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<document id="B7C9425B1309828B83D660B404840F88" ID-DOI="10.11646/zootaxa.3866.4.2" ID-GBIF-Dataset="4115b984-e9fd-44a6-ad86-05e88ab3b439" ID-ISSN="1175-5326" ID-Zenodo-Dep="249698" ID-ZooBank="D06E5477-4D5C-4402-B909-09A2AAFFB556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460919741933" checkinUser="plazi" docAuthor="Gapp, I. Wesley &amp; Lieberman, Bruce S." docDate="2014" docId="DC11878B3B47FFC8B1AF3F59E7D12001" docLanguage="en" docName="zt03866p498.pdf" docOrigin="Zootaxa 3866 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holmiella domackae Gapp &amp; Lieberman, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="492" masterDocId="2028FFF33B4DFFC5B1383913E4032219" masterDocTitle="New olenelloid trilobites from the Northwest Territories, Canada" masterLastPageNumber="498" masterPageNumber="479" pageNumber="489" updateTime="1698362066151" updateUser="plazi">
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<mods:title id="8A6846D081BB82EEFCCE2206388810FB">New olenelloid trilobites from the Northwest Territories, Canada</mods:title>
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<mods:namePart id="1BBA14F94C3B256F8892D36E188CC111">Gapp, I. Wesley</mods:namePart>
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<mods:namePart id="F4BE4BA4BE05DD418060BA8A45026036">Lieberman, Bruce S.</mods:namePart>
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<mods:date id="281E9220985BB1FBC22411060F47B483">2014</mods:date>
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<treatment id="DC11878B3B47FFC8B1AF3F59E7D12001" ID-DOI="http://doi.org/10.5281/zenodo.6136090" ID-GBIF-Taxon="119588945" ID-Zenodo-Dep="6136090" LSID="urn:lsid:plazi:treatment:DC11878B3B47FFC8B1AF3F59E7D12001" httpUri="http://treatment.plazi.org/id/DC11878B3B47FFC8B1AF3F59E7D12001" lastPageId="13" lastPageNumber="492" pageId="10" pageNumber="489">
<subSubSection id="1CA265163B47FFCFB1AF3F59E4DA249C" pageId="10" pageNumber="489" type="nomenclature">
<paragraph id="5407369D3B47FFCFB1AF3F59E5F5247C" blockId="10.[151,502,1610,1670]" box="[151,502,1610,1637]" pageId="10" pageNumber="489">
<heading id="0F4F81F13B47FFCFB1AF3F59E5F5247C" bold="true" box="[151,502,1610,1637]" fontSize="11" level="1" pageId="10" pageNumber="489" reason="1">
<emphasis id="66CCEA8F3B47FFCFB1AF3F59E5F5247C" bold="true" box="[151,502,1610,1637]" pageId="10" pageNumber="489">
<taxonomicName id="93B84D1E3B47FFCFB1AF3F59E593247D" box="[151,400,1610,1636]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="domackae" status="sp. nov.">
<emphasis id="66CCEA8F3B47FFCFB1AF3F59E593247D" bold="true" box="[151,400,1610,1636]" italics="true" pageId="10" pageNumber="489">Holmiella domackae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B47FFCFB0A03F58E5F5247C" box="[408,502,1611,1637]" pageId="10" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5407369D3B47FFCFB1AF3F7EE4DA249C" blockId="10.[151,502,1610,1670]" box="[151,217,1645,1670]" pageId="10" pageNumber="489">
<figureCitation id="CC832A183B47FFCFB1AF3F7EE4DA249C" box="[151,217,1645,1670]" captionStart="FIGURE 7" captionStartId="12.[151,250,1556,1578]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure@12.[151,1436,193,1534]" captionTargetPageId="12" captionText="FIGURE 7. Holmiella domackae sp. nov. 1, holotype PWNHC- 2013.23.23, x 3.2; 2, PWNHC 2013.23. 24 (latex cast), x 1.1; 3, KUMIP 355548, x 1.9." httpUri="https://zenodo.org/record/249705/files/figure.png" pageId="10" pageNumber="489">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1AF3FA6E16824D7" box="[151,1387,1717,1742]" pageId="10" pageNumber="489" type="materials_examined">
<paragraph id="5407369D3B47FFCFB1AF3FA6E16824D7" blockId="10.[151,1436,1717,1994]" box="[151,1387,1717,1742]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1AF3FA6E54624D7" bold="true" box="[151,325,1717,1742]" pageId="10" pageNumber="489">
<typeStatus id="8B03883F3B47FFCFB1AF3FA6E4D124D7" box="[151,210,1717,1742]" pageId="10" pageNumber="489">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8B03883F3B47FFCFB0743FA6E5BA24D7" box="[332,441,1717,1742]" pageId="10" pageNumber="489" type="holotype">Holotype</typeStatus>
: PWNHC-2013.23.23;
<typeStatus id="8B03883F3B47FFCFB3863FA5E72024D7" box="[702,803,1718,1742]" pageId="10" pageNumber="489" type="paratype">paratype</typeStatus>
:
<collectionCode id="32A9AE583B47FFCFB2153FA5E78424D4" box="[813,903,1718,1741]" pageId="10" pageNumber="489">KUMIP</collectionCode>
355548-355549, PWNHC-2013.23.2425.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3FCAE641250F" pageId="10" pageNumber="489" type="etymology">
<paragraph id="5407369D3B47FFCFB1FF3FCAE641250F" blockId="10.[151,1436,1717,1994]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3FCAE54D24EB" bold="true" box="[199,334,1753,1778]" pageId="10" pageNumber="489">Etymology.</emphasis>
In honour of Hamilton College paleontologist Cindy Domack, who had an important influence on the professional development of IWG.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3E32E7802547" pageId="10" pageNumber="489" type="diagnosis">
<paragraph id="5407369D3B47FFCFB1FF3E32E7802547" blockId="10.[151,1436,1717,1994]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3E32E5422523" bold="true" box="[199,321,1825,1850]" pageId="10" pageNumber="489">Diagnosis.</emphasis>
Cephalic border between intergenal spine and genal spine diverges anteriorly; genal spines have a sinuous shape; the posterolateral margins of LA diverge anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFC9B1FF3E7AE79925EF" lastPageId="12" lastPageNumber="491" pageId="10" pageNumber="489" type="description">
<paragraph id="5407369D3B47FFC9B1FF3E7AE79925EF" blockId="10.[151,1436,1717,1994]" lastBlockId="12.[151,1437,1652,2038]" lastPageId="12" lastPageNumber="491" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3E7AE55A259B" bold="true" box="[199,345,1897,1922]" pageId="10" pageNumber="489">Description.</emphasis>
Anterior border abaxial of sagittal line at an angle approximately 25 degrees to transverse line; length (sag.) anterior cephalic border directly anterior LA approximately 0.5 length of L3, developed as a flattened shelf; anterior portion of genal spines attached to cephalon at point directly abaxial to midpoint of LA along a sagittal line; genal spine deflected back rapidly and slightly outward creating an S-shape; length (sag.) LA approximately equal to length (sag.) of LO and L1; lateral margins of LA abaxial to lateral margins of LO; S3 convex anteriorly; ocular lobe separated from glabella by furrow; line between antero- and posteromedial tips of ocular lobe creates an angle with sagittal line of approximately 0-10 degrees; posterior ends of ocular lobes extend back to adaxial part of L1; width of ocular lobe at adaxial point about 60-70% length (sag.) of L1; width (tr.) of extraocular area opposite L1 approximately 5565% interocular area; S3 gently convex and does not conjoin adaxially; S2 gently convex and not conjoined adaxially; S1 gently convex and not conjoined adaxially; L1, L2, and L3 are approximately equal in length (sag.); lateral margins of L2, L3, and LA constricted posteriorly; width (tr.) of LO approximately equal to width of L3, length (sag.) approximately 1.5 times length (sag.) of L1; small spine on posterior margin of LO; intergenal spines positioned directly abaxial of occipital spine; posterior border between LO and intergenal spine directed back posteriorly abaxially.
</paragraph>
</subSubSection>
<caption id="00C766153B46FFCEB1AF3EB2E65B25CF" httpUri="https://zenodo.org/record/249704/files/figure.png" pageId="11" pageNumber="490" targetBox="[171,1416,193,1930]" targetPageId="11">
<paragraph id="5407369D3B46FFCEB1AF3EB2E65B25CF" blockId="11.[151,1436,1953,2006]" pageId="11" pageNumber="490">
<emphasis id="66CCEA8F3B46FFCEB1AF3EB2E51025AE" bold="true" box="[151,275,1953,1975]" pageId="11" pageNumber="490">FIGURE 6.</emphasis>
<taxonomicName id="93B84D1E3B46FFCEB0213EB2E5C625AE" box="[281,453,1953,1975]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="11" pageNumber="490" phylum="Ascomycota" rank="species" species="taurus" status="sp. nov.">
<emphasis id="66CCEA8F3B46FFCEB0213EB2E5C625AE" box="[281,453,1953,1975]" italics="true" pageId="11" pageNumber="490">Holmiella taurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B46FFCEB0F43EB2E61F25AE" box="[460,540,1953,1975]" pageId="11" pageNumber="490" rank="species">
<emphasis id="66CCEA8F3B46FFCEB0F43EB2E61F25AE" bold="true" box="[460,540,1953,1975]" pageId="11" pageNumber="490">sp. nov.</emphasis>
</taxonomicNameLabel>
1, holotype KUMIP 355545, x1.4; 2, PWNHC 2013.23.21 (latex cast), x2.1; 3, KUMIP 355546, x1.7; 4, PWNHC-2013.23.22, x0.9.
</paragraph>
</caption>
<caption id="00C766153B41FFC9B1AF3F07E5782451" httpUri="https://zenodo.org/record/249705/files/figure.png" pageId="12" pageNumber="491" targetBox="[151,1436,193,1534]" targetPageId="12">
<paragraph id="5407369D3B41FFC9B1AF3F07E5782451" blockId="12.[151,1436,1556,1608]" pageId="12" pageNumber="491">
<emphasis id="66CCEA8F3B41FFC9B1AF3F07E5102433" bold="true" box="[151,275,1556,1578]" pageId="12" pageNumber="491">FIGURE 7.</emphasis>
<taxonomicName id="93B84D1E3B41FFC9B0213F07E5EF2433" box="[281,492,1556,1578]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="12" pageNumber="491" phylum="Ascomycota" rank="species" species="domackae" status="sp. nov.">
<emphasis id="66CCEA8F3B41FFC9B0213F07E5EF2433" box="[281,492,1556,1578]" italics="true" pageId="12" pageNumber="491">Holmiella domackae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B41FFC9B0CA3F07E6402433" box="[498,579,1556,1578]" pageId="12" pageNumber="491" rank="species">
<emphasis id="66CCEA8F3B41FFC9B0CA3F07E6402433" bold="true" box="[498,579,1556,1578]" pageId="12" pageNumber="491">sp. nov.</emphasis>
</taxonomicNameLabel>
1, holotype PWNHC-2013.23.23, x3.2; 2, PWNHC 2013.23.24 (latex cast), x1.1; 3, KUMIP 355548, x1.9.
</paragraph>
</caption>
<subSubSection id="1CA265163B40FFC8B1FF3984E7D12001" pageId="13" pageNumber="492" type="discussion">
<paragraph id="5407369D3B40FFC8B1FF3984E6E923B5" blockId="13.[151,1436,151,536]" pageId="13" pageNumber="492">
<emphasis id="66CCEA8F3B40FFC8B1FF3984E54F22A9" bold="true" box="[199,332,151,176]" pageId="13" pageNumber="492">Discussion.</emphasis>
This species can be distinguished from
<taxonomicName id="93B84D1E3B40FFC8B210398AE79F22A9" box="[808,924,152,176]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="species" species="falcuta">
<emphasis id="66CCEA8F3B40FFC8B210398AE79F22A9" box="[808,924,152,176]" italics="true" pageId="13" pageNumber="492">H. falcuta</emphasis>
</taxonomicName>
and
<taxonomicName id="93B84D1E3B40FFC8B2E2398AE07722A9" box="[986,1140,153,176]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="species" species="preancora">
<emphasis id="66CCEA8F3B40FFC8B2E2398AE07722A9" box="[986,1140,153,176]" italics="true" pageId="13" pageNumber="492">H. preancora</emphasis>
</taxonomicName>
by its much wider genal spines, which are directed abaxially before being directed posteriorly (similarly to
<taxonomicName id="93B84D1E3B40FFC8B56E39AEE0A522CD" box="[1110,1190,188,212]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="species" species="falx">
<emphasis id="66CCEA8F3B40FFC8B56E39AEE0A522CD" box="[1110,1190,188,212]" italics="true" pageId="13" pageNumber="492">H. falx</emphasis>
</taxonomicName>
), and have a slightly sinuous shape. The posterior cephalic border between the intergenal spine and the genal spine diverges anteriorly in
<taxonomicName id="93B84D1E3B40FFC8B1AF3816E5282305" box="[151,299,260,284]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="species" species="domackae">
<emphasis id="66CCEA8F3B40FFC8B1AF3816E5282305" box="[151,299,260,284]" italics="true" pageId="13" pageNumber="492">H. domackae</emphasis>
</taxonomicName>
unlike in most other species of
<taxonomicName id="93B84D1E3B40FFC8B3B53817E6FE2305" box="[653,765,260,284]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="genus">
<emphasis id="66CCEA8F3B40FFC8B3B53817E6FE2305" box="[653,765,260,284]" italics="true" pageId="13" pageNumber="492">Holmiella</emphasis>
</taxonomicName>
(though see
<bibRefCitation id="30294B6C3B40FFC8B2B53817E0012305" author="Fritz" box="[909,1026,260,285]" pageId="13" pageNumber="492" refString="Fritz, W. H. (1972) Lower Cambrian trilobites from the Sekwi Formation type section, Mackenzie Mountains, northwestern Canada. Geological Survey of Canada, Bulletin, 212, 1 - 90." type="journal article" year="1972">Fritz 1972</bibRefCitation>
, pl. 7, figs. 7, 12), which are parallel to subparallel. Lastly, the posterolateral margin of LA diverges anteriorly unlike
<taxonomicName id="93B84D1E3B40FFC8B50E383AE0A12359" box="[1078,1186,297,320]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="13" pageNumber="492" phylum="Ascomycota" rank="species" species="taurus" status="sp. nov.">
<emphasis id="66CCEA8F3B40FFC8B50E383AE0A12359" box="[1078,1186,297,320]" italics="true" pageId="13" pageNumber="492">H. taurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B40FFC8B5933834E1042359" box="[1195,1287,295,320]" pageId="13" pageNumber="492" rank="species">
<emphasis id="66CCEA8F3B40FFC8B5933834E1042359" bold="true" box="[1195,1287,295,320]" pageId="13" pageNumber="492">sp. nov.</emphasis>
</taxonomicNameLabel>
This species may be biostratigraphically significant as it appears to represent the only species of holmiid that survives into the
<taxonomicName id="93B84D1E3B40FFC8B1AF3863E53E239E" authority="Zone" authorityName="Zone" box="[151,317,368,392]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="13" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B40FFC8B1AF3863E5002391" box="[151,259,368,392]" italics="true" pageId="13" pageNumber="492">Olenellus</emphasis>
zone
</taxonomicName>
or Dyeran stage and thus crosses the Montezuman-Dyeran stage boundary
<emphasis id="66CCEA8F3B40FFC8B5BC3862E0C12391" box="[1156,1218,369,392]" italics="true" pageId="13" pageNumber="492">sensu</emphasis>
<bibRefCitation id="30294B6C3B40FFC8B5F1387CE1792391" author="Webster" box="[1225,1402,367,392]" pageId="13" pageNumber="492" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian &quot; Lower Cambrian &quot;) in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
,
<bibRefCitation id="30294B6C3B40FFC8B4BD387CE1972391" author="Webster" box="[1413,1428,367,392]" pageId="13" pageNumber="492" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
) and Webster
<emphasis id="66CCEA8F3B40FFC8B0103886E56223B5" box="[296,353,404,428]" italics="true" pageId="13" pageNumber="492">et al.</emphasis>
(2011) (M. Webster, pers. comm.).
</paragraph>
<paragraph id="5407369D3B40FFC8B1FF38A4E7D12001" blockId="13.[151,1436,151,536]" pageId="13" pageNumber="492">
<emphasis id="66CCEA8F3B40FFC8B1FF38A4E55A23C9" bold="true" box="[199,345,439,464]" pageId="13" pageNumber="492">Occurrence.</emphasis>
<taxonomicName id="93B84D1E3B40FFC8B05A38ABE60923D6" authority="Zone" authorityName="Zone" box="[354,522,440,464]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="13" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B40FFC8B05A38ABE5CE23C9" box="[354,461,440,464]" italics="true" pageId="13" pageNumber="492">Olenellus</emphasis>
zone
</taxonomicName>
or Waucoban Series, Dyeran stage,
<emphasis id="66CCEA8F3B40FFC8B29038AAE7E523C9" box="[936,998,441,464]" italics="true" pageId="13" pageNumber="492">sensu</emphasis>
<bibRefCitation id="30294B6C3B40FFC8B2D638A4E0A223C9" author="Webster" box="[1006,1185,439,464]" pageId="13" pageNumber="492" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian &quot; Lower Cambrian &quot;) in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
,
<bibRefCitation id="30294B6C3B40FFC8B59638A4E0BE23C9" author="Webster" box="[1198,1213,439,464]" pageId="13" pageNumber="492" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
) and Webster
<emphasis id="66CCEA8F3B40FFC8B45A38AAE19F23C9" box="[1378,1436,440,464]" italics="true" pageId="13" pageNumber="492">et al.</emphasis>
(2011), early Cambrian, Sekwi Formation, Mackenzie Mountains, Northwest Territories,
<collectingCountry id="2CAF760D3B40FFC8B59A38CFE0FA23EC" box="[1186,1273,476,501]" name="Canada" pageId="13" pageNumber="492">Canada</collectingCountry>
, Section 2, in float, Section 3, 700
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above the base of section, Section
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float.
</paragraph>
</subSubSection>
</treatment>
</document>