonomicName>
|
||
; Cappetta: 346.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC6908B1AC23A7" blockId="5.[159,781,1023,1787]" pageId="5" pageNumber="1523">
|
||
2014
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE908B6908B1522344" authority="Heckel, 1851" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[232,639,1634,1657]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE908B6908B2FD2344" box="[232,464,1634,1657]" italics="true" pageId="5" pageNumber="1523">Platyrhina bolcensis</emphasis>
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE91876908B1522344" author="Heckel, M. J." box="[484,639,1634,1657]" pageId="5" pageNumber="1523" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
; Carnevale, Bannikov, Marram̀a, Tyler & Zorzin: 41.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE90FC69C9B2FE23C6" blockId="5.[159,781,1023,1787]" pageId="5" pageNumber="1523">
|
||
2018c ‘
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE916169C9B2D52387" authorityName="Heckel" authorityYear="1851" baseAuthorityName="Agassiz" box="[258,504,1698,1722]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis">
|
||
<emphasis id="B958B80CFFE8FFCE916169C9B2592387" box="[258,372,1699,1722]" italics="true" pageId="5" pageNumber="1523">Platyrhina</emphasis>
|
||
’
|
||
<emphasis id="B958B80CFFE8FFCE91F669C9B2D52387" box="[405,504,1699,1722]" italics="true" pageId="5" pageNumber="1523">bolcensis</emphasis>
|
||
</taxonomicName>
|
||
; Marram̀a, Carnevale, Engelbrecht, Claeson, Zorzin, Fornasiero & Carnevale: 287, fig. 12C.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3363795FFE8FFCE90FC687FB77C2735" pageId="5" pageNumber="1523" type="materials_examined">
|
||
<paragraph id="8B93641EFFE8FFCE90FC687FB1A22250" blockId="5.[159,781,1813,1901]" pageId="5" pageNumber="1523">
|
||
<materialsCitation id="3B446E43FFE8FFCE90FC687FB1A22250" collectionCode="DW" pageId="5" pageNumber="1523" specimenCode="MGP-PD 8873" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B958B80CFFE8FFCE90FC687FB2202211" bold="true" box="[159,269,1813,1836]" pageId="5" pageNumber="1523">
|
||
<typeStatus id="5497DABCFFE8FFCE90FC687FB2242211" box="[159,265,1813,1836]" pageId="5" pageNumber="1523" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DB8ACC65FFE8FFCE9175687FB2ED2211" box="[278,448,1813,1836]" collectionCode="MGP-PD" pageId="5" pageNumber="1523">MGP-PD 8873</specimenCode>
|
||
C/8874C, articulated skeleton in part and counterpart, lacking the caudal fin,
|
||
<quantity id="4CD4C9FBFFE8FFCE90FC683CB2252250" box="[159,264,1878,1901]" metricMagnitude="-1" metricUnit="m" metricValue="3.385" pageId="5" pageNumber="1523" unit="mm" value="338.5">338.5 mm</quantity>
|
||
disc width (
|
||
<collectionCode id="ED3DFCDBFFE8FFCE91F0683CB2EE2250" box="[403,451,1878,1901]" pageId="5" pageNumber="1523">DW</collectionCode>
|
||
, hereafter;
|
||
<figureCitation id="1317789BFFE8FFCE925C683CB1522250" box="[575,639,1878,1901]" captionStart="Figure 2" captionStartId="6.[159,226,1315,1336]" captionTargetBox="[159,1466,189,1297]" captionTargetId="figure-269@6.[159,1467,188,1298]" captionTargetPageId="6" captionText="Figure 2. The holotype of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site in part and counterpart. A, MGP-PD 8873C; B, MGP-PD 8874C. Scale bars = 100mm." figureDoi="http://doi.org/10.5281/zenodo.10932531" httpUri="https://zenodo.org/record/10932531/files/figure.png" pageId="5" pageNumber="1523">Fig. 2</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE932F6FD7B0B3244B" blockId="5.[844,1467,189,374]" pageId="5" pageNumber="1523">
|
||
<emphasis id="B958B80CFFE8FFCE932F6FD7B71C25E9" bold="true" box="[844,1073,189,212]" pageId="5" pageNumber="1523">Referred material.</emphasis>
|
||
<collectionCode id="ED3DFCDBFFE8FFCE94596FD7B7B225E9" box="[1082,1183,189,212]" pageId="5" pageNumber="1523">MGP-PD</collectionCode>
|
||
26279C/26280C, completely articulated skeleton in part and counterpart,
|
||
<quantity id="4CD4C9FBFFE8FFCE932F6F94B09B2428" box="[844,950,254,277]" metricMagnitude="-1" metricUnit="m" metricValue="3.842" pageId="5" pageNumber="1523" unit="mm" value="384.2">384.2 mm</quantity>
|
||
DW,
|
||
<quantity id="4CD4C9FBFFE8FFCE946F6F94B75B2428" box="[1036,1142,254,277]" metricMagnitude="-1" metricUnit="m" metricValue="8.403" pageId="5" pageNumber="1523" unit="mm" value="840.3">840.3 mm</quantity>
|
||
TL (
|
||
<figureCitation id="1317789BFFE8FFCE94A06F94B6642428" box="[1219,1353,254,277]" captionStart="Figure 3" captionStartId="7.[159,226,782,803]" captionTargetBox="[159,1467,188,765]" captionTargetId="figure-528@7.[159,1467,188,765]" captionTargetPageId="7" captionText="Figure 3. Specimens of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C; B, original historical plate of the specimen MGP-PD 26279C, illustrated by Achille de Zigno (1813–1892). Photo: courtesy of Universit̀a degli Studi di Padova; C, specimen MGGC 7449; D, MGGC 7449 under ultraviolet light. Scale bars = 100mm." figureDoi="http://doi.org/10.5281/zenodo.10932533" httpUri="https://zenodo.org/record/10932533/files/figure.png" pageId="5" pageNumber="1523">Fig. 3A, B</figureCitation>
|
||
);
|
||
<collectionCode id="ED3DFCDBFFE8FFCE95096F94B6972428" box="[1386,1466,254,277]" pageId="5" pageNumber="1523">MGGC</collectionCode>
|
||
7449/7450, articulated skeleton in part and counterpart, lacking dorsal and caudal fins,
|
||
<quantity id="4CD4C9FBFFE8FFCE94AD6E55B615246B" box="[1230,1336,319,342]" metricMagnitude="-1" metricUnit="m" metricValue="3.7939999999999996" pageId="5" pageNumber="1523" unit="mm" value="379.4">379.4 mm</quantity>
|
||
DW (
|
||
<figureCitation id="1317789BFFE8FFCE95F36E55B0A3244B" captionStart="Figure 3" captionStartId="7.[159,226,782,803]" captionTargetBox="[159,1467,188,765]" captionTargetId="figure-528@7.[159,1467,188,765]" captionTargetPageId="7" captionText="Figure 3. Specimens of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C; B, original historical plate of the specimen MGP-PD 26279C, illustrated by Achille de Zigno (1813–1892). Photo: courtesy of Universit̀a degli Studi di Padova; C, specimen MGGC 7449; D, MGGC 7449 under ultraviolet light. Scale bars = 100mm." figureDoi="http://doi.org/10.5281/zenodo.10932533" httpUri="https://zenodo.org/record/10932533/files/figure.png" pageId="5" pageNumber="1523">Fig. 3C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE932F6EE5B77C2735" blockId="5.[844,1467,399,520]" pageId="5" pageNumber="1523">
|
||
<emphasis id="B958B80CFFE8FFCE932F6EE5B7A1249B" bold="true" box="[844,1164,399,422]" pageId="5" pageNumber="1523">Type locality and horizon.</emphasis>
|
||
Monte Postale site, Bolca Konservat-Lagerst¨atte,
|
||
<collectingCountry id="F33B248EFFE8FFCE94376EC5B7A524FB" box="[1108,1160,431,454]" name="Italy" pageId="5" pageNumber="1523">Italy</collectingCountry>
|
||
; early Eocene, Ypresian, middle Cuisian, SBZ 11 (
|
||
<collectionCode id="ED3DFCDBFFE8FFCE940A6EBAB7A024DA" box="[1129,1165,464,487]" pageId="5" pageNumber="1523">NP</collectionCode>
|
||
13, CNE 5); 50.7–48.9 Ma (
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE93376E9BB76C2735" author="Papazzoni, C. A. & Fornaciari, E. & Giusberti, L. & Vescogni, A. & Fornaciari, B." box="[852,1089,497,520]" pageId="5" pageNumber="1523" pagination="1 - 12" refId="ref16756" refString="Papazzoni, C. A., Fornaciari, E., Giusberti, L., Vescogni, A. & Fornaciari, B. 2017. Integrating shallow benthic and calcareous nannofossil zones: the lower Eocene of the Monte Postale section (Northern Italy). Palaios, 32, 1 - 12. doi: 10.2110 / palo. 2016.014" type="journal article" year="2017">
|
||
Papazzoni
|
||
<emphasis id="B958B80CFFE8FFCE93AF6E9BB0D02735" box="[972,1021,497,520]" italics="true" pageId="5" pageNumber="1523">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3363795FFE8FFCE932F6D4AB7AA270A" box="[844,1159,544,567]" pageId="5" pageNumber="1523" type="diagnosis">
|
||
<paragraph id="8B93641EFFE8FFCE932F6D4AB7AA270A" blockId="5.[844,1159,544,567]" box="[844,1159,544,567]" pageId="5" pageNumber="1523">
|
||
<emphasis id="B958B80CFFE8FFCE932F6D4AB0EC270A" bold="true" box="[844,961,544,567]" pageId="5" pageNumber="1523">Diagnosis.</emphasis>
|
||
As for the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3363795FFE8FFC1932F6D19B2962379" lastPageId="10" lastPageNumber="1528" pageId="5" pageNumber="1523" type="description">
|
||
<paragraph id="8B93641EFFE8FFCE932F6D19B0F027B0" blockId="5.[844,1467,627,1203]" box="[844,989,627,653]" pageId="5" pageNumber="1523">
|
||
<emphasis id="B958B80CFFE8FFCE932F6D19B0F027B0" bold="true" box="[844,989,627,653]" pageId="5" pageNumber="1523">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCE932F6DFEB7F1218E" blockId="5.[844,1467,627,1203]" pageId="5" pageNumber="1523">
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE93396DFFB6232791" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[858,1294,661,684]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE8FFCE93396DFFB7752791" box="[858,1112,661,684]" italics="true" pageId="5" pageNumber="1523">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE940E6DFFB6292791" author="Heckel, M. J." box="[1133,1284,661,684]" pageId="5" pageNumber="1523" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE8FFCE957F6DFFB6B62791" box="[1308,1435,661,684]" pageId="5" pageNumber="1523" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is represented by three partially complete articulated specimens in part and counterpart (
|
||
<figureCitation id="1317789BFFE8FFCE94C46DBFB7C327D1" box="[1191,1262,725,748]" captionStart="Figure 2" captionStartId="6.[159,226,1315,1336]" captionTargetBox="[159,1466,189,1297]" captionTargetId="figure-269@6.[159,1467,188,1298]" captionTargetPageId="6" captionText="Figure 2. The holotype of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site in part and counterpart. A, MGP-PD 8873C; B, MGP-PD 8874C. Scale bars = 100mm." figureDoi="http://doi.org/10.5281/zenodo.10932531" httpUri="https://zenodo.org/record/10932531/files/figure.png" pageId="5" pageNumber="1523">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="1317789BFFE8FFCE95626DBFB62327D1" box="[1281,1294,725,748]" captionStart="Figure 3" captionStartId="7.[159,226,782,803]" captionTargetBox="[159,1467,188,765]" captionTargetId="figure-528@7.[159,1467,188,765]" captionTargetPageId="7" captionText="Figure 3. Specimens of †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C; B, original historical plate of the specimen MGP-PD 26279C, illustrated by Achille de Zigno (1813–1892). Photo: courtesy of Universit̀a degli Studi di Padova; C, specimen MGGC 7449; D, MGGC 7449 under ultraviolet light. Scale bars = 100mm." figureDoi="http://doi.org/10.5281/zenodo.10932533" httpUri="https://zenodo.org/record/10932533/files/figure.png" pageId="5" pageNumber="1523">3</figureCitation>
|
||
), including the
|
||
<typeStatus id="5497DABCFFE8FFCE932F6D9CB0842630" box="[844,937,758,781]" pageId="5" pageNumber="1523" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED3DFCDBFFE8FFCE93A36D9CB70A2630" box="[960,1063,758,781]" pageId="5" pageNumber="1523">MGP-PD</collectionCode>
|
||
8873C/8874C) and two additional specimens (
|
||
<collectionCode id="ED3DFCDBFFE8FFCE93AF6C7CB71E2610" box="[972,1075,790,813]" pageId="5" pageNumber="1523">MGP-PD</collectionCode>
|
||
26279C/26280C and
|
||
<collectionCode id="ED3DFCDBFFE8FFCE95476C7CB6592610" box="[1316,1396,790,813]" pageId="5" pageNumber="1523">MGGC</collectionCode>
|
||
7449/ 7450). Counts and measurements are listed in the Supplemental material (File 1,
|
||
<tableCitation id="C6AE51A5FFE8FFCE94C76C3DB62B2653" box="[1188,1286,855,878]" captionStart="Table 1" captionStartId="14.[159,218,707,728]" captionTargetPageId="14" captionText="Table 1. Tree statistics for parsimony analyses. Abbreviations: CH, coding follows Compagno (1999); CI, consistency index; DS, coding follows Da Silva et al. (2018); RI, retention index. In CH-85 and DS-85 analyses were run excluding the character 85. In CH-HetSqua and DS-HetSqua the taxa Heterodontus and Squalus were excluded from -85 nexus files." httpUri="http://table.plazi.org/id/DF533496FFE3FFC590FC6DA9B7BF2633" pageId="5" pageNumber="1523" tableUuid="DF533496FFE3FFC590FC6DA9B7BF2633">Table S1</tableCitation>
|
||
). The examined specimens are similar in size. The largest one measures
|
||
<quantity id="4CD4C9FBFFE8FFCE932F6CF2B0A02692" box="[844,909,920,943]" metricMagnitude="-1" metricUnit="m" metricValue="8.4" pageId="5" pageNumber="1523" unit="cm" value="84.0">84 cm</quantity>
|
||
TL and
|
||
<quantity id="4CD4C9FBFFE8FFCE94746CF2B77A2692" box="[1047,1111,920,943]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="1523" unit="cm" value="38.0">38 cm</quantity>
|
||
DW. The pectoral disc of †
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE93396CD3B0CA26ED" authority="Marram̀ & Carneval & Claeso & Naylo & Kriwe, 2020" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" box="[858,999,953,976]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B958B80CFFE8FFCE93396CD3B0CA26ED" box="[858,999,953,976]" italics="true" pageId="5" pageNumber="1523">Eoplatyrhina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE8FFCE93906CD3B77126ED" box="[1011,1116,953,976]" pageId="5" pageNumber="1523" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is notably expanded, ovoid or shovel shaped, slightly longer than wide and reaching its maximum width just posterior to its mid-length. The snout is broad and rounded. The tail is not very stout, slightly longer than disc length, with two dorsal fins inserting posteriorly on the tail. The overall body shape and proportions are similar to those of the extant thornbacks
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE93F16BF6B729218E" authorityName="Muller & Henle" authorityYear="1838" box="[914,1028,1180,1203]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE93F16BF6B729218E" box="[914,1028,1180,1203]" italics="true" pageId="5" pageNumber="1523">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE945F6BF6B7F8218E" authorityName="Garman" authorityYear="1881" box="[1084,1237,1180,1203]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE945F6BF6B7F8218E" box="[1084,1237,1180,1203]" italics="true" pageId="5" pageNumber="1523">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE8FFCD932F6BA6B0C12326" blockId="5.[844,1467,1228,1901]" lastBlockId="6.[844,1467,1411,1563]" lastPageId="6" lastPageNumber="1524" pageId="5" pageNumber="1523">
|
||
<emphasis id="B958B80CFFE8FFCE932F6BA6B0D621DE" bold="true" box="[844,1019,1228,1251]" pageId="5" pageNumber="1523">Neurocranium.</emphasis>
|
||
The rostral cartilage fails to reach the anterior margin of the disc, as in all platyrhinids. This element is long and tapers gradually anteriorly (
|
||
<figureCitation id="1317789BFFE8FFCE950E6A67B6992019" box="[1389,1460,1293,1316]" captionStart="Figure 4" captionStartId="8.[159,226,1157,1178]" captionTargetBox="[159,1460,188,1140]" captionTargetId="figure-295@8.[159,1467,188,1140]" captionTargetPageId="8" captionText="Figure 4. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGGC 7449, close-up of the head and pectoral girdle under UV light; B, reconstruction. Abbreviations: af, anterior fontanel; ao, antorbital cartilage; cb5, fifth ceratobranchial; hp, horn-like process; hyo, hyomandibula; mc, Meckel’s cartilage; mes, mesopterygium; met, metapterygium; nc, nasal capsule; pq, palatoquadrate; pro, propterygium; ro, rostral cartilage; rp, rostral process; sca, scapulocoracoid; syn, synarcual. Scale bars = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10932537" httpUri="https://zenodo.org/record/10932537/files/figure.png" pageId="5" pageNumber="1523">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="1317789BFFE8FFCE932F6A44B0402078" box="[844,877,1326,1349]" captionStart="Figure 5" captionStartId="9.[159,226,951,972]" captionTargetBox="[159,1467,188,933]" captionTargetId="figure-57@9.[159,1467,188,933]" captionTargetPageId="9" captionText="Figure 5. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C, close-up of the rostral cartilage; B, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula; C, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin. Abbreviations: af, anterior fontanel; co, coracoid bar of the scapulocoracoid; mes, mesopterygium; met, metapterygium; pro, propterygium; ro, rostral cartilage; scap, scapular process of the scapulocoracoid; ss, suprascapulae; syn, synarcual. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10932541" httpUri="https://zenodo.org/record/10932541/files/figure.png" pageId="5" pageNumber="1523">5A</figureCitation>
|
||
), resembling the condition typical of
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE95416A44B6962078" authorityName="Garman" authorityYear="1881" box="[1314,1467,1326,1349]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE95416A44B6962078" box="[1314,1467,1326,1349]" italics="true" pageId="5" pageNumber="1523">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE93F16A24B72C2058" authorityName="de Carvalho" authorityYear="2004" box="[914,1025,1358,1381]" class="Elasmobranchii" family="Rhinobatidae" genus="Tethybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE93F16A24B72C2058" box="[914,1025,1358,1381]" italics="true" pageId="5" pageNumber="1523">Tethybatis</emphasis>
|
||
</taxonomicName>
|
||
, and differs from the short rostrum observed in †
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE93886A05B74820BB" authorityName="Claeson, Underwood & Ward" authorityYear="2013" box="[1003,1125,1391,1414]" class="Elasmobranchii" family="Rhinobatidae" genus="Tingitanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE8FFCE93886A05B74820BB" box="[1003,1125,1391,1414]" italics="true" pageId="5" pageNumber="1523">Tingitanius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B958B80CFFE8FFCE94C56A05B60D20BB" box="[1190,1312,1391,1414]" italics="true" pageId="5" pageNumber="1523">
|
||
<taxonomicName id="4C2C1F9DFFE8FFCE94C56A05B63120BB" authorityName="Muller & Henle" authorityYear="1838" box="[1190,1308,1391,1414]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="5" pageNumber="1523" phylum="Chordata" rank="genus">Platyrhina</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Unlike other platyrhinids, the anterior margin of the rostral cartilage is not pointed but trough-shaped, with the rostral node slightly expanded laterally (
|
||
<figureCitation id="1317789BFFE8FFCE94E66ABAB7E620DA" box="[1157,1227,1488,1511]" captionStart="Figure 4" captionStartId="8.[159,226,1157,1178]" captionTargetBox="[159,1460,188,1140]" captionTargetId="figure-295@8.[159,1467,188,1140]" captionTargetPageId="8" captionText="Figure 4. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGGC 7449, close-up of the head and pectoral girdle under UV light; B, reconstruction. Abbreviations: af, anterior fontanel; ao, antorbital cartilage; cb5, fifth ceratobranchial; hp, horn-like process; hyo, hyomandibula; mc, Meckel’s cartilage; mes, mesopterygium; met, metapterygium; nc, nasal capsule; pq, palatoquadrate; pro, propterygium; ro, rostral cartilage; rp, rostral process; sca, scapulocoracoid; syn, synarcual. Scale bars = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10932537" httpUri="https://zenodo.org/record/10932537/files/figure.png" pageId="5" pageNumber="1523">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="1317789BFFE8FFCE94BD6ABAB7D220DA" box="[1246,1279,1488,1511]" captionStart="Figure 5" captionStartId="9.[159,226,951,972]" captionTargetBox="[159,1467,188,933]" captionTargetId="figure-57@9.[159,1467,188,933]" captionTargetPageId="9" captionText="Figure 5. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C, close-up of the rostral cartilage; B, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula; C, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin. Abbreviations: af, anterior fontanel; co, coracoid bar of the scapulocoracoid; mes, mesopterygium; met, metapterygium; pro, propterygium; ro, rostral cartilage; scap, scapular process of the scapulocoracoid; ss, suprascapulae; syn, synarcual. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10932541" httpUri="https://zenodo.org/record/10932541/files/figure.png" pageId="5" pageNumber="1523">5A</figureCitation>
|
||
). Rostral appendices at the tip of the rostrum are absent. A small rod-like process lateral to the rostral cartilage and just anterior to the nasal capsule in
|
||
<collectionCode id="ED3DFCDBFFE8FFCE94F36958B7CD2374" box="[1168,1248,1586,1609]" pageId="5" pageNumber="1523">MGGC</collectionCode>
|
||
7449/7450 can be interpreted as one of the two rostral processes, which are uniquely present in extant thornbacks. Although
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE932F69F9B75E2397" author="McEachran, J. D. & Dunn, K. A. & Miyake, T." box="[844,1139,1683,1706]" pageId="5" pageNumber="1523" refId="ref16313" refString="McEachran, J. D., Dunn, K. A. & Miyake, T. 1996. Interrelationships of the batoid fishes (Chondrichthyes: Batoidea). Pp. 63 - 84 in M. L. J. Stassney, L. R. Parenti & G. D. Johnson (eds) Interrelationships of fishes. Academic Press, San Diego." type="book" year="1996">
|
||
McEachran
|
||
<emphasis id="B958B80CFFE8FFCE93BA69F9B73E2397" box="[985,1043,1683,1706]" italics="true" pageId="5" pageNumber="1523">et al</emphasis>
|
||
. (1996)
|
||
</bibRefCitation>
|
||
considered these structures homologous to the rostral appendices of skates and guitarfishes,
|
||
<bibRefCitation id="EFBD19EFFFE8FFCE93DF69BEB75E23D6" author="Carvalho, M. R. de" box="[956,1139,1748,1771]" pageId="5" pageNumber="1523" refId="ref14177" refString="Carvalho, M. R. de. 2004. A Late Cretaceous thornback ray from southern Italy, with a phylogenetic reappraisal of the Platyrhinidae (Chondrichthyes: Batoidea). Pp. 75 - 100 in G. Arratia & A. Tintori (eds) Mesozoic fishes 3 - Systematics, paleoenvironments and biodiversity. Verlag Dr Friedrich Pfeil, Munich." type="book" year="2004">Carvalho (2004)</bibRefCitation>
|
||
pointed out that the rostral processes of platyrhinids, originating ventral to the rostral cartilage, might represent outgrowths of the lamina orbitonasalis, unlike the rostral appendices that are secondary chondrifications fused laterally to the rostral node. The nasal capsules are ovoid, laterally expanded, and at right angles to the rostrum, as in †
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD921A6AC9B1C52087" authorityName="de Carvalho" authorityYear="2004" box="[633,744,1443,1466]" class="Elasmobranchii" family="Rhinobatidae" genus="Tethybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEBFFCD921A6AC9B1C52087" box="[633,744,1443,1466]" italics="true" pageId="6" pageNumber="1524">Tethybatis</emphasis>
|
||
</taxonomicName>
|
||
. A single small horn-like process (= tab-like process of
|
||
<bibRefCitation id="EFBD19EFFFEBFFCD90FC6A89B25120C7" author="Claeson, K. M. & Underwood, C. J. & Ward, D. J." box="[159,380,1507,1530]" pageId="6" pageNumber="1524" pagination="1019 - 1036" refId="ref14384" refString="Claeson, K. M., Underwood, C. J. & Ward, D. J. 2013. Tingitanius tenuimandibulus, a new platyrhinid batoid from the Turonian (Cretaceous) of Morocco and the Cretaceous radiation of the Platyrhinidae. Journal of Vertebrate Paleontology, 33, 1019 - 1036. doi: 10.1080 / 02724634.2013. 767266" type="journal article" year="2013">
|
||
Claeson
|
||
<emphasis id="B958B80CFFEBFFCD91636A89B21920C7" box="[256,308,1507,1530]" italics="true" pageId="6" pageNumber="1524">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
) can be recognized on the anterior margin of each nasal capsule, similar to the extant platyrhinids and †
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD912B6949B2EC2307" authorityName="Claeson, Underwood & Ward" authorityYear="2013" box="[328,449,1571,1594]" class="Elasmobranchii" family="Rhinobatidae" genus="Tingitanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEBFFCD912B6949B2EC2307" box="[328,449,1571,1594]" italics="true" pageId="6" pageNumber="1524">Tingitanius</emphasis>
|
||
</taxonomicName>
|
||
. The antorbital cartilages are well developed and plate-like and have an irregular outline. They project laterally from the postero-lateral margin of the nasal capsules and articulate distally with the propterygia. It is difficult to distinguish the preorbital process or the jugal arch, but a small and narrow postorbital process can be recognized in the otic region, just posterior to the supraorbital crest. The orbital region is longer than wide. The anterior fontanel extends through almost the entire length of the rostral cartilage and resembles an isosceles triangle with a close and concave posterior border, similar to the condition seen in †
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD93396AAEB0F920E6" authorityName="Claeson, Underwood & Ward" authorityYear="2013" box="[858,980,1476,1499]" class="Elasmobranchii" family="Rhinobatidae" genus="Tingitanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEBFFCD93396AAEB0F920E6" box="[858,980,1476,1499]" italics="true" pageId="6" pageNumber="1524">Tingitanius</emphasis>
|
||
</taxonomicName>
|
||
, and in contrast to the oval-shaped fontanel of
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD93116A8EB0C920C6" authorityName="Muller & Henle" authorityYear="1838" box="[882,996,1508,1531]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEBFFCD93116A8EB0C920C6" box="[882,996,1508,1531]" italics="true" pageId="6" pageNumber="1524">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
, or to the figure-eight shape typical of
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD932F696EB0C82326" authorityName="Garman" authorityYear="1881" box="[844,997,1540,1563]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEBFFCD932F696EB0C82326" box="[844,997,1540,1563]" italics="true" pageId="6" pageNumber="1524">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF533496FFEBFFCD90FC6A49B03E206E" ID-DOI="http://doi.org/10.5281/zenodo.10932531" ID-Zenodo-Dep="10932531" httpUri="https://zenodo.org/record/10932531/files/figure.png" pageId="6" pageNumber="1524" startId="6.[159,226,1315,1336]" targetBox="[159,1466,189,1297]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B93641EFFEBFFCD90FC6A49B03E206E" blockId="6.[159,1468,1315,1364]" pageId="6" pageNumber="1524">
|
||
<emphasis id="B958B80CFFEBFFCD90FC6A49B3D02005" bold="true" box="[159,253,1315,1336]" pageId="6" pageNumber="1524">Figure 2.</emphasis>
|
||
The holotype of †
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD91DE6A49B01B2005" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[445,822,1315,1336]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFEBFFCD91DE6A49B1B12005" box="[445,668,1315,1336]" italics="true" pageId="6" pageNumber="1524">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFEBFFCD92C86A49B0032005" author="Heckel, M. J." box="[683,814,1315,1336]" pageId="6" pageNumber="1524" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFEBFFCD935C6A49B0832005" box="[831,942,1315,1336]" pageId="6" pageNumber="1524" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site in part and counterpart.
|
||
<emphasis id="B958B80CFFEBFFCD90FC6A54B39D206E" bold="true" box="[159,176,1342,1363]" pageId="6" pageNumber="1524">A</emphasis>
|
||
, MGP-PD 8873C;
|
||
<emphasis id="B958B80CFFEBFFCD910D6A54B253206E" bold="true" box="[366,382,1342,1363]" pageId="6" pageNumber="1524">B</emphasis>
|
||
, MGP-PD 8874C. Scale bars = 100 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93641EFFEBFFCC932F695EB23521E1" blockId="6.[844,1467,1588,1901]" lastBlockId="7.[159,781,900,1244]" lastPageId="7" lastPageNumber="1525" pageId="6" pageNumber="1524">
|
||
<emphasis id="B958B80CFFEBFFCD932F695EB62F2376" bold="true" box="[844,1282,1588,1611]" pageId="6" pageNumber="1524">Jaws, hyoid and gill arches.</emphasis>
|
||
Specimens of †
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD9339693EB7712356" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[858,1116,1620,1643]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFEBFFCD9339693EB7712356" box="[858,1116,1620,1643]" italics="true" pageId="6" pageNumber="1524">Eoplatyrhina bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFEBFFCD940D693EB7DC2356" box="[1134,1265,1620,1643]" pageId="6" pageNumber="1524" rank="species">comb. nov.</taxonomicNameLabel>
|
||
are mostly preserved in dorsal view, obscuring the jaws, which are displaced and difficult to describe (
|
||
<figureCitation id="1317789BFFEBFFCD94B169FFB63F2391" box="[1234,1298,1685,1708]" captionStart="Figure 4" captionStartId="8.[159,226,1157,1178]" captionTargetBox="[159,1460,188,1140]" captionTargetId="figure-295@8.[159,1467,188,1140]" captionTargetPageId="8" captionText="Figure 4. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGGC 7449, close-up of the head and pectoral girdle under UV light; B, reconstruction. Abbreviations: af, anterior fontanel; ao, antorbital cartilage; cb5, fifth ceratobranchial; hp, horn-like process; hyo, hyomandibula; mc, Meckel’s cartilage; mes, mesopterygium; met, metapterygium; nc, nasal capsule; pq, palatoquadrate; pro, propterygium; ro, rostral cartilage; rp, rostral process; sca, scapulocoracoid; syn, synarcual. Scale bars = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10932537" httpUri="https://zenodo.org/record/10932537/files/figure.png" pageId="6" pageNumber="1524">Fig. 4</figureCitation>
|
||
). For the same reason, teeth are not exposed in any specimen, and therefore their morphology remains unknown. It is also unclear whether the labial cartilages are present, as in mature specimens of
|
||
<emphasis id="B958B80CFFEBFFCD945B687CB79C2210" box="[1080,1201,1814,1837]" italics="true" pageId="6" pageNumber="1524">
|
||
<taxonomicName id="4C2C1F9DFFEBFFCD945B687CB7832210" authorityName="Muller & Henle" authorityYear="1838" box="[1080,1198,1814,1837]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="6" pageNumber="1524" phylum="Chordata" rank="genus">Platyrhina</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The hyomandibulae are stout, robust and slightly arched, with a concave inner margin, narrow at their medial section. They project anterolaterally. As in †
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC91FC6CEFB12326A1" authorityName="de Carvalho" authorityYear="2004" box="[415,526,901,924]" class="Elasmobranchii" family="Rhinobatidae" genus="Tethybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC91FC6CEFB12326A1" box="[415,526,901,924]" italics="true" pageId="7" pageNumber="1525">Tethybatis</emphasis>
|
||
</taxonomicName>
|
||
, there is a large space between the hyomandibulae and mandibular arch, which is interpreted by
|
||
<bibRefCitation id="EFBD19EFFFEAFFCC910B6CAFB13226E1" author="Carvalho, M. R. de" box="[360,543,965,988]" pageId="7" pageNumber="1525" refId="ref14177" refString="Carvalho, M. R. de. 2004. A Late Cretaceous thornback ray from southern Italy, with a phylogenetic reappraisal of the Platyrhinidae (Chondrichthyes: Batoidea). Pp. 75 - 100 in G. Arratia & A. Tintori (eds) Mesozoic fishes 3 - Systematics, paleoenvironments and biodiversity. Verlag Dr Friedrich Pfeil, Munich." type="book" year="2004">Carvalho (2004)</bibRefCitation>
|
||
as indicative of the presence of a large spiracular opening. In radiographs, this space is not present in
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC91B26B6FB1472121" authorityName="Garman" authorityYear="1881" box="[465,618,1029,1052]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC91B26B6FB1472121" box="[465,618,1029,1052]" italics="true" pageId="7" pageNumber="1525">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC92F76B6FB02B2121" authorityName="Muller & Henle" authorityYear="1838" box="[660,774,1029,1052]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC92F76B6FB02B2121" box="[660,774,1029,1052]" italics="true" pageId="7" pageNumber="1525">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
, while it is present in
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC91F46B4FB1252101" authorityName="Garman" authorityYear="1913" box="[407,520,1061,1084]" class="Elasmobranchii" family="Rhinobatidae" genus="Zanobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC91F46B4FB1252101" box="[407,520,1061,1084]" italics="true" pageId="7" pageNumber="1525">Zanobatus</emphasis>
|
||
</taxonomicName>
|
||
. The distal part of the hyomandibulae appears taphonomically separated from the Meckel’ s cartilage. The fifth ceratobranchials articulate with the anterior margin of the scapulocoracoid, and the remaining gill arches are poorly preserved or missing.
|
||
</paragraph>
|
||
<caption id="DF533496FFEAFFCC90FC6C64B67A2664" ID-DOI="http://doi.org/10.5281/zenodo.10932533" ID-Zenodo-Dep="10932533" httpUri="https://zenodo.org/record/10932533/files/figure.png" pageId="7" pageNumber="1525" startId="7.[159,226,782,803]" targetBox="[159,1467,188,765]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B93641EFFEAFFCC90FC6C64B67A2664" blockId="7.[159,1467,782,857]" pageId="7" pageNumber="1525">
|
||
<emphasis id="B958B80CFFEAFFCC90FC6C64B3D3261E" bold="true" box="[159,254,782,803]" pageId="7" pageNumber="1525">Figure 3.</emphasis>
|
||
Specimens of †
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC91C76C64B033261E" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[420,798,782,803]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFEAFFCC91C76C64B1AF261E" box="[420,642,782,803]" italics="true" pageId="7" pageNumber="1525">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFEAFFCC92F06C64B03B261E" author="Heckel, M. J." box="[659,790,782,803]" pageId="7" pageNumber="1525" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFEAFFCC934B6C64B0BA261E" box="[808,919,782,803]" pageId="7" pageNumber="1525" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site.
|
||
<emphasis id="B958B80CFFEAFFCC94A76C64B7F8261E" bold="true" box="[1220,1237,782,803]" pageId="7" pageNumber="1525">A</emphasis>
|
||
, MGP-PD 26279C;
|
||
<emphasis id="B958B80CFFEAFFCC95C66C64B698261E" bold="true" box="[1445,1461,782,803]" pageId="7" pageNumber="1525">B</emphasis>
|
||
, original historical plate of the specimen MGP-PD 26279C, illustrated by Achille de Zigno (1813–1892). Photo: courtesy of Universit̀a degli Studi di Padova;
|
||
<emphasis id="B958B80CFFEAFFCC91896C2EB2D62664" bold="true" box="[490,507,836,857]" pageId="7" pageNumber="1525">C</emphasis>
|
||
, specimen MGGC 7449;
|
||
<emphasis id="B958B80CFFEAFFCC929B6C2EB0242664" bold="true" box="[760,777,836,857]" pageId="7" pageNumber="1525">D</emphasis>
|
||
, MGGC 7449 under ultraviolet light. Scale bars = 100 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93641EFFEAFFCC90FC6B9FB0F52681" blockId="7.[159,782,1269,1901]" lastBlockId="7.[844,1467,901,956]" pageId="7" pageNumber="1525">
|
||
<emphasis id="B958B80CFFEAFFCC90FC6B9FB1332031" bold="true" box="[159,542,1269,1292]" pageId="7" pageNumber="1525">Synarcual and vertebral column.</emphasis>
|
||
Although the synarcual can be identified as a tubular mineralized structure between the neurocranium and scapulocoracoid, its morphology remains ambiguous. The dorsally exposed specimens obscure the pattern of free centra. In †
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC90CF6AFFB20B2091" authorityName="Claeson, Underwood & Ward" authorityYear="2013" box="[172,294,1429,1452]" class="Elasmobranchii" family="Rhinobatidae" genus="Tingitanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC90CF6AFFB20B2091" box="[172,294,1429,1452]" italics="true" pageId="7" pageNumber="1525">Tingitanius</emphasis>
|
||
</taxonomicName>
|
||
, the first exposed vertebral centrum of the synarcual is located posterior to the articulation of the suprascapular cartilage with the synarcual. In
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC90FC6A9CB23C2330" authorityName="Muller & Henle" authorityYear="1838" box="[159,273,1526,1549]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC90FC6A9CB23C2330" box="[159,273,1526,1549]" italics="true" pageId="7" pageNumber="1525">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
, the first free centrum is situated at the level of the scapulocoracoid articulation with the synarcual. In
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC90A3695CB2742370" authorityName="Garman" authorityYear="1881" box="[192,345,1590,1613]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFEAFFCC90A3695CB2742370" box="[192,345,1590,1613]" italics="true" pageId="7" pageNumber="1525">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
, the first free centrum is rostral to the scapulocoracoid articulation with the synarcual. The vertebral column of †
|
||
<taxonomicName id="4C2C1F9DFFEAFFCC911D691CB15023B0" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[382,637,1654,1677]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="7" pageNumber="1525" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFEAFFCC911D691CB15023B0" box="[382,637,1654,1677]" italics="true" pageId="7" pageNumber="1525">Eoplatyrhina bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFEAFFCC92EE691CB02023B0" box="[653,781,1654,1677]" pageId="7" pageNumber="1525" rank="species">comb. nov.</taxonomicNameLabel>
|
||
consists of about 132 vertebral centra, in the most complete specimen
|
||
<collectionCode id="ED3DFCDBFFEAFFCC913F69DCB2EC23F0" box="[348,449,1718,1741]" pageId="7" pageNumber="1525">MGP-PD</collectionCode>
|
||
26279C/26280C. There are 20–24 trunk centra (from the first distinguishable centrum to the anterior margin of the puboischiadic bar), and 113–118 from the puboischiadic bar to the tip of the tail (of these, about 23 are caudal). The vertebral centra are highly calcified, sub-rectangular in shape and anteroposteriorly compressed. There are about 15 or 16 pairs of ribs.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFEAFFC3932F6CBFB2F320DF" blockId="7.[844,1467,981,1902]" lastBlockId="8.[159,781,1350,1901]" lastPageId="8" lastPageNumber="1526" pageId="7" pageNumber="1525">
|
||
<emphasis id="B958B80CFFEAFFCC932F6CBFB7C926D1" bold="true" box="[844,1252,981,1004]" pageId="7" pageNumber="1525">Appendicular skeleton and fins.</emphasis>
|
||
It is difficult to describe the morphology of the coracoid bar because the specimens are mostly exposed in dorsal view, but the scapular processes of the scapulocoracoid seem to be short in
|
||
<collectionCode id="ED3DFCDBFFEAFFCC93CA6B3FB7232151" box="[937,1038,1109,1132]" pageId="7" pageNumber="1525">MGP-PD</collectionCode>
|
||
26279C/80C (
|
||
<figureCitation id="1317789BFFEAFFCC94D16B3FB6292151" box="[1202,1284,1109,1132]" captionStart="Figure 5" captionStartId="9.[159,226,951,972]" captionTargetBox="[159,1467,188,933]" captionTargetId="figure-57@9.[159,1467,188,933]" captionTargetPageId="9" captionText="Figure 5. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C, close-up of the rostral cartilage; B, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula; C, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin. Abbreviations: af, anterior fontanel; co, coracoid bar of the scapulocoracoid; mes, mesopterygium; met, metapterygium; pro, propterygium; ro, rostral cartilage; scap, scapular process of the scapulocoracoid; ss, suprascapulae; syn, synarcual. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10932541" httpUri="https://zenodo.org/record/10932541/files/figure.png" pageId="7" pageNumber="1525">Fig. 5B</figureCitation>
|
||
). This specimen shows a small medially fused suprascapular cartilage; this cartilage is hourglass-shaped, with concave anterior and posterior borders, exhibiting deep indentations into which the distal edges of the scapular processes of the scapulocoracoid fit. Laterally, the scapulocoracoid articulates with the proximal portion of the pterygia through equidistant condyles. The propterygium is long and arched, tapers distally and extends to the anterior disc margin (
|
||
<figureCitation id="1317789BFFEAFFCC93CD6A1FB0DE20B1" box="[942,1011,1397,1420]" captionStart="Figure 4" captionStartId="8.[159,226,1157,1178]" captionTargetBox="[159,1460,188,1140]" captionTargetId="figure-295@8.[159,1467,188,1140]" captionTargetPageId="8" captionText="Figure 4. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGGC 7449, close-up of the head and pectoral girdle under UV light; B, reconstruction. Abbreviations: af, anterior fontanel; ao, antorbital cartilage; cb5, fifth ceratobranchial; hp, horn-like process; hyo, hyomandibula; mc, Meckel’s cartilage; mes, mesopterygium; met, metapterygium; nc, nasal capsule; pq, palatoquadrate; pro, propterygium; ro, rostral cartilage; rp, rostral process; sca, scapulocoracoid; syn, synarcual. Scale bars = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10932537" httpUri="https://zenodo.org/record/10932537/files/figure.png" pageId="7" pageNumber="1525">Fig. 4</figureCitation>
|
||
). The propterygium is segmented, with the first segment lying anterior to the mouth, close to the level of the antorbital cartilage. The proximal section of the propterygium does not extend far posteriorly to the procondyle, and does not articulate with the scapulocoracoid. A single unsegmented mesopterygium seems to be present. The metapterygium is as long and curved as the propterygium, but it is unclear whether it is segmented distally. The pectoral fins are clearly of the plesodic
|
||
<typeStatus id="5497DABCFFEAFFCC93BA69FCB7272390" box="[985,1034,1686,1709]" pageId="7" pageNumber="1525">type</typeStatus>
|
||
, with radials reaching the external border of the pectoral disc. All the radials articulate with the pterygia. Each pectoral radial contains 10–12 segments and bifurcates distally only once at about the eighth segment. There are approximately 81–87 pectoral radials, of which 35–38 are propterygial, 8–10 mesopterygial, and 38–41 metapterygial. The pectoral radials of †
|
||
<taxonomicName id="4C2C1F9DFFE5FFC390CF6A2DB26C2063" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[172,321,1351,1374]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="8" pageNumber="1526" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE5FFC390CF6A2DB26C2063" box="[172,321,1351,1374]" italics="true" pageId="8" pageNumber="1526">E. bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE5FFC3913F6A2DB2CA2063" box="[348,487,1351,1374]" pageId="8" pageNumber="1526" rank="species">comb. nov.</taxonomicNameLabel>
|
||
are robust, stiff and completely covered by mineralized tissue, forming the so-called ‘crustal calcification’ typical of most of batoids except the benthic stingrays and skates (
|
||
<bibRefCitation id="EFBD19EFFFE5FFC390C56AA1B2E320DF" author="Schaefer, J. T. & Summers, A. P." box="[166,462,1483,1506]" pageId="8" pageNumber="1526" pagination="298 - 313" refId="ref16848" refString="Schaefer, J. T. & Summers, A. P. 2005. Batoid wing skeletal structure: novel morphologies, mechanical implications, and phylogenetic patterns. Journal of Morphology, 264, 298 - 313. doi: 10.1002 / jmor. 10331" type="journal article" year="2005">Schaefer & Summers 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF533496FFE5FFC390FC6BEFB2412038" ID-DOI="http://doi.org/10.5281/zenodo.10932537" ID-Zenodo-Dep="10932537" httpUri="https://zenodo.org/record/10932537/files/figure.png" pageId="8" pageNumber="1526" startId="8.[159,226,1157,1178]" targetBox="[159,1460,188,1140]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B93641EFFE5FFC390FC6BEFB2412038" blockId="8.[159,1468,1157,1285]" pageId="8" pageNumber="1526">
|
||
<emphasis id="B958B80CFFE5FFC390FC6BEFB3D321A7" bold="true" box="[159,254,1157,1178]" pageId="8" pageNumber="1526">Figure 4.</emphasis>
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE5FFC391756BEFB1BF21A7" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[278,658,1157,1178]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="8" pageNumber="1526" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE5FFC391756BEFB2DB21A7" box="[278,502,1157,1178]" italics="true" pageId="8" pageNumber="1526">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE5FFC392656BEFB1A421A7" author="Heckel, M. J." box="[518,649,1157,1178]" pageId="8" pageNumber="1526" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE5FFC392FF6BEFB02121A7" box="[668,780,1157,1178]" pageId="8" pageNumber="1526" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site.
|
||
<emphasis id="B958B80CFFE5FFC3945F6BEFB76021A7" bold="true" box="[1084,1101,1157,1178]" pageId="8" pageNumber="1526">A</emphasis>
|
||
, MGGC 7449, close-up of the head and pectoral girdle under UV light;
|
||
<emphasis id="B958B80CFFE5FFC3927C6BCAB1022188" bold="true" box="[543,559,1184,1205]" pageId="8" pageNumber="1526">B</emphasis>
|
||
, reconstruction.
|
||
<emphasis id="B958B80CFFE5FFC392BE6BCAB0B02188" bold="true" box="[733,925,1184,1205]" pageId="8" pageNumber="1526">Abbreviations: af,</emphasis>
|
||
anterior fontanel;
|
||
<emphasis id="B958B80CFFE5FFC394016BCAB7AD2188" bold="true" box="[1122,1152,1184,1205]" pageId="8" pageNumber="1526">ao,</emphasis>
|
||
antorbital cartilage;
|
||
<emphasis id="B958B80CFFE5FFC395386BCAB6A92188" bold="true" box="[1371,1412,1184,1205]" pageId="8" pageNumber="1526">cb5,</emphasis>
|
||
fifth ceratobranchial;
|
||
<emphasis id="B958B80CFFE5FFC391216BD1B24F21ED" bold="true" box="[322,354,1211,1232]" pageId="8" pageNumber="1526">hp,</emphasis>
|
||
horn-like process;
|
||
<emphasis id="B958B80CFFE5FFC392456BD1B17C21ED" bold="true" box="[550,593,1211,1232]" pageId="8" pageNumber="1526">hyo,</emphasis>
|
||
hyomandibula;
|
||
<emphasis id="B958B80CFFE5FFC392976BD1B03421ED" bold="true" box="[756,793,1211,1232]" pageId="8" pageNumber="1526">mc,</emphasis>
|
||
Meckel’ s cartilage;
|
||
<emphasis id="B958B80CFFE5FFC3938A6BD1B73B21ED" bold="true" box="[1001,1046,1211,1232]" pageId="8" pageNumber="1526">mes,</emphasis>
|
||
mesopterygium;
|
||
<emphasis id="B958B80CFFE5FFC394A66BD1B7DF21ED" bold="true" box="[1221,1266,1211,1232]" pageId="8" pageNumber="1526">met,</emphasis>
|
||
metapterygium;
|
||
<emphasis id="B958B80CFFE5FFC395FE6BD1B69621ED" bold="true" box="[1437,1467,1211,1232]" pageId="8" pageNumber="1526">nc,</emphasis>
|
||
nasal capsule;
|
||
<emphasis id="B958B80CFFE5FFC391526BBFB27C21D7" bold="true" box="[305,337,1237,1258]" pageId="8" pageNumber="1526">pq,</emphasis>
|
||
palatoquadrate;
|
||
<emphasis id="B958B80CFFE5FFC391956BBFB10D21D7" bold="true" box="[502,544,1237,1258]" pageId="8" pageNumber="1526">pro,</emphasis>
|
||
propterygium;
|
||
<emphasis id="B958B80CFFE5FFC392D86BBFB1F521D7" bold="true" box="[699,728,1237,1258]" pageId="8" pageNumber="1526">ro,</emphasis>
|
||
rostral cartilage;
|
||
<emphasis id="B958B80CFFE5FFC393EA6BBFB08A21D7" bold="true" box="[905,935,1237,1258]" pageId="8" pageNumber="1526">rp,</emphasis>
|
||
rostral process;
|
||
<emphasis id="B958B80CFFE5FFC3942D6BBFB75921D7" bold="true" box="[1102,1140,1237,1258]" pageId="8" pageNumber="1526">sca,</emphasis>
|
||
scapulocoracoid;
|
||
<emphasis id="B958B80CFFE5FFC3954B6BBFB67D21D7" bold="true" box="[1320,1360,1237,1258]" pageId="8" pageNumber="1526">syn,</emphasis>
|
||
synarcual. Scale bars = 50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93641EFFE5FFC390DA6A81B0D323BC" blockId="8.[159,781,1350,1901]" lastBlockId="8.[845,1467,1351,1665]" pageId="8" pageNumber="1526">
|
||
The puboischiadic bar is partly recognizable in
|
||
<collectionCode id="ED3DFCDBFFE5FFC390FC6966B3DC231E" box="[159,241,1548,1571]" pageId="8" pageNumber="1526">MGGC</collectionCode>
|
||
7449/7450, where it seems straight or slightly bent, narrow and plate-like (
|
||
<figureCitation id="1317789BFFE5FFC391926947B11A2379" box="[497,567,1581,1604]" captionStart="Figure 6" captionStartId="9.[159,226,1661,1682]" captionTargetBox="[160,1466,1145,1643]" captionTargetId="figure-150@9.[159,1467,1144,1644]" captionTargetPageId="9" captionText="Figure 6. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, close-up of the pelvic girdle and fins in MGGC 7449; B, the same area under ultraviolet light; C, detail of one of the claspers. Abbreviations: ax, axial cartilage; cl, clasper; pf, pelvic fin radials; pub, puboischiadic bar; vc, vertebral centra. Scale bars: A, B = 50mm; C = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10932543" httpUri="https://zenodo.org/record/10932543/files/figure.png" pageId="8" pageNumber="1526">Fig. 6</figureCitation>
|
||
). It is difficult to recognize the postpelvic processes on the posterior margin of the puboischiadic bar that are typical for living platyrhinids. There are about 18–21 pelvic radials. The structure of the first pelvic radial is unclear but the pelvic condyles seem close together and not separated as in skates. All the specimens show straight and stout claspers, whose length represents about 10% TL (
|
||
<figureCitation id="1317789BFFE5FFC391CF685FB2D92271" box="[428,500,1845,1868]" captionStart="Figure 6" captionStartId="9.[159,226,1661,1682]" captionTargetBox="[160,1466,1145,1643]" captionTargetId="figure-150@9.[159,1467,1144,1644]" captionTargetPageId="9" captionText="Figure 6. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, close-up of the pelvic girdle and fins in MGGC 7449; B, the same area under ultraviolet light; C, detail of one of the claspers. Abbreviations: ax, axial cartilage; cl, clasper; pf, pelvic fin radials; pub, puboischiadic bar; vc, vertebral centra. Scale bars: A, B = 50mm; C = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10932543" httpUri="https://zenodo.org/record/10932543/files/figure.png" pageId="8" pageNumber="1526">Fig. 6</figureCitation>
|
||
). As in
|
||
<taxonomicName id="4C2C1F9DFFE5FFC39206685FB02B2271" authorityName="Garman" authorityYear="1881" box="[613,774,1845,1868]" class="Elasmobranchii" family="Rhinobatidae" genus="Platyrhinoidis" kingdom="Animalia" order="Rhinopristiformes" pageId="8" pageNumber="1526" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE5FFC39206685FB02B2271" box="[613,774,1845,1868]" italics="true" pageId="8" pageNumber="1526">Platyrhinoidis</emphasis>
|
||
</taxonomicName>
|
||
, their distal extremity does not reach the origin of the first dorsal fin; they differ from those characteristic of
|
||
<taxonomicName id="4C2C1F9DFFE5FFC393136A0DB0C52043" authorityName="Muller & Henle" authorityYear="1838" box="[880,1000,1383,1406]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="8" pageNumber="1526" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE5FFC393136A0DB0C52043" box="[880,1000,1383,1406]" italics="true" pageId="8" pageNumber="1526">Platyrhina</emphasis>
|
||
</taxonomicName>
|
||
, whose clasper tips can extend beyond the first dorsal-fin origin (e.g.
|
||
<bibRefCitation id="EFBD19EFFFE5FFC394CC6AEDB64820A3" author="Last, P. R. & White, W. & Carvalho, M. R. de & Stehmann, M. & Naylor, G. J. P." box="[1199,1381,1415,1438]" pageId="8" pageNumber="1526" refId="ref15487" refString="Last, P. R., White, W., Carvalho, M. R. de, S´eret, B., Stehmann, M. & Naylor, G. J. P. 2016. Rays of the world. CSIRO Publishing, Clayton North, 790 pp." type="book" year="2016">
|
||
Last
|
||
<emphasis id="B958B80CFFE5FFC3948A6AEDB60920A3" box="[1257,1316,1415,1438]" italics="true" pageId="8" pageNumber="1526">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD19EFFFE5FFC395146AEDB0F22082" author="White, W. T. & Last, P. R." pageId="8" pageNumber="1526" pagination="533 - 544" refId="ref17323" refString="White, W. T. & Last, P. R. 2016. Platyrhina psomadakisi sp. nov., a new species of fanray (Batoidea: Platyrhinidae) from the Andaman Sea, the first record of this family in the Indian Ocean. Zootaxa, 4121, 533 - 544. doi: 10.11646 / zootaxa. 4121.5.3" type="journal article" year="2016">White & Last 2016</bibRefCitation>
|
||
). The clasper glands are almost entirely covered by dermal denticles, and consequently their skeletal morphology is difficult to describe. However, the axial cartilage is rod-like, possibly calcified over most of its length, and extends and inserts over the complete length of the clasper to the ventral terminal cartilage.
|
||
</paragraph>
|
||
<paragraph id="8B93641EFFE5FFC2932E69FEB7122260" blockId="8.[845,1467,1684,1901]" lastBlockId="9.[845,1467,1797,1885]" lastPageId="9" lastPageNumber="1527" pageId="8" pageNumber="1526">
|
||
<emphasis id="B958B80CFFE5FFC3932E69FEB7732396" bold="true" box="[845,1118,1684,1707]" pageId="8" pageNumber="1526">Dorsal and caudal fins.</emphasis>
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE5FFC3941769FEB6432396" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[1140,1390,1684,1707]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="8" pageNumber="1526" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE5FFC3941769FEB6432396" box="[1140,1390,1684,1707]" italics="true" pageId="8" pageNumber="1526">Eoplatyrhina bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE5FFC3951A69FEB05123F1" pageId="8" pageNumber="1526" rank="species">comb. nov.</taxonomicNameLabel>
|
||
possesses two dorsal fins located in the posterior half of the tail. The extent of the fin radial cartilages into the fin web is not precisely ascertainable, but they are possibly aplesodic. The base of the dorsal fins has a length of about 5% TL. No impression of dorsal-fin radials is visible. The caudal fin is only preserved in
|
||
<collectionCode id="ED3DFCDBFFE4FFC290FC686CB2292220" box="[159,260,1798,1821]" pageId="9" pageNumber="1527">MGP-PD</collectionCode>
|
||
26279C/80C (
|
||
<figureCitation id="1317789BFFE4FFC291D2686CB1242220" box="[433,521,1798,1821]" captionStart="Figure 5" captionStartId="9.[159,226,951,972]" captionTargetBox="[159,1467,188,933]" captionTargetId="figure-57@9.[159,1467,188,933]" captionTargetPageId="9" captionText="Figure 5. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, MGP-PD 26279C, close-up of the rostral cartilage; B, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula; C, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin. Abbreviations: af, anterior fontanel; co, coracoid bar of the scapulocoracoid; mes, mesopterygium; met, metapterygium; pro, propterygium; ro, rostral cartilage; scap, scapular process of the scapulocoracoid; ss, suprascapulae; syn, synarcual. Scale bars = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10932541" httpUri="https://zenodo.org/record/10932541/files/figure.png" pageId="9" pageNumber="1527">Fig. 5C</figureCitation>
|
||
). It is about 11% TL and contains about 23 vertebrae not reaching the posterior-most border of the caudal fin. There are about 20–25 caudal-fin radials on the ventral and dorsal sides (
|
||
<quantity id="4CD4C9FBFFE4FFC29515686CB04F2200" metricMagnitude="0" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="9" pageNumber="1527" unit="in" value="45.0" valueMax="50.0" valueMin="40.0">40–50 in</quantity>
|
||
total), which do not reach the external margin of the caudal fin (aplesodic).
|
||
</paragraph>
|
||
<caption id="DF533496FFE4FFC290FC6CDDB04C210B" ID-DOI="http://doi.org/10.5281/zenodo.10932541" ID-Zenodo-Dep="10932541" httpUri="https://zenodo.org/record/10932541/files/figure.png" pageId="9" pageNumber="1527" startId="9.[159,226,951,972]" targetBox="[159,1467,188,933]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B93641EFFE4FFC290FC6CDDB04C210B" blockId="9.[159,1466,950,1079]" pageId="9" pageNumber="1527">
|
||
<emphasis id="B958B80CFFE4FFC290FC6CDDB3D226F1" bold="true" box="[159,255,951,972]" pageId="9" pageNumber="1527">Figure 5.</emphasis>
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE4FFC291756CDDB1B926F1" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[278,660,951,972]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="9" pageNumber="1527" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE4FFC291756CDDB2DB26F1" box="[278,502,951,972]" italics="true" pageId="9" pageNumber="1527">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE4FFC292646CDDB1A126F1" author="Heckel, M. J." box="[519,652,951,972]" pageId="9" pageNumber="1527" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE4FFC292FC6CDDB02226F1" box="[671,783,951,972]" pageId="9" pageNumber="1527" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site.
|
||
<emphasis id="B958B80CFFE4FFC294216CDDB77E26F1" bold="true" box="[1090,1107,951,972]" pageId="9" pageNumber="1527">A</emphasis>
|
||
, MGP-PD 26279C, close-up of the rostral cartilage;
|
||
<emphasis id="B958B80CFFE4FFC291276CBBB27926DB" bold="true" box="[324,340,977,998]" pageId="9" pageNumber="1527">B</emphasis>
|
||
, close-up of the pectoral girdle (dorsal view) showing the position of the suprascapula;
|
||
<emphasis id="B958B80CFFE4FFC294D86CBBB7E126DB" bold="true" box="[1211,1228,977,998]" pageId="9" pageNumber="1527">C</emphasis>
|
||
, caudal fin of MGP-PD 26279C; the dashed line shows the original genuine outline of the fin.
|
||
<emphasis id="B958B80CFFE4FFC293006C86B732213C" bold="true" box="[867,1055,1004,1025]" pageId="9" pageNumber="1527">Abbreviations: af,</emphasis>
|
||
anterior fontanel;
|
||
<emphasis id="B958B80CFFE4FFC294BB6C86B7D8213C" bold="true" box="[1240,1269,1004,1025]" pageId="9" pageNumber="1527">co,</emphasis>
|
||
coracoid bar of the scapulocoracoid;
|
||
<emphasis id="B958B80CFFE4FFC291296B6DB2552121" bold="true" box="[330,376,1031,1052]" pageId="9" pageNumber="1527">mes,</emphasis>
|
||
mesopterygium;
|
||
<emphasis id="B958B80CFFE4FFC292446B6DB1792121" bold="true" box="[551,596,1031,1052]" pageId="9" pageNumber="1527">met,</emphasis>
|
||
metapterygium;
|
||
<emphasis id="B958B80CFFE4FFC2929C6B6DB0042121" bold="true" box="[767,809,1031,1052]" pageId="9" pageNumber="1527">pro,</emphasis>
|
||
propterygium;
|
||
<emphasis id="B958B80CFFE4FFC293A66B6DB0CF2121" bold="true" box="[965,994,1031,1052]" pageId="9" pageNumber="1527">ro,</emphasis>
|
||
rostral cartilage;
|
||
<emphasis id="B958B80CFFE4FFC294F56B6DB7E42121" bold="true" box="[1174,1225,1031,1052]" pageId="9" pageNumber="1527">scap,</emphasis>
|
||
scapular process of the scapulocoracoid;
|
||
<emphasis id="B958B80CFFE4FFC291246B4BB24D210B" bold="true" box="[327,352,1057,1078]" pageId="9" pageNumber="1527">ss,</emphasis>
|
||
suprascapulae;
|
||
<emphasis id="B958B80CFFE4FFC291986B4BB109210B" bold="true" box="[507,548,1057,1078]" pageId="9" pageNumber="1527">syn,</emphasis>
|
||
synarcual. Scale bars = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF533496FFE4FFC290FC6917B62A23FA" ID-DOI="http://doi.org/10.5281/zenodo.10932543" ID-Zenodo-Dep="10932543" httpUri="https://zenodo.org/record/10932543/files/figure.png" pageId="9" pageNumber="1527" startId="9.[159,226,1661,1682]" targetBox="[160,1466,1145,1643]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B93641EFFE4FFC290FC6917B62A23FA" blockId="9.[159,1467,1660,1736]" pageId="9" pageNumber="1527">
|
||
<emphasis id="B958B80CFFE4FFC290FC6917B3D023AF" bold="true" box="[159,253,1661,1682]" pageId="9" pageNumber="1527">Figure 6.</emphasis>
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE4FFC291766917B1A023AF" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[277,653,1661,1682]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="9" pageNumber="1527" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE4FFC291766917B2DE23AF" box="[277,499,1661,1682]" italics="true" pageId="9" pageNumber="1527">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE4FFC292616917B1A823AF" author="Heckel, M. J." box="[514,645,1661,1682]" pageId="9" pageNumber="1527" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE4FFC292F56917B02923AF" box="[662,772,1661,1682]" pageId="9" pageNumber="1527" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site.
|
||
<emphasis id="B958B80CFFE4FFC2944E6917B71323AF" bold="true" box="[1069,1086,1661,1682]" pageId="9" pageNumber="1527">A</emphasis>
|
||
, close-up of the pelvic girdle and fins in MGGC 7449;
|
||
<emphasis id="B958B80CFFE4FFC2912C69FDB2722391" bold="true" box="[335,351,1687,1708]" pageId="9" pageNumber="1527">B</emphasis>
|
||
, the same area under ultraviolet light;
|
||
<emphasis id="B958B80CFFE4FFC2928D69FDB1D22391" bold="true" box="[750,767,1687,1708]" pageId="9" pageNumber="1527">C</emphasis>
|
||
, detail of one of the claspers.
|
||
<emphasis id="B958B80CFFE4FFC2945E69FDB7D32391" bold="true" box="[1085,1278,1687,1708]" pageId="9" pageNumber="1527">Abbreviations: ax,</emphasis>
|
||
axial cartilage;
|
||
<emphasis id="B958B80CFFE4FFC295C069FDB6962391" bold="true" box="[1443,1467,1687,1708]" pageId="9" pageNumber="1527">cl,</emphasis>
|
||
clasper;
|
||
<emphasis id="B958B80CFFE4FFC2909369D8B22623FA" bold="true" box="[240,267,1714,1735]" pageId="9" pageNumber="1527">pf,</emphasis>
|
||
pelvic fin radials;
|
||
<emphasis id="B958B80CFFE4FFC291A669D8B2DE23FA" bold="true" box="[453,499,1714,1735]" pageId="9" pageNumber="1527">pub,</emphasis>
|
||
puboischiadic bar;
|
||
<emphasis id="B958B80CFFE4FFC292D069D8B1FD23FA" bold="true" box="[691,720,1714,1735]" pageId="9" pageNumber="1527">vc,</emphasis>
|
||
vertebral centra. Scale bars: A, B = 50 mm; C = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF533496FFE7FFC190FC6C02B66B26AA" ID-DOI="http://doi.org/10.5281/zenodo.10932545" ID-Zenodo-Dep="10932545" httpUri="https://zenodo.org/record/10932545/files/figure.png" pageId="10" pageNumber="1528" startId="10.[159,226,872,893]" targetBox="[159,1446,204,854]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8B93641EFFE7FFC190FC6C02B66B26AA" blockId="10.[159,1467,871,920]" pageId="10" pageNumber="1528">
|
||
<emphasis id="B958B80CFFE7FFC190FC6C02B22D2640" bold="true" box="[159,256,872,893]" pageId="10" pageNumber="1528">Figure 7.</emphasis>
|
||
†
|
||
<taxonomicName id="4C2C1F9DFFE7FFC191746C02B1BA2640" authority="(Heckel, 1851)" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[279,663,872,893]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE7FFC191746C02B2D42640" box="[279,505,872,893]" italics="true" pageId="10" pageNumber="1528">Eoplatyrhina bolcensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD19EFFFE7FFC192696C02B1A22640" author="Heckel, M. J." box="[522,655,872,893]" pageId="10" pageNumber="1528" pagination="281 - 333" refId="ref15152" refString="Heckel, M. J. 1851. Bericht einer auf Kosten der kais. Akademie der Wissenschaften durch Oberosterreich nach Salzburg, Munchen, Innsbruck, Botzen, Verona, Padua, Venedig und Triest unternommenen Reise. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Klasse, 7, 281 - 333." type="journal article" year="1851">Heckel, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE7FFC192C06C02B03E2640" box="[675,787,872,893]" pageId="10" pageNumber="1528" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Monte Postale site.
|
||
<emphasis id="B958B80CFFE7FFC194286C02B7712640" bold="true" box="[1099,1116,872,893]" pageId="10" pageNumber="1528">A</emphasis>
|
||
, dermal denticles from the tail of MGP-PD 26279C (dorsal view);
|
||
<emphasis id="B958B80CFFE7FFC191816CE8B2DF26AA" bold="true" box="[482,498,898,919]" pageId="10" pageNumber="1528">B</emphasis>
|
||
, dermal denticles from the tail of MGP-PD 26279C (basal view). Scale bar = 400 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93641EFFE7FFC190FC6CB3B2962379" blockId="10.[159,781,985,1604]" pageId="10" pageNumber="1528">
|
||
<emphasis id="B958B80CFFE7FFC190FC6CB3B25A26CD" bold="true" box="[159,375,985,1008]" pageId="10" pageNumber="1528">Dermal denticles.</emphasis>
|
||
As in extant platyrhinids (see
|
||
<bibRefCitation id="EFBD19EFFFE7FFC190FC6C90B21D212C" author="Deynat, P. P." box="[159,304,1018,1041]" pageId="10" pageNumber="1528" pagination="75 - 84" refId="ref14675" refString="Deynat, P. P. 2005. Characteristics of the dermal covering in Platyrhinidae (Chondrichthyes, Rhinobatiformes). Biociencias, 13, 75 - 84." type="journal article" year="2005">Deynat 2005</bibRefCitation>
|
||
), the entire body of †
|
||
<taxonomicName id="4C2C1F9DFFE7FFC192506C90B193212C" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[563,702,1018,1041]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE7FFC192506C90B193212C" box="[563,702,1018,1041]" italics="true" pageId="10" pageNumber="1528">E. bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE7FFC192AB6C90B3FD210F" pageId="10" pageNumber="1528" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is covered with numerous small dermal denticles that form a continuous and regular covering (
|
||
<figureCitation id="1317789BFFE7FFC192D46B56B1D1216E" box="[695,764,1084,1107]" captionStart="Figure 7" captionStartId="10.[159,226,872,893]" captionTargetBox="[159,1446,204,854]" captionTargetId="figure-443@10.[159,1467,188,854]" captionTargetPageId="10" captionText="Figure 7. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, dermal denticles from the tail of MGP-PD 26279C (dorsal view); B, dermal denticles from the tail of MGP-PD 26279C (basal view). Scale bar = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.10932545" httpUri="https://zenodo.org/record/10932545/files/figure.png" pageId="10" pageNumber="1528">Fig. 7</figureCitation>
|
||
). Denticle size is quite uniform across the body. Some denticles were extracted from the dorsal side of the disc of
|
||
<collectionCode id="ED3DFCDBFFE7FFC1909A6BCAB24C218A" box="[249,353,1184,1207]" pageId="10" pageNumber="1528">MGP-PD</collectionCode>
|
||
26279C/80C for a detailed analysis. Their crown is about 200 µm wide and rhomboidal or lozenge-shaped (
|
||
<figureCitation id="1317789BFFE7FFC191076B88B28721C4" box="[356,426,1250,1273]" captionStart="Figure 7" captionStartId="10.[159,226,872,893]" captionTargetBox="[159,1446,204,854]" captionTargetId="figure-443@10.[159,1467,188,854]" captionTargetPageId="10" captionText="Figure 7. †Eoplatyrhina bolcensis (Heckel, 1851) comb. nov. from the Monte Postale site. A, dermal denticles from the tail of MGP-PD 26279C (dorsal view); B, dermal denticles from the tail of MGP-PD 26279C (basal view). Scale bar = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.10932545" httpUri="https://zenodo.org/record/10932545/files/figure.png" pageId="10" pageNumber="1528">Fig. 7</figureCitation>
|
||
). The denticle root is deeper than the crown height and a nutritive foramen can be recognized near the centre. Extant thornbacks and †
|
||
<taxonomicName id="4C2C1F9DFFE7FFC190CF6A2FB2012061" authorityName="Claeson, Underwood & Ward" authorityYear="2013" box="[172,300,1349,1372]" class="Elasmobranchii" family="Rhinobatidae" genus="Tingitanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE7FFC190CF6A2FB2012061" box="[172,300,1349,1372]" italics="true" pageId="10" pageNumber="1528">Tingitanius</emphasis>
|
||
</taxonomicName>
|
||
possess parallel rows of enlarged dermal denticles (thorns) over the posterior part of the disc and tail, a condition that was regarded as diagnostic for platyrhinids. However, this is not the case for †
|
||
<taxonomicName id="4C2C1F9DFFE7FFC190CF6AA0B29920DC" authorityName="Marram̀ & Carneval & Claeso & Naylo & Kriwe" authorityYear="2020" baseAuthorityName="Heckel" baseAuthorityYear="1851" box="[172,436,1482,1505]" class="Elasmobranchii" family="Platyrhinidae" genus="Eoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="bolcensis" status="comb. nov.">
|
||
<emphasis id="B958B80CFFE7FFC190CF6AA0B29920DC" box="[172,436,1482,1505]" italics="true" pageId="10" pageNumber="1528">Eoplatyrhina bolcensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B0577FFE7FFC191A26AA0B16E20DC" box="[449,579,1482,1505]" pageId="10" pageNumber="1528" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and †
|
||
<taxonomicName id="4C2C1F9DFFE7FFC192F26AA0B02B20DC" authorityName="de Carvalho" authorityYear="2004" box="[657,774,1482,1505]" class="Elasmobranchii" family="Rhinobatidae" genus="Tethybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhinopristiformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958B80CFFE7FFC192F26AA0B02B20DC" box="[657,774,1482,1505]" italics="true" pageId="10" pageNumber="1528">Tethybatis</emphasis>
|
||
</taxonomicName>
|
||
, in which thorns are completely absent (
|
||
<bibRefCitation id="EFBD19EFFFE7FFC192C66A81B3F6231E" author="Carvalho, M. R. de" pageId="10" pageNumber="1528" refId="ref14177" refString="Carvalho, M. R. de. 2004. A Late Cretaceous thornback ray from southern Italy, with a phylogenetic reappraisal of the Platyrhinidae (Chondrichthyes: Batoidea). Pp. 75 - 100 in G. Arratia & A. Tintori (eds) Mesozoic fishes 3 - Systematics, paleoenvironments and biodiversity. Verlag Dr Friedrich Pfeil, Munich." type="book" year="2004">Carvalho 2004</bibRefCitation>
|
||
), possibly representing a feature supporting this sister-group relationship.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |