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<document id="B4502AAEED730DDFEA6633AE86AF6E64" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago &amp; Vaz-De, Fernando Z." docDate="2021" docId="03850669FFBBFFE0FE0123643892C881" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) subaeneus" docType="treatment" docVersion="8" lastPageNumber="47" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="44" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="06D7BC9E5218030F802D1FAECBCA07F3">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="801AD7F62911ADC65EC2C8C28351F49E">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="1357C8136D64BD60939ED8ED72024766">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="57EFD646CFCCD6B65C4DC623D114DE52" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="424339D7835ED033B5F71F6F99DDFF7F">Vaz-De, Fernando Z.</mods:namePart>
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<mods:title id="4BB824828341709D8E7AFAAC05203AF8">European Journal of Taxonomy</mods:title>
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<treatment id="03850669FFBBFFE0FE0123643892C881" ID-DOI="http://doi.org/10.5281/zenodo.4545861" ID-GBIF-Taxon="177420804" ID-Zenodo-Dep="4545861" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881" httpUri="http://treatment.plazi.org/id/03850669FFBBFFE0FE0123643892C881" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44">
<subSubSection id="C336E4F4FFBBFFE5FE012364382ECE0B" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8B93B77FFFBBFFE5FE012364382ECE0B" blockId="43.[428,1159,1313,1373]" pageId="43" pageNumber="44">
<taxonomicName id="4C2CCCFCFFBBFFE5FE0123643FDECE6A" ID-CoL="7ZN5G" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[428,1159,1313,1340]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus" subGenus="Luederwaldtinia">
<emphasis id="B9586B6DFFBBFFE5FE012364391ECE6A" bold="true" box="[428,583,1313,1340]" italics="true" pageId="43" pageNumber="44">Dichotomius</emphasis>
(
<emphasis id="B9586B6DFFBBFFE5FDF523643846CE6A" bold="true" box="[600,799,1313,1340]" italics="true" pageId="43" pageNumber="44">Luederwaldtinia</emphasis>
)
<emphasis id="B9586B6DFFBBFFE5FC9D236438F6CE6A" bold="true" box="[816,943,1313,1340]" italics="true" pageId="43" pageNumber="44">subaeneus</emphasis>
(
<bibRefCitation id="EFBDCA8EFFBBFFE5FC1223643F26CE6A" author="Castelnau M." box="[959,1151,1313,1340]" pageId="43" pageNumber="44" pagination="1 - 564" refId="ref29645" refString="Castelnau M. 1840. Histoire naturelle des coleopteres. In: Blanchard E. (eds) Histoire naturelle des animaux articules. Vol. 2: 1 - 564. Dumenil, Paris." type="journal article" year="1840">Castelnau, 1840</bibRefCitation>
)
</taxonomicName>
<figureCitation id="1317ABFAFFBBFFE5FD1123073849CE0B" box="[700,784,1346,1373]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="43" pageNumber="44">Fig. 15</figureCitation>
, Map 13
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBBFFE5FF1023C739EECEAA" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8B93B77FFFBBFFE5FF1023C73825CECB" blockId="43.[189,892,1410,1437]" box="[189,892,1410,1437]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB3C9051FFBBFFE5FF1023C73825CECB" box="[189,892,1410,1437]" pageId="43" pageNumber="44">
<treatmentCitation id="0A8D916EFFBBFFE5FF1023C7392DCECA" author="Castelnau M." box="[189,628,1410,1436]" page="79" pageId="43" pageNumber="44" year="1840">
<taxonomicName id="4C2CCCFCFFBBFFE5FF1023C7392DCECA" ID-CoL="7VDMN" authority="Castelnau, 1840: 79" authorityName="Castelnau" authorityPageNumber="79" authorityYear="1840" box="[189,628,1410,1436]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaenea">
<emphasis id="B9586B6DFFBBFFE5FF1023C73ADACECA" box="[189,387,1410,1436]" italics="true" pageId="43" pageNumber="44">Copris subaenea</emphasis>
<bibRefCitation id="EFBDCA8EFFBBFFE5FE2723C7392DCECA" author="Castelnau M." box="[394,628,1410,1436]" pageId="43" pageNumber="44" pagination="1 - 564" refId="ref29645" refString="Castelnau M. 1840. Histoire naturelle des coleopteres. In: Blanchard E. (eds) Histoire naturelle des animaux articules. Vol. 2: 1 - 564. Dumenil, Paris." type="journal article" year="1840">Castelnau, 1840: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFFBBFFE5FF1023873F4DCE8D" blockId="43.[189,1044,1473,1533]" box="[189,1044,1473,1500]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB3C9051FFBBFFE5FF1023873F4DCE8D" box="[189,1044,1473,1500]" pageId="43" pageNumber="44">
<taxonomicName id="4C2CCCFCFFBBFFE5FF1023873AC1CE8D" box="[189,408,1473,1499]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBBFFE5FF1023873AC1CE8D" box="[189,408,1473,1499]" italics="true" pageId="43" pageNumber="44">Pinotus subaeneus</emphasis>
</taxonomicName>
<treatmentCitation id="0A8D916EFFBBFFE5FE1823843993CE8D" author="Luederwaldt H." box="[437,714,1473,1499]" page="136" pageId="43" pageNumber="44" year="1929">
<bibRefCitation id="EFBDCA8EFFBBFFE5FE1823843993CE8D" author="Luederwaldt H." box="[437,714,1473,1499]" pageId="43" pageNumber="44" pagination="1 - 173" refId="ref30128" refString="Luederwaldt H. 1929. As especies Brasileiras do genero Pinotus. Revista do Museu Paulista 16: 1 - 173." type="journal article" year="1929">Luederwaldt 1929: 136</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A8D916EFFBBFFE5FD5623843F49CE8D" author="Blackwelder R. E." box="[763,1040,1473,1499]" page="208" pageId="43" pageNumber="44" year="1944">
<bibRefCitation id="EFBDCA8EFFBBFFE5FD5623843F49CE8D" author="Blackwelder R. E." box="[763,1040,1473,1499]" pageId="43" pageNumber="44" pagination="189 - 341" refId="ref29526" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFFBBFFE5FF1023A739EECEAA" blockId="43.[189,1044,1473,1533]" box="[189,695,1506,1533]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB3C9051FFBBFFE5FF1023A739EECEAA" box="[189,695,1506,1533]" pageId="43" pageNumber="44">
<taxonomicName id="4C2CCCFCFFBBFFE5FF1023A73A8DCEAA" ID-CoL="7ZN5G" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[189,468,1506,1532]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBBFFE5FF1023A73A8DCEAA" box="[189,468,1506,1532]" italics="true" pageId="43" pageNumber="44">Dichotomius subaeneus</emphasis>
</taxonomicName>
<treatmentCitation id="0A8D916EFFBBFFE5FE5F23A639EACEAA" author="Larsen T. H." box="[498,691,1506,1533]" page="92" pageId="43" pageNumber="44" year="2011">
<bibRefCitation id="EFBDCA8EFFBBFFE5FE5F23A639EACEAA" author="Larsen T. H." box="[498,691,1506,1533]" pageId="43" pageNumber="44" pagination="63" refId="ref30006" refString="Larsen T. H. 2011. Dung beetles of the Kwamalasamutu region, Suriname (Coleoptera: Scarabaeidae: Scarabaeinae) In: O'Shea B. J., Alonso L. E. &amp; Larsen T. H. (eds) A Rapid biological Assessment of the Kwamalasamutu Region, Southwestern Suriname. RAP Bulletin of Biological Assessment 63. Conservation International, Arlington, VA." type="book chapter" year="2011">Larsen 2011: 92</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBBFFE5FF10206439EDCDBA" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8B93B77FFFBBFFE5FF1020643A6FCD6D" blockId="43.[189,310,1569,1595]" box="[189,310,1569,1595]" pageId="43" pageNumber="44">
<heading id="D0DB0013FFBBFFE5FF1020643A6FCD6D" bold="true" box="[189,310,1569,1595]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B9586B6DFFBBFFE5FF1020643A6FCD6D" bold="true" box="[189,310,1569,1595]" pageId="43" pageNumber="44">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBBFFE5FF10200B39EDCDBA" blockId="43.[189,1400,1614,1772]" pageId="43" pageNumber="44">
<taxonomicName id="4C2CCCFCFFBBFFE5FF10200B3A8FCD3E" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[189,470,1614,1640]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBBFFE5FF10200B3A8FCD3E" box="[189,470,1614,1640]" italics="true" pageId="43" pageNumber="44">Dichotomius subaeneus</emphasis>
</taxonomicName>
is separated from other species in the group by the following combination of characters: copper colour. Sparse or absence of setae on body surface. Cephalic carina narrow, wider than high. Ventral clypeal process greatly emarginate. Pronotum not modified. Elytra striae displaying strong, well-imprinted punctuation separated between them by one time diameter of a puncture on elytral disc. Protibia apical spine bifurcate.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBBFFE2FF102154384FC8EC" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8B93B77FFFBBFFE5FF1021543AF3CC7D" blockId="43.[189,426,1809,1835]" box="[189,426,1809,1835]" pageId="43" pageNumber="44">
<heading id="D0DB0013FFBBFFE5FF1021543AF3CC7D" bold="true" box="[189,426,1809,1835]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B9586B6DFFBBFFE5FF1021543AF3CC7D" bold="true" box="[189,426,1809,1835]" pageId="43" pageNumber="44">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBBFFE5FF4721073E2ECC2A" blockId="43.[189,1399,1855,2025]" pageId="43" pageNumber="44">
<materialsCitation id="3B44BD22FFBBFFE5FF4721073E2ECC2A" ID-GBIF-Occurrence="3037418374" collectionCode="BMNH" country="French Guiana" location="FRENCH GUIANA" pageId="43" pageNumber="44" specimenCount="1" typeStatus="lectotype">
<emphasis id="B9586B6DFFBBFFE5FF4721073A3FCC0A" bold="true" box="[234,358,1858,1884]" pageId="43" pageNumber="44">
<typeStatus id="549709DDFFBBFFE5FF4721073A3FCC0A" box="[234,358,1858,1884]" pageId="43" pageNumber="44" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<specimenCount id="9D2A7CF6FFBBFFE5FED7217A3AD7CC0D" box="[378,398,1855,1883]" pageId="43" pageNumber="44" type="male"></specimenCount>
, here designated)
<collectingCountry id="F33BF7EFFFBBFFE5FF1021263AF7CC2B" box="[189,430,1891,1917]" name="French Guiana" pageId="43" pageNumber="44">FRENCH GUIANA</collectingCountry>
<emphasis id="B9586B6DFFBBFFE5FE6521263A87CC2B" bold="true" box="[456,478,1891,1917]" pageId="43" pageNumber="44"></emphasis>
;
<emphasis id="B9586B6DFFBBFFE5FE5D21263959CC2B" box="[496,512,1891,1917]" italics="true" pageId="43" pageNumber="44"></emphasis>
SYNTYPE” [white, rounded with blue margin]/ “67·45” [white, p] /“364”
</materialsCitation>
</paragraph>
<paragraph id="8B93B77FFFBBFFE5FF1221C2383CCCBF" blockId="43.[189,1399,1855,2025]" pageId="43" pageNumber="44">
[grey, hw]/ “
<taxonomicName id="4C2CCCFCFFBBFFE5FEF521C339D3CCF6" authority="Dej. Cayenne" authorityName="Dej. Cayenne" box="[344,650,1926,1952]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBBFFE5FEF521C33A8BCCF6" box="[344,466,1926,1952]" italics="true" pageId="43" pageNumber="44">subaeneus</emphasis>
Dej. Cayenne
</taxonomicName>
” [white, hw] / “
<taxonomicName id="4C2CCCFCFFBBFFE5FCE121C33FD1CCF6" authority="Cast. J. Gillet." box="[844,1160,1926,1953]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBBFFE5FCE121C3389FCCF6" box="[844,966,1926,1952]" italics="true" pageId="43" pageNumber="44">subaeneus</emphasis>
Cast. J. Gillet.
</taxonomicName>
det. 1912” [white, phw] / “LECTOTYPE” [red with black margin, p]/ “
<taxonomicName id="4C2CCCFCFFBBFFE5FC9D21EF3F59CC92" authority="LECTOT." authorityName="LECTOT." box="[816,1024,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9586B6DFFBBFFE5FC9D21EF38D9CC92" box="[816,896,1962,1988]" italics="true" pageId="43" pageNumber="44">Copris</emphasis>
LECTOT.
</taxonomicName>
<taxonomicName id="4C2CCCFCFFBBFFE5FBA421EF3FE7CC92" authority="Cast." authorityName="Castelnau" authorityYear="1840" box="[1033,1214,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subaenea">
<emphasis id="B9586B6DFFBBFFE5FBA421EF3F21CC92" box="[1033,1144,1962,1988]" italics="true" pageId="43" pageNumber="44">subaenea</emphasis>
Cast.
</taxonomicName>
det, F. Vaz-de- Mello, 2011”/ “NHMUK 010847024” [white, p];
<collectionCode id="ED3D2FBAFFBBFFE5FCA8218A3838CCBF" box="[773,865,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="43" pageNumber="44">BMNH</collectionCode>
.
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF47274D3AB8CA74" blockId="44.[189,1399,264,539]" box="[234,481,264,290]" pageId="44" pageNumber="45">
<emphasis id="B9586B6DFFBCFFE2FF47274D3AB8CA74" bold="true" box="[234,481,264,290]" pageId="44" pageNumber="45">Additional material</emphasis>
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1027693FE7C94D" blockId="44.[189,1399,264,539]" pageId="44" pageNumber="45">
<materialsCitation id="3B44BD22FFBCFFE2FF1027693F8ACA10" ID-GBIF-Occurrence="3037418630" box="[189,1235,300,326]" collectingDate="1997-11" collectionCode="CEMT" collectorName="E. Andressen" country="Brazil" location="Amazonas" pageId="44" pageNumber="45" specimenCount="7" specimenCount-female="7" stateProvince="Amazonas">
<collectingCountry id="F33BF7EFFFBCFFE2FF1027693A7FCA10" box="[189,294,300,326]" name="Brazil" pageId="44" pageNumber="45">BRAZIL</collectingCountry>
<emphasis id="B9586B6DFFBCFFE2FEE927693A9FCA10" bold="true" box="[324,454,300,326]" pageId="44" pageNumber="45">
<collectingRegion id="49E8799DFFBCFFE2FEE927693A9FCA10" box="[324,454,300,326]" country="Brazil" name="Amazonas" pageId="44" pageNumber="45">Amazonas</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FFBCFFE2FE4F2769397FCA10" box="[482,550,300,326]" pageId="44" pageNumber="45" type="female">7 ♀♀</specimenCount>
;
<locationDeviation id="9A21AE5DFFBCFFE2FD9B27693850CA10" box="[566,777,300,326]" pageId="44" pageNumber="45">
<quantity id="4CD41A9AFFBCFFE2FD9B276939D8CA10" box="[566,641,300,326]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="km" value="90.0">90 km</quantity>
N Manaus
</locationDeviation>
;
<date id="FF9291BFFFBCFFE2FCBB276938CDCA10" box="[790,916,300,326]" pageId="44" pageNumber="45" value="1997-11">
<collectingDate id="EFD66857FFBCFFE2FCBB276938CDCA10" box="[790,916,300,326]" pageId="44" pageNumber="45" value="1997-11">Nov. 1997</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FC0F27693F67CA10" box="[930,1086,300,326]" pageId="44" pageNumber="45">E. Andressen</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FB2F27693F8ACA10" box="[1154,1235,300,326]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FB4227693E3DCA3F" ID-GBIF-Occurrence="3037418433" collectingDate="1966-03-04" collectingDateMax="1966-03-12" collectingDateMin="1966-03-04" collectionCode="CEMT" collectorName="W. D. Edmonds" country="Brazil" location="Res. Florest. Ducke" municipality="Res. Florest. Ducke" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="2" stateProvince="Amazonas">
<specimenCount id="9D2A7CF6FFBCFFE2FB4227693E6ACA10" box="[1263,1331,300,326]" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="6BF72D05FFBCFFE2FAEE27693A37CA3F" pageId="44" pageNumber="45">Res. Florest. Ducke</collectingMunicipality>
,
<locationDeviation id="9A21AE5DFFBCFFE2FED7270A3930CA3C" box="[378,617,335,362]" location="Manaus" pageId="44" pageNumber="45">
<quantity id="4CD41A9AFFBCFFE2FED7270A3A9CCA3F" box="[378,453,335,361]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="44" pageNumber="45" unit="km" value="26.0">26 km</quantity>
E of Manaus
</locationDeviation>
, Am. [Amazonas];
<date id="FF9291BFFFBCFFE2FCE2270A3F56CA3F" box="[847,1039,335,362]" pageId="44" pageNumber="45" value="1966-03-04" valueMax="1966-03-12" valueMin="1966-03-04">
<collectingDate id="EFD66857FFBCFFE2FCE2270A3F56CA3F" box="[847,1039,335,362]" pageId="44" pageNumber="45" value="1966-03-04" valueMax="1966-03-12" valueMin="1966-03-04">412 Mar. 1966</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FBB627153F96CA3F" box="[1051,1231,335,362]" pageId="44" pageNumber="45">W.D. Edmonds</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FABE270A3E3DCA3F" box="[1299,1380,335,361]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FF13273638ADCADB" ID-GBIF-Occurrence="3037418399" box="[190,1012,370,397]" collectingDate="1927-12-17" collectionCode="CEMT" country="Brazil" county="Manaos" location="Amaz." municipality="Rio Negro" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas">
<specimenCount id="9D2A7CF6FFBCFFE2FF1327363BB5CADB" box="[190,236,371,397]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62F2CFF3FFBCFFE2FF5027363A04CADB" box="[253,349,371,397]" pageId="44" pageNumber="45">Manáos</collectingCounty>
,
<collectingMunicipality id="6BF72D05FFBCFFE2FEC727363AB3CADB" box="[362,490,371,397]" pageId="44" pageNumber="45">Rio Negro</collectingMunicipality>
,
<location id="8EF3E1A4FFBCFFE2FE5A2736391DCADB" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FE5A2736391DCADB" box="[503,580,371,397]" country="Brazil" county="Manaos" municipality="Rio Negro" name="Amaz." pageId="44" pageNumber="45" stateProvince="Amazonas">Amaz.</location>
[Amazonas];
<date id="FF9291BFFFBCFFE2FD43273638CACADB" box="[750,915,371,397]" pageId="44" pageNumber="45" value="1927-12-17">
<collectingDate id="EFD66857FFBCFFE2FD43273638CACADB" box="[750,915,371,397]" pageId="44" pageNumber="45" value="1927-12-17">17 Dec. 1927</collectingDate>
</date>
;
<collectionCode id="ED3D2FBAFFBCFFE2FC0C273638ADCADB" box="[929,1012,371,397]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FFBCFFE2FBB42737398FCA82" ID-GBIF-Occurrence="3037418339" collectedFrom="rainforest" collectingDate="2005-04" collectingDateMax="2005-05-31" collectingDateMin="2005-04" collectingMethod="dung pitfall trap" collectionCode="CEMT" collectorName="T. A. Gardner &amp; M. I. M. Hernandez" country="Brazil" elevation="180" latitude="-0.6333333" location="Reserva do Pacanari" longLatPrecision="1308" longitude="-52.566666" municipality="Monte Dourado" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Para">
<emphasis id="B9586B6DFFBCFFE2FBB427373F0CCADB" bold="true" box="[1049,1109,370,397]" pageId="44" pageNumber="45">
<collectingRegion id="49E8799DFFBCFFE2FBB427373F0CCADB" box="[1049,1109,370,397]" pageId="44" pageNumber="45">Pará</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FFBCFFE2FBDE27363FFBCADB" box="[1139,1186,371,397]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BF72D05FFBCFFE2FB1F27363E2BCADB" box="[1202,1394,371,397]" pageId="44" pageNumber="45">Monte Dourado</collectingMunicipality>
,
<location id="8EF3E1A4FFBCFFE2FF1027D23AE9CAE6" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FF1027D23AE9CAE6" box="[189,432,406,433]" country="Brazil" latitude="-0.6333333" longLatPrecision="1308" longitude="-52.566666" municipality="Monte Dourado" name="Reserva do Pacanari" pageId="44" pageNumber="45" stateProvince="Para">Reserva do Pacanari</location>
;
<elevation id="0001504CFFBCFFE2FE1127D3391FCAE6" box="[444,582,406,433]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="44" pageNumber="45" unit="m" value="180.0">
<quantity id="4CD41A9AFFBCFFE2FE1127D3395FCAE7" box="[444,518,406,433]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="44" pageNumber="45" unit="m" value="180.0">180 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FDFE27D039F1CAE6" box="[595,680,405,433]" degrees="0" direction="south" minutes="38" orientation="latitude" pageId="44" pageNumber="45" precision="925" value="-0.6333333">0º38 S</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBCFFE2FD1827D0387DCAE7" box="[693,804,405,433]" degrees="52" direction="west" minutes="34" orientation="longitude" pageId="44" pageNumber="45" precision="925" value="-52.566666">52º34 W</geoCoordinate>
;
<collectedFrom id="3830F482FFBCFFE2FC9D27D338FDCAE6" box="[816,932,406,432]" pageId="44" pageNumber="45">rainforest</collectedFrom>
,
<collectingMethod id="526DCF68FFBCFFE2FC0227D33F29CAE7" box="[943,1136,406,433]" pageId="44" pageNumber="45">dung pitfall trap</collectingMethod>
;
<date id="FF9291BFFFBCFFE2FBD727D33E6ECAE7" box="[1146,1335,406,433]" pageId="44" pageNumber="45" value="2005-04" valueMax="2005-05-31" valueMin="2005-04-00">
<collectingDate id="EFD66857FFBCFFE2FBD727D33E6ECAE7" box="[1146,1335,406,433]" pageId="44" pageNumber="45" value="2005-04" valueMax="2005-05-31" valueMin="2005-04-00">Apr.May 2005</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FAEE27D33A44CA85" pageId="44" pageNumber="45">T.A. Gardner</collectorName>
and
<collectorName id="26D9D2A9FFBCFFE2FEF027FF3962CA85" box="[349,571,441,468]" pageId="44" pageNumber="45">M.I.M. Hernández</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FD2827FF398FCA82" box="[645,726,442,468]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FD5B27FF3A8FC94D" ID-GBIF-Occurrence="3037418536" collectedFrom="rainforest" collectingDate="2005-04" collectingDateMax="2005-05-31" collectingDateMin="2005-04" collectingMethod="dung pitfall trap" collectionCode="BMNH" collectorName="T. A. Gardner &amp; M. I. M. Hernandez" country="Brazil" elevation="180" latitude="-0.6333333" location="Reserva do Pacanari" longLatPrecision="1308" longitude="-52.566666" municipality="Monte Dourado" pageId="44" pageNumber="45" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Para">
<specimenCount id="9D2A7CF6FFBCFFE2FD5B27FF3864CA85" box="[758,829,439,468]" pageId="44" pageNumber="45" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D2A7CF6FFBCFFE2FCE327FC38CCCA82" box="[846,917,441,468]" pageId="44" pageNumber="45" type="female">5 ♀♀</specimenCount>
;
<collectingMunicipality id="6BF72D05FFBCFFE2FC0A27FF3F31CA85" box="[935,1128,441,468]" pageId="44" pageNumber="45">Monte Dourado</collectingMunicipality>
,
<location id="8EF3E1A4FFBCFFE2FBDA27FF3E2BCA85" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FBDA27FF3E2BCA85" box="[1143,1394,441,468]" country="Brazil" latitude="-0.6333333" longLatPrecision="1308" longitude="-52.566666" municipality="Monte Dourado" name="Reserva do Pacanari" pageId="44" pageNumber="45" stateProvince="Para">Reserva do Pacanari</location>
;
<elevation id="0001504CFFBCFFE2FF1027983A11CAA1" box="[189,328,477,504]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="44" pageNumber="45" unit="m" value="180.0">
<quantity id="4CD41A9AFFBCFFE2FF1027983A51CAAE" box="[189,264,477,504]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="44" pageNumber="45" unit="m" value="180.0">180 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FEFB279E3AF5CAA1" box="[342,428,475,503]" degrees="0" direction="south" minutes="38" orientation="latitude" pageId="44" pageNumber="45" precision="925" value="-0.6333333">0º38 S</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBCFFE2FE17279E3970CAAE" box="[442,553,475,504]" degrees="52" direction="west" minutes="34" orientation="longitude" pageId="44" pageNumber="45" precision="925" value="-52.566666">52º34 W</geoCoordinate>
;
<collectedFrom id="3830F482FFBCFFE2FD9B279839F3CAA1" box="[566,682,477,503]" pageId="44" pageNumber="45">rainforest</collectedFrom>
,
<collectingMethod id="526DCF68FFBCFFE2FD1B2798382ECAAE" box="[694,887,477,504]" pageId="44" pageNumber="45">dung pitfall trap</collectingMethod>
;
<date id="FF9291BFFFBCFFE2FC2F27983F19CAAE" box="[898,1088,477,504]" pageId="44" pageNumber="45" value="2005-04" valueMax="2005-05-31" valueMin="2005-04-00">
<collectingDate id="EFD66857FFBCFFE2FC2F27983F19CAAE" box="[898,1088,477,504]" pageId="44" pageNumber="45" value="2005-04" valueMax="2005-05-31" valueMin="2005-04-00">Apr.May 2005</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FBE127983FB1CAA1" box="[1100,1256,477,503]" pageId="44" pageNumber="45">T.A. Gardner</collectorName>
and
<collectorName id="26D9D2A9FFBCFFE2FA8F279B3A65C94C" pageId="44" pageNumber="45">M.I.M. Hernández</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FED324443A8FC94D" box="[382,470,513,539]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FE4224443FE3C94D" ID-GBIF-Occurrence="3037418437" box="[495,1210,510,539]" collectionCode="CEMT" collectorName="N. Degallier" country="Brazil" location="Rio Tocantins" municipality="Tucuruis" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins">
<specimenCount id="9D2A7CF6FFBCFFE2FE4224443945C94C" box="[495,540,510,539]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BF72D05FFBCFFE2FD87244539CDC94C" box="[554,660,512,538]" pageId="44" pageNumber="45">Tucuruis</collectingMunicipality>
, 8.84,
<location id="8EF3E1A4FFBCFFE2FD4F244538D0C94C" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FD4F244538D0C94C" box="[738,905,512,538]" country="Brazil" municipality="Tucuruis" name="Rio Tocantins" pageId="44" pageNumber="45" stateProvince="Tocantins">Rio Tocantins</location>
;
<collectorName id="26D9D2A9FFBCFFE2FC3924443F7CC94C" box="[916,1061,512,539]" pageId="44" pageNumber="45">N. Degallier</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FBCA24443FE3C94D" box="[1127,1210,513,539]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1024073A91C802" blockId="44.[189,1399,576,853]" pageId="44" pageNumber="45">
<materialsCitation id="3B44BD22FFBCFFE2FF1024073AE9C9D6" ID-GBIF-Occurrence="3037418480" collectingDate="1997-03" collectionCode="CEMT" collectorName="F. Feer." country="French Guiana" elevation="155" latitude="4.0833335" location="Nouragues" longLatPrecision="1307" longitude="-52.666668" pageId="44" pageNumber="45" specimenCount="3" specimenCount-female="1" stateProvince="Cayenne">
<collectingCountry id="F33BF7EFFFBCFFE2FF1024073AF2C90A" box="[189,427,578,604]" name="French Guiana" pageId="44" pageNumber="45">FRENCH GUIANA</collectingCountry>
<emphasis id="B9586B6DFFBCFFE2FE6124073960C90A" bold="true" box="[460,569,578,604]" pageId="44" pageNumber="45">
<collectingRegion id="49E8799DFFBCFFE2FE6124073960C90A" box="[460,569,578,604]" pageId="44" pageNumber="45">Cayenne</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FFBCFFE2FDFE240739CEC90A" box="[595,663,576,604]" pageId="44" pageNumber="45">2 ♂♂</specimenCount>
,
<specimenCount id="9D2A7CF6FFBCFFE2FD0B2407398DC90B" box="[678,724,578,605]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FD492406383EC90B" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FD492406383EC90B" box="[740,871,579,605]" country="French Guiana" latitude="4.0833335" longLatPrecision="1307" longitude="-52.666668" name="Nouragues" pageId="44" pageNumber="45" stateProvince="Cayenne">Nouragues</location>
;
<elevation id="0001504CFFBCFFE2FCDE24073F59C90A" box="[883,1024,578,605]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="44" pageNumber="45" unit="m" value="155.0">
<quantity id="4CD41A9AFFBCFFE2FCDE240738E6C90B" box="[883,959,578,605]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="44" pageNumber="45" unit="m" value="155.0">155 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FBA324053F33C90B" box="[1038,1130,576,605]" degrees="4" direction="north" minutes="05" orientation="latitude" pageId="44" pageNumber="45" precision="925" value="4.0833335">4º05 N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBCFFE2FBDA24053FBEC90B" box="[1143,1255,576,605]" degrees="52" direction="west" minutes="40" orientation="longitude" pageId="44" pageNumber="45" precision="925" value="-52.666668">52º40 W</geoCoordinate>
;
<date id="FF9291BFFFBCFFE2FB5824063E2BC90A" box="[1269,1394,578,605]" pageId="44" pageNumber="45" value="1997-03">
<collectingDate id="EFD66857FFBCFFE2FB5824063E2BC90A" box="[1269,1394,578,605]" pageId="44" pageNumber="45" value="1997-03">Mar. 1997</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FF1024233A4EC9D6" box="[189,279,614,640]" pageId="44" pageNumber="45">F. Feer.</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FEF224233AE9C9D6" box="[351,432,614,640]" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FE6224233A85C9F5" ID-GBIF-Occurrence="3037418595" collectingDate="2011-03-06" collectionCode="CEMT" collectorName="Seag" country="French Guiana" elevation="430" latitude="4.619167" location="Nouragues Camp." longLatPrecision="21" longitude="-53.40889" pageId="44" pageNumber="45" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Cayenne">
<specimenCount id="9D2A7CF6FFBCFFE2FE6224233AA6C929" box="[463,511,611,640]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D2A7CF6FFBCFFE2FDBC2423390EC9D6" box="[529,599,614,640]" pageId="44" pageNumber="45" type="female">3 ♀♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FDC42423381AC9D6" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FDC42423381AC9D6" box="[617,835,614,640]" country="French Guiana" latitude="4.619167" longLatPrecision="21" longitude="-53.40889" name="Nouragues Camp." pageId="44" pageNumber="45" stateProvince="Cayenne">Nouragues Camp.</location>
;
<elevation id="0001504CFFBCFFE2FCFF242338BAC929" box="[850,995,613,640]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="44" pageNumber="45" unit="m" value="430.0">
<quantity id="4CD41A9AFFBCFFE2FCFF242338F9C9D6" box="[850,928,614,640]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="44" pageNumber="45" unit="m" value="430.0">430 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FC5924213F35C9D6" box="[1012,1132,612,640]" degrees="4" direction="north" minutes="37" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="9" value="4.619167">4º379″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBCFFE2FBD124213E4EC9D6" box="[1148,1303,612,640]" degrees="53" direction="west" minutes="24" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="32" value="-53.40889">53º2432″ W</geoCoordinate>
;
<date id="FF9291BFFFBCFFE2FA8A24203BA3C9F5" pageId="44" pageNumber="45" value="2011-03-06">
<collectingDate id="EFD66857FFBCFFE2FA8A24203BA3C9F5" pageId="44" pageNumber="45" value="2011-03-06">6 Mar. 2011</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FEA424CC3A1BC9F5" box="[265,322,649,675]" pageId="44" pageNumber="45">Seag</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FE2624CC3A85C9F5" box="[395,476,649,675]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FE5124CC3AA5C990" ID-GBIF-Occurrence="3037418412" collectingDate="2014-05" collectionCode="CEMT" collectorName="F. Feer." country="French Guiana" elevation="35" latitude="3.9" location="Saint Georges de L' Oyapok" longLatPrecision="1307" longitude="-51.8" municipality="Saint Georges" pageId="44" pageNumber="45" specimenCount="5" specimenCount-female="5" stateProvince="Cayenne">
<specimenCount id="9D2A7CF6FFBCFFE2FE5124CC391BC9F5" box="[508,578,649,676]" pageId="44" pageNumber="45" type="female">5 ♀♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FDF924CC38E9C9F5" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FDF924CC38E9C9F5" box="[596,944,649,675]" country="French Guiana" latitude="3.9" longLatPrecision="1307" longitude="-51.8" municipality="Saint Georges" name="Saint Georges de L' Oyapok" pageId="44" pageNumber="45" stateProvince="Cayenne">Saint Georges de L Oyapok</location>
;
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a.s.l.
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;
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,
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;
<date id="FF9291BFFFBCFFE2FAEC24CC3BA2C990" pageId="44" pageNumber="45" value="2014-05">
<collectingDate id="EFD66857FFBCFFE2FAEC24CC3BA2C990" pageId="44" pageNumber="45" value="2014-05">May 2014</collectingDate>
</date>
;
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leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FE0424E93AA5C990" box="[425,508,684,710]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
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;
<collectingMunicipality id="6BF72D05FFBCFFE2FC9A24E93889C990" box="[823,976,684,710]" pageId="44" pageNumber="45">Sain Eugène</collectingMunicipality>
,
<location id="8EF3E1A4FFBCFFE2FC7324E83FBBC990" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FC7324E83FBBC990" box="[990,1250,684,711]" country="French Guiana" latitude="4.866667" longLatPrecision="1306" longitude="-53.066666" municipality="Sain Eugene" name="Barrage de Petit Saut" pageId="44" pageNumber="45" stateProvince="Guyane">Barrage de Petit Saut</location>
;
<elevation id="0001504CFFBCFFE2FB5D24E93E29C990" box="[1264,1392,684,711]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="m" value="90.0">
<quantity id="4CD41A9AFFBCFFE2FB5D24E93E77C991" box="[1264,1326,684,711]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="m" value="90.0">90 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FF10248B3A41C9BC" box="[189,280,718,746]" degrees="4" direction="north" minutes="52" orientation="latitude" pageId="44" pageNumber="45" precision="925" value="4.866667">4º52 N</geoCoordinate>
,
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;
<date id="FF9291BFFFBCFFE2FE3E24953956C9BC" box="[403,527,720,746]" pageId="44" pageNumber="45" value="1998-12">
<collectingDate id="EFD66857FFBCFFE2FE3E24953956C9BC" box="[403,527,720,746]" pageId="44" pageNumber="45" value="1998-12">Dec. 1998</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FDB124953934C9BC" box="[540,621,720,746]" pageId="44" pageNumber="45">F. Feer</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FD1D2495385AC9BC" box="[688,771,720,746]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FFBCFFE2FC8B248A39C3C85B" ID-GBIF-Occurrence="3037418442" collectingDate="1969-10" collectingDateMax="1969-11-31" collectingDateMin="1969-10" collectionCode="CEMT" collectorName="Balachowsky-Gruner" country="French Guiana" location="Guyanne-Mission" pageId="44" pageNumber="45" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Saul">
<emphasis id="B9586B6DFFBCFFE2FC8B248A3804C9BF" bold="true" box="[806,861,719,745]" pageId="44" pageNumber="45">
<collectingRegion id="49E8799DFFBCFFE2FC8B248A3804C9BF" box="[806,861,719,745]" pageId="44" pageNumber="45">Saül</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FFBCFFE2FCDA249538E2C9BC" box="[887,955,718,746]" pageId="44" pageNumber="45" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D2A7CF6FFBCFFE2FC6424953F54C9BC" box="[969,1037,720,746]" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FBB124953FABC9BC" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FBB124953FABC9BC" box="[1052,1266,720,746]" country="French Guiana" name="Guyanne-Mission" pageId="44" pageNumber="45" stateProvince="Saul">Guyanne-Mission</location>
;
<date id="FF9291BFFFBCFFE2FB5324953BA2C85B" pageId="44" pageNumber="45" value="1969-10" valueMax="1969-11-31" valueMin="1969-10-00">
<collectingDate id="EFD66857FFBCFFE2FB5324953BA2C85B" pageId="44" pageNumber="45" value="1969-10" valueMax="1969-11-31" valueMin="1969-10-00">Oct.Nov. 1969</collectingDate>
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leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FDE424B639C3C85B" box="[585,666,755,781]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
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<materialsCitation id="3B44BD22FFBCFFE2FD1B24B63F51C867" ID-GBIF-Occurrence="3037418398" collectingDate="2011-02-28" collectionCode="CEMT" collectorName="Saint Laurent du Maroni &amp; Seag" country="French Guiana" elevation="326" latitude="3.622778" location="Belvedere de Saul" longLatPrecision="21" longitude="-53.215836" pageId="44" pageNumber="45" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Saul">
<specimenCount id="9D2A7CF6FFBCFFE2FD1B24B639A2C85B" box="[694,763,753,781]" pageId="44" pageNumber="45" type="male">2 ♂♂</specimenCount>
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;
<collectorName id="26D9D2A9FFBCFFE2FCF224B63FD1C85B" box="[863,1160,755,782]" pageId="44" pageNumber="45">Saint Laurent du Maroni</collectorName>
,
<location id="8EF3E1A4FFBCFFE2FB3B24B63E2BC85B" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FB3B24B63E2BC85B" box="[1174,1394,755,781]" country="French Guiana" latitude="3.622778" longLatPrecision="21" longitude="-53.215836" name="Belvedere de Saul" pageId="44" pageNumber="45" stateProvince="Saul">Bélvédère de Saül</location>
;
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<quantity id="4CD41A9AFFBCFFE2FF1025533A51C867" box="[189,264,790,817]" metricMagnitude="2" metricUnit="m" metricValue="3.26" pageId="44" pageNumber="45" unit="m" value="326.0">326 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBCFFE2FEFA25503A85C867" box="[343,476,789,817]" degrees="3" direction="north" minutes="37" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="22" value="3.622778">3º3722″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBCFFE2FE44255039DBC867" box="[489,642,789,817]" degrees="53" direction="west" minutes="12" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="57" value="-53.215836">53º1257″ W</geoCoordinate>
;
<date id="FF9291BFFFBCFFE2FD2225523874C867" box="[655,813,791,817]" pageId="44" pageNumber="45" value="2011-02-28">
<collectingDate id="EFD66857FFBCFFE2FD2225523874C867" box="[655,813,791,817]" pageId="44" pageNumber="45" value="2011-02-28">28 Feb. 2011</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FC972552382AC867" box="[826,883,791,817]" pageId="44" pageNumber="45">Seag</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FC1A25523F51C867" box="[951,1032,791,817]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FB8E25523A9DC802" ID-GBIF-Occurrence="3037418583" collectingDate="1991-11" collectionCode="CEMT" collectorName="P. Arnaud" country="French Guiana" location="St. Laurent" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Saul">
<specimenCount id="9D2A7CF6FFBCFFE2FB8E25523F08C867" box="[1059,1105,791,817]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FBCD25523FBEC867" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FBCD25523FBEC867" box="[1120,1255,791,817]" country="French Guiana" name="St. Laurent" pageId="44" pageNumber="45" stateProvince="Saul">St. Laurent</location>
;
<date id="FF9291BFFFBCFFE2FB5E25523E2BC867" box="[1267,1394,791,817]" pageId="44" pageNumber="45" value="1991-11">
<collectingDate id="EFD66857FFBCFFE2FB5E25523E2BC867" box="[1267,1394,791,817]" pageId="44" pageNumber="45" value="1991-11">Nov. 1991</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FF10257E3A76C802" box="[189,303,826,853]" pageId="44" pageNumber="45">P. Arnaud</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FEDC257F3A9DC802" box="[369,452,826,852]" collectionName="CEMT" pageId="44" pageNumber="45">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF102539384FC8EC" blockId="44.[189,1398,890,954]" pageId="44" pageNumber="45">
<materialsCitation id="3B44BD22FFBCFFE2FF1025393FB2C8C0" ID-GBIF-Occurrence="3037418344" box="[189,1259,890,918]" collectionCode="BMNH" collectorName="H. Bedding" country="Guyana" location="Oronque and New River Head" municipality="River Head" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F33BF7EFFFBCFFE2FF1025393A68C8C0" box="[189,305,892,918]" name="Guyana" pageId="44" pageNumber="45">GUIANA</collectingCountry>
<specimenCount id="9D2A7CF6FFBCFFE2FEE525393A2CC8C0" box="[328,373,890,918]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FE2E253939B2C8C0" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FE2E253939B2C8C0" box="[387,747,892,918]" country="Guyana" municipality="River Head" name="Oronque and New River Head" pageId="44" pageNumber="45">Oronque and New River Head</location>
; 1938; 1938-346;
<collectorName id="26D9D2A9FFBCFFE2FC6D25393F1EC8C0" box="[960,1095,892,918]" pageId="44" pageNumber="45">H. Bedding</collectorName>
B.M;
<collectionCode id="ED3D2FBAFFBCFFE2FB3E25393FB2C8C0" box="[1171,1259,892,918]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBCFFE2FAA92539384BC8EC" ID-GBIF-Occurrence="3037418364" collectingDate="1938-03" collectingDateMax="1938-05-31" collectingDateMin="1938-03" collectionCode="BMNH" collectorName="C. A. Hudson" country="Guyana" location="New River" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2A7CF6FFBCFFE2FAA925393E68C8C0" box="[1284,1329,892,918]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<location id="8EF3E1A4FFBCFFE2FA9225393A5BC8EF" LSID="urn:lsid:plazi:treatment:03850669FFBBFFE0FE0123643892C881:8EF3E1A4FFBCFFE2FA9225393A5BC8EF" country="Guyana" name="New River" pageId="44" pageNumber="45">New River</location>
;
<date id="FF9291BFFFBCFFE2FEA325E53A97C8EF" box="[270,462,927,954]" pageId="44" pageNumber="45" value="1938-03" valueMax="1938-05-31" valueMin="1938-03-00">
<collectingDate id="EFD66857FFBCFFE2FEA325E53A97C8EF" box="[270,462,927,954]" pageId="44" pageNumber="45" value="1938-03" valueMax="1938-05-31" valueMin="1938-03-00">Mar.May 1938</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBCFFE2FE7725DA392DC8EF" box="[474,628,927,953]" pageId="44" pageNumber="45">C.A. Hudson</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBCFFE2FD1B25E5384BC8EC" box="[694,786,928,954]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBCFFE0FF1025A53917C999" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="8B93B77FFFBCFFE2FF1025A53A32C8AC" blockId="44.[189,363,992,1018]" box="[189,363,992,1018]" pageId="44" pageNumber="45">
<heading id="D0DB0013FFBCFFE2FF1025A53A32C8AC" bold="true" box="[189,363,992,1018]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B9586B6DFFBCFFE2FF1025A53A32C8AC" bold="true" box="[189,363,992,1018]" pageId="44" pageNumber="45">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF4722553A73CF7C" blockId="44.[189,1399,1040,1137]" box="[234,298,1040,1066]" pageId="44" pageNumber="45">
<emphasis id="B9586B6DFFBCFFE2FF4722553A73CF7C" bold="true" box="[234,298,1040,1066]" pageId="44" pageNumber="45">Male</emphasis>
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1022763855CF26" blockId="44.[189,1399,1040,1137]" pageId="44" pageNumber="45">
MEASUREMENTS. Length:
<quantity id="4CD41A9AFFBCFFE2FE4F22763916CF1B" box="[482,591,1075,1101]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="44" pageNumber="45" unit="mm" value="20.8">20.8 mm</quantity>
. Dorsal and ventral surface copper to dark copper, shiny with olive green reflection. Body with setae sparse or absent.
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1022DC3949CE4A" blockId="44.[188,1399,1176,1308]" pageId="44" pageNumber="45">
HEAD. Anterior surface rugose extending posteriorly not reaching base of cephalic carina. Cephalic carina narrow, wider than high; anterior surface smooth with scattered punctures. Median tubercle one and half times as high as lateral tubercles which are rounded apically. (
<figureCitation id="1317ABFAFFBCFFE2FC41229A3F01CFAC" box="[1004,1112,1247,1274]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="44" pageNumber="45">Fig. 15A</figureCitation>
). Ventral clypeal process greatly emarginate (
<figureCitation id="1317ABFAFFBCFFE2FE0B2347395ACE4A" box="[422,515,1282,1308]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="44" pageNumber="45">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1023013FA0CE9E" blockId="44.[189,1400,1348,1481]" pageId="44" pageNumber="45">PRONOTUM. Twice as wide as long and as wide as elytra. Anterior disc declivity weak. Pronotal medial disc shiny, with fine well-impressed punctures. Antero-lateral fossae with ocellate punctures. Ocellate punctures along anterior and posterior margins, never on lateral. Hypomeral disc smooth, without punctures; external margin bearing single row of organized long setae visible in dorsal view.</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1023B538D0CD89" blockId="44.[185,1401,1520,1759]" pageId="44" pageNumber="45">
ABDOMEN. Prosternum surface microsculpture shagreened, glabrous.Mesosternum surface microsculpture shagreened with ocellate setigerous punctures, setae short. Metepisternum surface smooth, displaying setigerous punctures. Metasternum antero-medial edge with superficial weak marked punctuation next to apical margins of mesocoxa (
<figureCitation id="1317ABFAFFBCFFE2FD92201F39F7CD23" box="[575,686,1626,1653]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="44" pageNumber="45">Fig. 15D</figureCitation>
); ocellate punctation restricted mainly to lateral edges, few setigerous punctures near metepisternal suture. Ocellate punctures along anterior margins of each sternite, denser in lateral areas and medially vanished. Pygidium surface displaying well-impressed simple punctures, basal margin with wide ocellate punctures.
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1021423BADCC31" blockId="44.[189,1399,1798,1895]" pageId="44" pageNumber="45">ELYTRA. Interstriae smooth, shiny with fine randomly separated punctures. Striae well delimited and distinctly punctate, punctures wider than striae and separated by twice diameter of a puncture on elytral disc.</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF1021CA3831CCFF" blockId="44.[189,872,1934,1961]" box="[189,872,1934,1961]" pageId="44" pageNumber="45">
LEGS. Metacoxa glabrous, fore calcar bifurcate (
<figureCitation id="1317ABFAFFBCFFE2FD4221CB3803CCFF" box="[751,858,1934,1961]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="44" pageNumber="45">Fig. 15C</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFFBCFFE2FF10218A3867CCBF" blockId="44.[189,830,1998,2025]" box="[189,830,1998,2025]" pageId="44" pageNumber="45">
PARAMERES AND LAMELLA COPULATRIX. As in
<figureCitation id="1317ABFAFFBCFFE2FD18218B385CCCBF" box="[693,773,1998,2025]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="44" pageNumber="45">Fig. 15</figureCitation>
EF.
</paragraph>
<caption id="DF53E7F7FFBDFFE3FF1021C238B8CCB1" ID-DOI="http://doi.org/10.5281/zenodo.4542137" ID-Zenodo-Dep="4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,232,1927,1953]" targetBox="[219,1375,274,1893]" targetPageId="45">
<paragraph id="8B93B77FFFBDFFE3FF1021C238B8CCB1" blockId="45.[188,1399,1925,2024]" pageId="45" pageNumber="46">
<emphasis id="B9586B6DFFBDFFE3FF1021C23A41CCF7" bold="true" box="[189,280,1927,1953]" pageId="45" pageNumber="46">Fig. 15.</emphasis>
<taxonomicName id="4C2CCCFCFFBDFFE3FEB321C23856CCF7" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[286,783,1927,1953]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="B9586B6DFFBDFFE3FEB321C2396DCCF7" box="[286,564,1927,1953]" italics="true" pageId="45" pageNumber="46">Dichotomius subaeneus</emphasis>
(
<bibRefCitation id="EFBDCA8EFFBDFFE3FDEE21C2385CCCF7" author="Castelnau M." box="[579,773,1927,1953]" pageId="45" pageNumber="46" pagination="1 - 564" refId="ref29645" refString="Castelnau M. 1840. Histoire naturelle des coleopteres. In: Blanchard E. (eds) Histoire naturelle des animaux articules. Vol. 2: 1 - 564. Dumenil, Paris." type="journal article" year="1840">Castelnau, 1840</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B9586B6DFFBDFFE3FCB421C23809CCF7" bold="true" box="[793,848,1927,1953]" pageId="45" pageNumber="46">AB</emphasis>
. Dorsal habitus.
<emphasis id="B9586B6DFFBDFFE3FBB821C23F73CCF7" bold="true" box="[1045,1066,1927,1953]" pageId="45" pageNumber="46">A</emphasis>
. ♂.
<emphasis id="B9586B6DFFBDFFE3FBF021C23F28CCF7" bold="true" box="[1117,1137,1927,1953]" pageId="45" pageNumber="46">B</emphasis>
. ♀.
<emphasis id="B9586B6DFFBDFFE3FB0F21C23FEECCF7" bold="true" box="[1186,1207,1927,1953]" pageId="45" pageNumber="46">C</emphasis>
. Lateral view of the aedeagus.
<emphasis id="B9586B6DFFBDFFE3FEF221EF3A2DCC92" bold="true" box="[351,372,1962,1988]" pageId="45" pageNumber="46">D</emphasis>
. Protibial calcar spine bifurcate.
<emphasis id="B9586B6DFFBDFFE3FD5921EF3851CC92" bold="true" box="[756,776,1962,1988]" pageId="45" pageNumber="46">E</emphasis>
. Ventral view of the aedeagus.
<emphasis id="B9586B6DFFBDFFE3FBDC21EF3FDACC92" bold="true" box="[1137,1155,1962,1988]" pageId="45" pageNumber="46">F</emphasis>
. Lamella copulatrix.
<emphasis id="B9586B6DFFBDFFE3FF1121883B8ACCB1" bold="true" box="[188,211,1997,2023]" pageId="45" pageNumber="46">G</emphasis>
. Metasternum pilosity sparse, some few setae on the lateral edges.
</paragraph>
</caption>
<paragraph id="8B93B77FFFBEFFE0FF10274D3AB7CA74" blockId="46.[189,494,264,290]" box="[189,494,264,290]" pageId="46" pageNumber="47">
<heading id="D0DB0013FFBEFFE0FF10274D3AB7CA74" bold="true" box="[189,494,264,290]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9586B6DFFBEFFE0FF10274D3AB7CA74" bold="true" box="[189,494,264,290]" pageId="46" pageNumber="47">Morphological variation</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF47277C3A8BCA02" blockId="46.[189,722,313,376]" box="[234,466,313,340]" pageId="46" pageNumber="47">
<emphasis id="B9586B6DFFBEFFE0FF47277C3A08CA05" bold="true" box="[234,337,313,339]" pageId="46" pageNumber="47">Females</emphasis>
(
<figureCitation id="1317ABFAFFBEFFE0FECC277C3A90CA02" box="[353,457,313,340]" captionStart="Fig" captionStartId="45.[189,232,1927,1953]" captionTargetBox="[219,1375,274,1893]" captionTargetId="figure-63@45.[211,1377,265,1900]" captionTargetPageId="45" captionText="Fig. 15. Dichotomius subaeneus (Castelnau, 1840). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Protibial calcar spine bifurcate. E. Ventral view of the aedeagus. F. Lamella copulatrix. G. Metasternum pilosity sparse, some few setae on the lateral edges." figureDoi="http://doi.org/10.5281/zenodo.4542137" httpUri="https://zenodo.org/record/4542137/files/figure.png" pageId="46" pageNumber="47">Fig. 15B</figureCitation>
)
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10271B398BCA2E" blockId="46.[189,722,313,376]" box="[189,722,350,376]" pageId="46" pageNumber="47">As males except for the following characters:</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF1027E7383AC953" blockId="46.[189,1401,417,517]" pageId="46" pageNumber="47">HEAD. Clypeus elongate. Anterior surface completely rugose, interocular surface with well-impressed flat punctures separated by twice diameter of a puncture. Cephalic carina quadri-dentate, triangular in shape. Median tubercles located forward lateral tubercles.</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10246A3FC7C91E" blockId="46.[189,1182,558,585]" box="[189,1182,558,585]" pageId="46" pageNumber="47">PRONOTUM. Shiny, with fine well-impressed punctures, denser on antero-lateral sides.</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10243738EBC9DA" blockId="46.[189,946,626,652]" box="[189,946,626,652]" pageId="46" pageNumber="47">ABDOMEN. Prosternum antero-medial edge with vertical rugosity.</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF1024F33917C999" blockId="46.[189,590,693,720]" box="[189,590,693,720]" pageId="46" pageNumber="47">LEGS. Profemur distal setae dense.</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBEFFE0FF1024BD3892C881" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8B93B77FFFBEFFE0FF1024BD3A76C844" blockId="46.[189,303,760,786]" box="[189,303,760,786]" pageId="46" pageNumber="47">
<heading id="D0DB0013FFBEFFE0FF1024BD3A76C844" bold="true" box="[189,303,760,786]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9586B6DFFBEFFE0FF1024BD3A76C844" bold="true" box="[189,303,760,786]" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10256C3892C881" blockId="46.[188,1399,809,983]" pageId="46" pageNumber="47">
This species is easily distinguished by its copper colour and sparse pilosity. As with
<taxonomicName id="4C2CCCFCFFBEFFE0FB74256C3E66C815" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[1241,1343,809,835]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="belus">
<emphasis id="B9586B6DFFBEFFE0FB74256C3E66C815" box="[1241,1343,809,835]" italics="true" pageId="46" pageNumber="47">D. belus</emphasis>
</taxonomicName>
, the protibial calcar spine is bifurcate.
<bibRefCitation id="EFBDCA8EFFBEFFE0FDE2250B3860C83E" author="Luederwaldt H." box="[591,825,846,872]" pageId="46" pageNumber="47" pagination="1 - 173" refId="ref30128" refString="Luederwaldt H. 1929. As especies Brasileiras do genero Pinotus. Revista do Museu Paulista 16: 1 - 173." type="journal article" year="1929">Luederwaldt (1929)</bibRefCitation>
mentioned the similarity of these two species to
<taxonomicName id="4C2CCCFCFFBEFFE0FF1325313950C8DB" authority="(Felsche, 1901)" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[190,521,883,909]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cuprinus">
<emphasis id="B9586B6DFFBEFFE0FF1325313A13C8DB" box="[190,330,884,909]" italics="true" pageId="46" pageNumber="47">D. cuprinus</emphasis>
(
<bibRefCitation id="EFBDCA8EFFBEFFE0FEF425363959C8DB" author="Felsche C." box="[345,512,883,909]" pageId="46" pageNumber="47" pagination="135 - 155" refId="ref29807" refString="Felsche C. 1901. Beschreibungen coprophager Scarabaeiden. Deutsche Entomologische Zeitschrift 2: 135 - 155." type="journal article" year="1901">Felsche, 1901</bibRefCitation>
)
</taxonomicName>
and transferred it from the
<taxonomicName id="4C2CCCFCFFBEFFE0FCFF253638F4C8DB" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[850,941,883,909]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="inachus">
<emphasis id="B9586B6DFFBEFFE0FCFF253638F4C8DB" box="[850,941,883,909]" italics="true" pageId="46" pageNumber="47">inachus</emphasis>
</taxonomicName>
group to the
<taxonomicName id="4C2CCCFCFFBEFFE0FBE225363F87C8DB" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1103,1246,883,909]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="B9586B6DFFBEFFE0FBE225363F87C8DB" box="[1103,1246,883,909]" italics="true" pageId="46" pageNumber="47">carbonarius</emphasis>
</taxonomicName>
group. After a detailed inspection of the
<typeStatus id="549709DDFFBEFFE0FDAA25DD392AC8E4" box="[519,627,920,946]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
and other specimens, we suggest that this species belongs to the
<taxonomicName id="4C2CCCFCFFBEFFE0FF1025FB3A6DC881" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[189,308,958,983]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="B9586B6DFFBEFFE0FF1025FB3A6DC881" box="[189,308,958,983]" italics="true" pageId="46" pageNumber="47">D. agenor</emphasis>
</taxonomicName>
species group based on the shape of the cephalic carina.
</paragraph>
</subSubSection>
</treatment>
</document>