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<mods:title id="C79AF9E8D5871D78ADD5BB4103D01E0B">Revision of the Eocene Platyrhina species from the Bolca Lagerst atte (Italy) reveals the first panray (Batomorphii: Zanobatidae) in the fossil record</mods:title>
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<mods:namePart id="C6395B1BDA2A7B9D6951C20C380BFFFF">Marram ̀, Giuseppe</mods:namePart>
<mods:affiliation id="43DBEE3D362E537DE415267F861FD340">Universita degli Studi di Torino, Dipartimento di Scienze della Terra, via Valperga Caluso 35, 10125 Torino, Italy; &amp; University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:namePart id="5AF5F0497DF9BB6E1B65F72DACB356A3">Carneval, Giorgio</mods:namePart>
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<mods:namePart id="737031D67554C211A9E72BC450F908F6">Claeso, Kerin M.</mods:namePart>
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<mods:namePart id="60976F322FC6B05BFFFE82471F1942F3">Naylo, Gavin J. P.</mods:namePart>
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<subSubSection id="C3363795FFE7FFC1917A69F9B1BE2397" box="[281,659,1682,1706]" pageId="10" pageNumber="1528" type="nomenclature">
<paragraph id="8B93641EFFE7FFC1917A69F9B1BE2397" blockId="10.[279,660,1648,1706]" box="[281,659,1682,1706]" pageId="10" pageNumber="1528">
<heading id="D0DBD372FFE7FFC1917A69F9B1BE2397" box="[281,659,1682,1706]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="1528" reason="8">
Genus †
<taxonomicName id="4C2C1F9DFFE7FFC1911069F8B1082397" authority="Marram̀ &amp; Carneval &amp; Claeso &amp; Naylo &amp; Kriwe, 2020" authorityName="Marram̀ &amp; Carneval &amp; Claeso &amp; Naylo &amp; Kriwe" authorityYear="2020" box="[371,549,1682,1706]" class="Elasmobranchii" family="Zanobatidae" genus="Plesiozanobatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B958B80CFFE7FFC1911069F8B1082397" bold="true" box="[371,549,1682,1706]" italics="true" pageId="10" pageNumber="1528">Plesiozanobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B0577FFE7FFC1924D69F9B1BE2397" box="[558,659,1683,1706]" pageId="10" pageNumber="1528" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="8B93641EFFE7FFC190FC69A9B1E823E7" blockId="10.[159,709,1730,1754]" box="[159,709,1730,1754]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC190FC69A9B21A23E7" bold="true" box="[159,311,1731,1754]" pageId="10" pageNumber="1528">Type species.</emphasis>
<taxonomicName id="4C2C1F9DFFE7FFC1912E69A9B19323E7" authority="De Zigno, 1876" authorityName=", De Zigno" authorityYear="1876" box="[333,702,1731,1754]" class="Elasmobranchii" family="Torpedinidae" genus="Torpedo" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="egertoni">
<emphasis id="B958B80CFFE7FFC1912E69A9B12423E7" box="[333,521,1731,1754]" italics="true" pageId="10" pageNumber="1528">Torpedo egertoni</emphasis>
<bibRefCitation id="EFBD19EFFFE7FFC1927169A9B19323E7" author="De Zigno, A." box="[530,702,1731,1754]" pageId="10" pageNumber="1528" pagination="445 - 455" refId="ref14740" refString="De Zigno, A. 1876. Annotazioni paleontologiche. Aggiunte alla ittiologia dell' epoca Eocena. Memorie del Reale Istituto Veneto di Science, Lettere ed Arti, 20, 445 - 455." type="journal article" year="1876">De Zigno, 1876</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<paragraph id="8B93641EFFE7FFC190FC6999B61621E9" blockId="10.[159,781,1779,1901]" lastBlockId="10.[844,1467,985,1236]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC190FC6999B2392237" bold="true" box="[159,276,1779,1802]" pageId="10" pageNumber="1528">Diagnosis.</emphasis>
Pectoral disc large and roughly rounded, representing 5670% TL; tail stout and short, distinctly demarcated from the disc; two dorsal fins and caudal fin present; densely, closely set small dermal denticles forming a continuous pavement; large, rounded, scattered thorns covering the entire disc and tail; rostral cartilage absent; nasal capsules laterally expanded without horn-like processes; long propterygia extending near the anterior margin of the disc; mesopterygium absent; about 6575 pectoral radials; puboischiadic bar narrow and moderately arched; approximately 20 pelvic-fin rays; 8090 vertebrae; about 10 pairs of ribs.
</paragraph>
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<subSubSection id="C3363795FFE7FFC1932F6B87B779205B" pageId="10" pageNumber="1528" type="etymology">
<paragraph id="8B93641EFFE7FFC1932F6B87B779205B" blockId="10.[844,1467,1261,1382]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC1932F6B87B7682039" bold="true" box="[844,1093,1261,1284]" pageId="10" pageNumber="1528">Derivation of name.</emphasis>
From the Ancient Greek word
<emphasis id="B958B80CFFE7FFC1932F6A65B0A12018" bold="true" box="[844,908,1293,1317]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC1932F6A65B0A92018" bold="true" box="[844,900,1295,1317]" italics="true" pageId="10" pageNumber="1528">pλGR</emphasis>
ί
</emphasis>
o
<emphasis id="B958B80CFFE7FFC193FF6A64B0842018" box="[924,937,1294,1317]" italics="true" pageId="10" pageNumber="1528">u</emphasis>
(
<emphasis id="B958B80CFFE7FFC193A96A64B7352018" box="[970,1048,1293,1317]" italics="true" pageId="10" pageNumber="1528">pl̄eśıon</emphasis>
) meaning near or close, and
<taxonomicName id="4C2C1F9DFFE7FFC1932F6A44B0902078" authorityName="Garman" authorityYear="1913" box="[844,957,1326,1349]" class="Elasmobranchii" family="Rhinobatidae" genus="Zanobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus">
<emphasis id="B958B80CFFE7FFC1932F6A44B0902078" box="[844,957,1326,1349]" italics="true" pageId="10" pageNumber="1528">Zanobatus</emphasis>
</taxonomicName>
, to remark upon its close relationship with the living panray genus.
</paragraph>
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<paragraph id="8B93641EFFE7FFC1932F6A15B7C820AB" blockId="10.[844,1253,1407,1430]" box="[844,1253,1407,1430]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC1932F6A15B73D20AB" bold="true" box="[844,1040,1407,1430]" pageId="10" pageNumber="1528">Included species.</emphasis>
<typeStatus id="5497DABCFFE7FFC1947B6A15B76220AB" box="[1048,1103,1407,1430]" pageId="10" pageNumber="1528">Type</typeStatus>
species only.
</paragraph>
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<subSubSection id="C3363795FFE7FFC1932F6AC5B73D2250" pageId="10" pageNumber="1528" type="discussion">
<paragraph id="8B93641EFFE7FFC1932F6AC5B73D2250" blockId="10.[844,1467,1455,1901]" pageId="10" pageNumber="1528">
<emphasis id="B958B80CFFE7FFC1932F6AC5B09620FB" bold="true" box="[844,955,1455,1478]" pageId="10" pageNumber="1528">Remarks.</emphasis>
<bibRefCitation id="EFBD19EFFFE7FFC193A76AC5B7AE20FB" author="De Zigno, A." box="[964,1155,1455,1478]" pageId="10" pageNumber="1528" pagination="445 - 455" refId="ref14740" refString="De Zigno, A. 1876. Annotazioni paleontologiche. Aggiunte alla ittiologia dell' epoca Eocena. Memorie del Reale Istituto Veneto di Science, Lettere ed Arti, 20, 445 - 455." type="journal article" year="1876">De Zigno (1876)</bibRefCitation>
considered that the overall similarity of the disc shape and the absence of a tail sting on the holotypic specimen
<collectionCode id="ED3DFCDBFFE7FFC194D06A9AB635233A" box="[1203,1304,1520,1543]" pageId="10" pageNumber="1528">MGP-PD</collectionCode>
154Z justified the assignment of this species to the genus
<taxonomicName id="4C2C1F9DFFE7FFC1953A697AB69E231A" box="[1369,1459,1552,1575]" class="Elasmobranchii" family="Torpedinidae" genus="Torpedo" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus">
<emphasis id="B958B80CFFE7FFC1953A697AB69E231A" box="[1369,1459,1552,1575]" italics="true" pageId="10" pageNumber="1528">Torpedo</emphasis>
</taxonomicName>
. Later,
<bibRefCitation id="EFBD19EFFFE7FFC193F8695BB71B2375" author="Jaekel, O." box="[923,1078,1585,1608]" pageId="10" pageNumber="1528" refId="ref15363" refString="Jaekel, O. 1894. Die eocanen Selachier vom Monte Bolca: ein Beitrag zur Morphogenie der Wirbelthiere. J. Springer, Berlin, 176 pp." type="book" year="1894">Jaekel (1894)</bibRefCitation>
, analysing additional, better preserved material, assigned the species †
<taxonomicName id="4C2C1F9DFFE7FFC195746938B6BB2354" authorityName=", De Zigno" authorityYear="1876" box="[1303,1430,1618,1641]" class="Elasmobranchii" family="Torpedinidae" genus="Torpedo" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="egertoni">
<emphasis id="B958B80CFFE7FFC195746938B6BB2354" box="[1303,1430,1618,1641]" italics="true" pageId="10" pageNumber="1528">T. egertoni</emphasis>
</taxonomicName>
to
<taxonomicName id="4C2C1F9DFFE7FFC1932F6918B09323B4" authorityName="Muller &amp; Henle" authorityYear="1838" box="[844,958,1650,1673]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="genus">
<emphasis id="B958B80CFFE7FFC1932F6918B09323B4" box="[844,958,1650,1673]" italics="true" pageId="10" pageNumber="1528">Platyrhina</emphasis>
</taxonomicName>
. However, he noticed that the fossil species from Bolca might have been more closely related to
<taxonomicName id="4C2C1F9DFFE7FFC1932F69D9B77F23F7" authorityName="Muller &amp; Henle" authorityYear="1841" box="[844,1106,1715,1738]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="schoenleinii">
<emphasis id="B958B80CFFE7FFC1932F69D9B77F23F7" box="[844,1106,1715,1738]" italics="true" pageId="10" pageNumber="1528">Platyrhina schoenleinii</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C2C1F9DFFE7FFC194A869D9B69623F7" box="[1227,1467,1715,1738]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="sinesensis">
<emphasis id="B958B80CFFE7FFC194A869D9B69623F7" box="[1227,1467,1715,1738]" italics="true" pageId="10" pageNumber="1528">Platyrhina sinesensis</emphasis>
</taxonomicName>
because of the general shape and proportions of the body and disc, as well as the arrangement of the pectoral radials and gill arches.
<taxonomicName id="4C2C1F9DFFE7FFC194F8687FB6B62211" authorityName="Muller &amp; Henle" authorityYear="1841" box="[1179,1435,1813,1836]" class="Elasmobranchii" family="Platyrhinidae" genus="Platyrhina" kingdom="Animalia" order="Torpediniformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="schoenleinii">
<emphasis id="B958B80CFFE7FFC194F8687FB6B62211" box="[1179,1435,1813,1836]" italics="true" pageId="10" pageNumber="1528">Platyrhina schoenleinii</emphasis>
</taxonomicName>
is currently recognized as
<taxonomicName id="4C2C1F9DFFE7FFC19416685CB6562270" baseAuthorityName="Muller &amp; Henle" baseAuthorityYear="1841" box="[1141,1403,1846,1869]" class="Elasmobranchii" family="Rhinobatidae" genus="Zanobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="10" pageNumber="1528" phylum="Chordata" rank="species" species="schoenleinii">
<emphasis id="B958B80CFFE7FFC19416685CB6562270" box="[1141,1403,1846,1869]" italics="true" pageId="10" pageNumber="1528">Zanobatus schoenleinii</emphasis>
</taxonomicName>
(see Compagno 1999).
</paragraph>
</subSubSection>
</treatment>
</document>