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<mods:title id="3E64A00C02305FB86BCF39C130108B82">Correcting sponge names: nomenclatural update of lower taxa level Porifera</mods:title>
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<subSubSection id="7FFDDBE2FFC631729786FE92BFE0066F" box="[151,642,330,357]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="37588869FFC631729786FE92BFE0066F" blockId="33.[151,642,330,393]" box="[151,642,330,357]" pageId="33" pageNumber="34">
<heading id="6C103F05FFC631729786FE92BFE0066F" bold="true" box="[151,642,330,357]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<taxonomicName id="F0E7F3EAFFC631729786FE92BFE0066F" ID-CoL="5P4H4" authority="Dendy, 1922" authorityName="Dendy" authorityYear="1922" box="[151,642,330,357]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="variety" species="carnosa" variety="laxa">
<emphasis id="0593547BFFC631729786FE92BFE0066F" bold="true" box="[151,642,330,357]" pageId="33" pageNumber="34">
<emphasis id="0593547BFFC631729786FE92BC0D066E" bold="true" box="[151,367,330,357]" italics="true" pageId="33" pageNumber="34">Gelliodes carnosa</emphasis>
var.
<emphasis id="0593547BFFC6317296BCFE92BC83066E" bold="true" box="[429,481,330,357]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
<bibRefCitation id="5376F598FFC6317296F9FE93BFE0066F" author="Dendy, A." box="[488,642,330,357]" pageId="33" pageNumber="34" pagination="1 - 164" refId="ref93529" refString="Dendy, A. (1922) Report on the Sigmatotetraxonida collected by H. M. S. ' Sealark' in the Indian Ocean. In: Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. Vol. 7. Transactions of the Linnean Society of London, 18 (1), 1 - 164, pls. 1 - 18. https: // doi. org / 10.1111 / j. 1096 - 3642.1922. tb 00547. x" type="journal article" year="1922">Dendy, 1922</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="7FFDDBE2FFC631729786FEB7BD9D0682" box="[151,255,367,393]" pageId="33" pageNumber="34" type="description">
<paragraph id="37588869FFC631729786FEB7BD9D0682" blockId="33.[151,642,330,393]" box="[151,255,367,393]" pageId="33" pageNumber="34">
(
<figureCitation id="AFDC94ECFFC63172978EFEB7BD950682" box="[159,247,367,393]" captionStart="FIGURE 3" captionStartId="30.[151,250,1830,1854]" captionTargetBox="[222,1366,181,1806]" captionTargetId="figure-24@30.[222,1366,181,1806]" captionTargetPageId="30" captionText="FIGURE 3. A. Amorphina grossa f. latissima Czerniavsky, 1880: pl. III (VII) fig. 23a, BE. Callyspongia rigida Desqueyroux- Faúndez, 1984, B. spicules fig. 25, C. habitus fig. 182, D E. skeleton figs. 183184, F. Chalinissa elongata var. tenuispicula Thum, 1903: figs. 89, G. Gelliodes benedeni var. fortior Topsent, 1917: pl. VI fig. 22, H. Gelliodes carnosa var. laxa Dendy, 1922: pl. 2 fig. 1, I. Gellius carduus and G. carduus var. magellanica Ridley &amp; Dendy, 1886, Ridley &amp; Dendy, 1887: pl. XIII figs. 67, J. Gellius glacialis var. nivea Ridley &amp; Dendy, 1886, Ridley &amp; Dendy, 1887: pl. VIII fig. 8, K. Gellius raphidiophorus Brøndsted, 1933: fig. 7, L. Sigmadocia tubuloramosus; Bergquist &amp; Warne, 1980: pl. 9a." figureDoi="http://doi.org/10.5281/zenodo.10494182" httpUri="https://zenodo.org/record/10494182/files/figure.png" pageId="33" pageNumber="34">Fig. 3H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7FFDDBE2FFC631729786FE6BBFBA06C0" box="[151,728,435,459]" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="37588869FFC631729786FE6BBFBA06C0" blockId="33.[151,728,435,459]" box="[151,728,435,459]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="17F7AF47FFC631729786FE6BBFBA06C0" box="[151,728,435,459]" pageId="33" pageNumber="34">
<treatmentCitation id="B646AE78FFC631729786FE6BBF3D06C0" author="Dendy, A." box="[151,607,435,459]" page="29" pageId="33" pageNumber="34" year="1922">
<taxonomicName id="F0E7F3EAFFC631729786FE6BBF3D06C0" ID-CoL="5P4H4" authority="Dendy, 1922: 29" authorityName="Dendy" authorityPageNumber="29" authorityYear="1922" box="[151,607,435,459]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="variety" species="carnosa" variety="laxa">
<emphasis id="0593547BFFC631729786FE6BBC3306C0" box="[151,337,435,459]" italics="true" pageId="33" pageNumber="34">Gelliodes carnosa</emphasis>
var.
<emphasis id="0593547BFFC631729693FE6BBCCC06C0" box="[386,430,435,459]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
<bibRefCitation id="5376F598FFC6317296A5FE6BBF3D06C0" author="Dendy, A." box="[436,607,435,459]" pageId="33" pageNumber="34" pagination="1 - 164" refId="ref93529" refString="Dendy, A. (1922) Report on the Sigmatotetraxonida collected by H. M. S. ' Sealark' in the Indian Ocean. In: Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. Vol. 7. Transactions of the Linnean Society of London, 18 (1), 1 - 164, pls. 1 - 18. https: // doi. org / 10.1111 / j. 1096 - 3642.1922. tb 00547. x" type="journal article" year="1922">Dendy, 1922: 29</bibRefCitation>
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, pl. 2 fig. 1.
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</paragraph>
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<subSubSection id="7FFDDBE2FFC631729786FE21BF920470" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="37588869FFC631729786FE21BF920470" blockId="33.[151,1437,505,892]" pageId="33" pageNumber="34">
The variety was described by Dendy from Cargados Carajos, approximate coordinates
<geoCoordinate id="52D3EEAEFFC63172934CFE21B9B1051F" box="[1117,1235,505,532]" degrees="16.5667" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="-16.5667">16.5667°S</geoCoordinate>
<geoCoordinate id="52D3EEAEFFC6317293C8FE21B836051F" box="[1241,1364,505,532]" degrees="59.6167" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="59.6167">59.6167°E</geoCoordinate>
, depth
<quantity id="F01F258CFFC631729786FDC5BDB00533" box="[151,210,541,568]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="33" pageNumber="34" unit="m" value="48.0">48 m</quantity>
, and from Amirante,
<collectingCountry id="4FF0C8F9FFC6317296AFFDC5BF54053C" box="[446,566,541,567]" name="Seychelles" pageId="33" pageNumber="34">Seychelles</collectingCountry>
(with
<typeStatus id="E85C36CBFFC63172956FFDC6BF820533" box="[638,736,542,568]" pageId="33" pageNumber="34" type="syntype">syntypes</typeStatus>
BMNH 1921.11.7.16, and 1925.11.1.934 &amp; 1177). The typical variety from Tuticorin, Gulf of Mannar, approximate coordinates
<geoCoordinate id="52D3EEAEFFC631729484FD99BEBA0550" box="[917,984,577,603]" degrees="8.9" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="5555" value="8.9">8.9°N</geoCoordinate>
<geoCoordinate id="52D3EEAEFFC6317294F2FD99B9520550" box="[995,1072,577,603]" degrees="78.2" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5555" value="78.2">78.2°E</geoCoordinate>
(
<typeStatus id="E85C36CBFFC631729352FD99B9C50550" box="[1091,1191,577,603]" pageId="33" pageNumber="34" type="syntype">syntypes</typeStatus>
BMNH 1889.1.21.6, 4445 &amp; 78) as described by
<bibRefCitation id="5376F598FFC6317296FBFDBDBFD60574" author="Dendy, A." box="[490,692,613,639]" pageId="33" pageNumber="34" pagination="73 - 99" refId="ref93249" refString="Dendy, A. (1889) Report on a Second Collection of Sponges from the Gulf of Manaar. Annals and Magazine of Natural History, Series 6, 3, 73 - 99, pls. III - V. [https: // www. biodiversitylibrary. org / page / 15621866] https: // doi. org / 10.1080 / 00222938909460303" type="journal article" year="1889">Dendy (1889: 83)</bibRefCitation>
appears rather more similar to the
<taxonomicName id="F0E7F3EAFFC631729357FDBEB9CA0574" ID-CoL="5P4H4" authorityName="Dendy" authorityYear="1922" box="[1094,1192,613,640]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="variety" species="carnosa" variety="laxa">
var.
<emphasis id="0593547BFFC631729369FDBDB9CA0574" box="[1144,1192,613,639]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
</taxonomicName>
than to the specimen pictured and described by
<bibRefCitation id="5376F598FFC6317296D4FD51BFF405A8" author="Dendy, A." box="[453,662,649,676]" pageId="33" pageNumber="34" pagination="57 - 246" refId="ref93381" refString="Dendy, A. (1905) Report on the sponges collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, 3 (Supplement 18), pp. 57 - 246, pls. I - XVI. [https: // biodiversitylibrary. org / page / 1865481]" type="journal article" year="1905">Dendy (1905: 137</bibRefCitation>
, pl. VII fig. 5, from Cod Bay, Trincomalee,
<collectingCountry id="4FF0C8F9FFC631729380FD51B86605A8" box="[1169,1284,649,675]" name="Sri Lanka" pageId="33" pageNumber="34">Sri Lanka</collectingCountry>
, approximate coordinates
<geoCoordinate id="52D3EEAEFFC63172960EFD75BC0A05C3" box="[287,360,685,712]" degrees="8" direction="north" minutes="57" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="8.95">8°57N</geoCoordinate>
<geoCoordinate id="52D3EEAEFFC63172967EFD75BCA305CC" box="[367,449,685,711]" degrees="81.2" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5555" value="81.2">81.2°E</geoCoordinate>
), as it is more squat and its oxeas are indicated to become vestigial. Although the 1889 type description is not accompanied by an illustration of the habitus, it does not seem to be clearly different in shape and skeletal features from the
<taxonomicName id="F0E7F3EAFFC6317296F9FD2EBF2A0404" ID-CoL="5P4H4" authorityName="Dendy" authorityYear="1922" box="[488,584,757,784]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="variety" species="carnosa" variety="laxa">
var.
<emphasis id="0593547BFFC631729509FD2DBF2A0404" box="[536,584,757,783]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
</taxonomicName>
. I believe the three reports of this species by Dendy (twice named
<taxonomicName id="F0E7F3EAFFC631729220FD2DBD930438" ID-CoL="3FJTF" authorityName="Dendy" authorityYear="1889" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="carnosa">
<emphasis id="0593547BFFC631729220FD2DBD930438" italics="true" pageId="33" pageNumber="34">Gelliodes carnosa</emphasis>
</taxonomicName>
, once given the varietal name
<emphasis id="0593547BFFC631729544FCC1BFE70438" box="[597,645,793,819]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
) are all conspecific and represent variations of the same Central and Western Indian Ocean
<taxonomicName id="F0E7F3EAFFC631729686FCE5BF00045C" box="[407,610,829,855]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="undetermined">
<emphasis id="0593547BFFC631729686FCE5BF60045C" box="[407,514,829,855]" italics="true" pageId="33" pageNumber="34">Gelliodes</emphasis>
species.
</taxonomicName>
Accordingly, I propose here to merge the
<taxonomicName id="F0E7F3EAFFC63172932BFCE6B9F9045C" ID-CoL="5P4H4" authorityName="Dendy" authorityYear="1922" box="[1082,1179,829,856]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="variety" species="carnosa" variety="laxa">
var.
<emphasis id="0593547BFFC63172937AFCE5B9F9045C" box="[1131,1179,829,855]" italics="true" pageId="33" pageNumber="34">laxa</emphasis>
</taxonomicName>
and the typical variety into a single species,
<taxonomicName id="F0E7F3EAFFC631729697FCB9BF890470" ID-CoL="3FJTF" authority="Dendy, 1889" authorityName="Dendy" authorityYear="1889" box="[390,747,865,891]" class="Demospongiae" family="Niphatidae" genus="Gelliodes" kingdom="Animalia" order="Haplosclerida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="carnosa">
<emphasis id="0593547BFFC631729697FCB9BF300470" box="[390,594,865,891]" italics="true" pageId="33" pageNumber="34">Gelliodes carnosa</emphasis>
<bibRefCitation id="5376F598FFC631729548FCB9BF890470" author="Dendy, A." box="[601,747,865,891]" pageId="33" pageNumber="34" pagination="73 - 99" refId="ref93249" refString="Dendy, A. (1889) Report on a Second Collection of Sponges from the Gulf of Manaar. Annals and Magazine of Natural History, Series 6, 3, 73 - 99, pls. III - V. [https: // www. biodiversitylibrary. org / page / 15621866] https: // doi. org / 10.1080 / 00222938909460303" type="journal article" year="1889">Dendy, 1889</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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