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<document id="9376E2672DA68DDE351EAAF254E87359" ID-DOI="10.11646/zootaxa.3866.4.2" ID-GBIF-Dataset="4115b984-e9fd-44a6-ad86-05e88ab3b439" ID-ISSN="1175-5326" ID-Zenodo-Dep="249698" ID-ZooBank="D06E5477-4D5C-4402-B909-09A2AAFFB556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460919741933" checkinUser="plazi" docAuthor="Gapp, I. Wesley &amp; Lieberman, Bruce S." docDate="2014" docId="DC11878B3B45FFCCB1AF3AD2E19A2001" docLanguage="en" docName="zt03866p498.pdf" docOrigin="Zootaxa 3866 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elliptocephala jaredi Gapp &amp; Lieberman, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="488" masterDocId="2028FFF33B4DFFC5B1383913E4032219" masterDocTitle="New olenelloid trilobites from the Northwest Territories, Canada" masterLastPageNumber="498" masterPageNumber="479" pageNumber="487" updateTime="1698362066151" updateUser="plazi">
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<mods:title id="B197166A54A6BB1F7D30FD01041A311C">New olenelloid trilobites from the Northwest Territories, Canada</mods:title>
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<mods:name id="BE2D880859C49CEC2FEBA52ABFFD52DE" type="personal">
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<mods:namePart id="0D0B5204E9BF13AFAB6D21BA14A49A98">Gapp, I. Wesley</mods:namePart>
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<mods:namePart id="E71FE265CDD1A6A7643B9B9368CD2B63">Lieberman, Bruce S.</mods:namePart>
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<mods:title id="E03B0A823375C9956B05412B049D22D7">Zootaxa</mods:title>
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<mods:date id="541DE0FD8A92EEA8DEBC9095371526D2">2014</mods:date>
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<treatment id="DC11878B3B45FFCCB1AF3AD2E19A2001" ID-DOI="http://doi.org/10.5281/zenodo.6136080" ID-GBIF-Taxon="119588914" ID-Zenodo-Dep="6136080" LSID="urn:lsid:plazi:treatment:DC11878B3B45FFCCB1AF3AD2E19A2001" httpUri="http://treatment.plazi.org/id/DC11878B3B45FFCCB1AF3AD2E19A2001" lastPageId="9" lastPageNumber="488" pageId="8" pageNumber="487">
<subSubSection id="1CA265163B45FFCDB1AF3AD2E58321E4" pageId="8" pageNumber="487" type="nomenclature">
<paragraph id="5407369D3B45FFCDB1AF3AD2E5FA21C2" blockId="8.[151,505,961,1021]" box="[151,505,961,987]" pageId="8" pageNumber="487">
<heading id="0F4F81F13B45FFCDB1AF3AD2E5FA21C2" bold="true" box="[151,505,961,987]" fontSize="11" level="1" pageId="8" pageNumber="487" reason="1">
<emphasis id="66CCEA8F3B45FFCDB1AF3AD2E5FA21C2" bold="true" box="[151,505,961,987]" pageId="8" pageNumber="487">
<taxonomicName id="93B84D1E3B45FFCDB1AF3AD2E59721C2" box="[151,404,961,987]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="jaredi" status="sp. nov.">
<emphasis id="66CCEA8F3B45FFCDB1AF3AD2E59721C2" bold="true" box="[151,404,961,987]" italics="true" pageId="8" pageNumber="487">Elliptocephala jaredi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B45FFCDB0A33AD2E5FA21C2" box="[411,505,961,987]" pageId="8" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="5407369D3B45FFCDB1AF3AF7E58321E4" blockId="8.[151,505,961,1021]" box="[151,384,996,1021]" pageId="8" pageNumber="487">Fig. 1.1, 1.2, 1.4, 1.5</paragraph>
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<subSubSection id="1CA265163B45FFCDB1AF3D3FE145265C" box="[151,1350,1068,1093]" pageId="8" pageNumber="487" type="materials_examined">
<paragraph id="5407369D3B45FFCDB1AF3D3FE145265C" blockId="8.[151,1437,1068,2029]" box="[151,1350,1068,1093]" pageId="8" pageNumber="487">
<emphasis id="66CCEA8F3B45FFCDB1AF3D3FE546265C" bold="true" box="[151,325,1068,1093]" pageId="8" pageNumber="487">
<typeStatus id="8B03883F3B45FFCDB1AF3D3FE4D1265C" box="[151,210,1068,1093]" pageId="8" pageNumber="487">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8B03883F3B45FFCDB0743D3FE5B7265C" box="[332,436,1068,1093]" pageId="8" pageNumber="487" type="holotype">Holotype</typeStatus>
<collectionCode id="32A9AE583B45FFCDB0833D3EE616265D" box="[443,533,1069,1092]" pageId="8" pageNumber="487">KUMIP</collectionCode>
355534.
<typeStatus id="8B03883F3B45FFCDB3463D3EE6E8265C" box="[638,747,1069,1093]" pageId="8" pageNumber="487" type="paratype">Paratypes</typeStatus>
<collectionCode id="32A9AE583B45FFCDB3CA3D3EE74F265D" box="[754,844,1069,1092]" pageId="8" pageNumber="487">KUMIP</collectionCode>
355535-355544 and PWNHC-2013.23.6-16.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B45FFCDB1FF3D5CE73B2671" box="[199,824,1103,1128]" pageId="8" pageNumber="487" type="etymology">
<paragraph id="5407369D3B45FFCDB1FF3D5CE73B2671" blockId="8.[151,1437,1068,2029]" box="[199,824,1103,1128]" pageId="8" pageNumber="487">
<emphasis id="66CCEA8F3B45FFCDB1FF3D5CE54D2671" bold="true" box="[199,334,1103,1128]" pageId="8" pageNumber="487">Etymology.</emphasis>
In honour of Jared Lieberman, son of BSL.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B45FFCDB1FF3D67E68526CC" pageId="8" pageNumber="487" type="diagnosis">
<paragraph id="5407369D3B45FFCDB1FF3D67E68526CC" blockId="8.[151,1437,1068,2029]" pageId="8" pageNumber="487">
<emphasis id="66CCEA8F3B45FFCDB1FF3D67E5422694" bold="true" box="[199,321,1140,1165]" pageId="8" pageNumber="487">Diagnosis.</emphasis>
LA contacts anterior cephalic margin; LA approximately 1.5 times combined length of LO and L1; glabellar furrows not conjoined; S3 strongly convex; ocular lobes terminate posteriorly opposite SO; posterior cephalic margin parallel to a transverse line.
</paragraph>
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<subSubSection id="1CA265163B45FFCDB1FF3DCCE7E324D4" pageId="8" pageNumber="487" type="description">
<paragraph id="5407369D3B45FFCDB1FF3DCCE7E324D4" blockId="8.[151,1437,1068,2029]" pageId="8" pageNumber="487">
<emphasis id="66CCEA8F3B45FFCDB1FF3DCCE55A26E1" bold="true" box="[199,345,1247,1272]" pageId="8" pageNumber="487">Description.</emphasis>
Anterior cephalic margin forms smooth, semi-circular arc; length of anterior cephalic border is short to moderately long (length = 0.51 times length [sag.] of LO); LA contacts anterior cephalic margin; anterior margins of the frontal lobes at each side of the midline deflected posteriorly at roughly 40 degree angle relative to a transverse line; length (sag.) of LA long, equal to 1.5 times length of LO and L1 adaxially; LA does not expand prominently dorsally; lateral margins of LA proximal to lateral margins of LO; ocular lobes contact frontal lobe at posterolateral margins of frontal lobe; ocular furrow present; line from posterior tip of the ocular lobe to junction of posterior margin of lobe with glabella forms 10-20 degree angle with sagittal line; posterior tips of the ocular lobes developed opposite SO; anterodistal margins of L3 formed by ocular lobes; SO, S1, and S2 are convex, S3 very convex; none of the axial furrows are conjoined adaxially; L2 and L3 do not merge abaxially; lateral lobes present on LO; extraocular area approximately two-thirds width of glabella, gently convex; polygonal fracturing visible on some cephala; posterior cephalic border parallel with transverse line or directed slightly posteriorly; intergenal spine and intergenal angle not present; genal spines directed posteriorly approximately 10 degrees relative to a sagittal line, extend back approximately three thoracic segments (inferred based on typical relative length [sag.] of LO as compared to typical relative length [sag.] of prothoracic axial rings).
</paragraph>
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<subSubSection id="1CA265163B45FFCDB1FF3FC4E6EB25D0" pageId="8" pageNumber="487" type="discussion">
<paragraph id="5407369D3B45FFCDB1FF3FC4E6EB25D0" blockId="8.[151,1437,1068,2029]" pageId="8" pageNumber="487">
<emphasis id="66CCEA8F3B45FFCDB1FF3FC4E54F24E9" bold="true" box="[199,332,1751,1776]" pageId="8" pageNumber="487">Discussion.</emphasis>
This species is assigned to the genus
<taxonomicName id="93B84D1E3B45FFCDB2393FCBE7A524E9" box="[769,934,1752,1776]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B45FFCDB2393FCBE7A524E9" box="[769,934,1752,1776]" italics="true" pageId="8" pageNumber="487">Elliptocephala</emphasis>
</taxonomicName>
on the basis of character states such as: the length of LA (sag.) is approximately 1.5 times the length (sag.) of LO and L1; the anterior margins of the frontal lobe at each side of the midline deflects posteriorly at approximately 40 degrees relative to a transverse line; L2 and L3 do not merge abaxially; the abaxial margins of L2 diverge anteriorly; S2 is convex anteriorly; the ocular lobe has a prominent ocular furrow; the line from the posterior tip of the ocular lobe to the junction of the lobe with the glabella forms a 10 to 20 degree angle with the sagittal line; the anterodistal margins of L3 are formed by the ocular lobes; and polygonal fracturing is visible on cephala.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B45FFCCB1FF3EC7E19A2001" lastPageId="9" lastPageNumber="488" pageId="8" pageNumber="487" type="reference_group">
<paragraph id="5407369D3B45FFCCB1FF3EC7E73D23B5" blockId="8.[151,1437,1068,2029]" lastBlockId="9.[151,1437,151,536]" lastPageId="9" lastPageNumber="488" pageId="8" pageNumber="487">
<taxonomicName id="93B84D1E3B45FFCDB1FF3EC7E5B425F5" box="[199,439,2004,2028]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="jaredi">
<emphasis id="66CCEA8F3B45FFCDB1FF3EC7E5B425F5" box="[199,439,2004,2028]" italics="true" pageId="8" pageNumber="487">Elliptocephala jaredi</emphasis>
</taxonomicName>
shares many similarities with
<taxonomicName id="93B84D1E3B45FFCDB2223EC6E0C925F5" authority="Wanner, 1901" authorityName="Wanner" authorityYear="1901" box="[794,1226,2004,2028]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="walcottana">
<emphasis id="66CCEA8F3B45FFCDB2223EC6E00E25F5" box="[794,1037,2004,2028]" italics="true" pageId="8" pageNumber="487">Wanneria walcottana</emphasis>
(
<bibRefCitation id="30294B6C3B45FFCDB5253EC7E0C225F5" author="Wanner" box="[1053,1217,2004,2028]" pageId="8" pageNumber="487" refString="Wanner, A. (1901) A new species of Olenellus from the Lower Cambrian of York County, Pennsylvania. Proceedings of the Washington Academy of Sciences, 3, 267 - 272." type="journal article" year="1901">Wanner, 1901</bibRefCitation>
)
</taxonomicName>
. This includes the fact that none of the glabellar furrows are conjoined adaxially, S3 is strongly convex, there is an absence of anastomosing ridges on the extraocular area, the frontal lobe contacts the anterior cephalic border, and the abaxial margins of the glabella at L1 relative to LO are constricted. Differences include the fact that the ocular lobes extend further posteriorly in
<taxonomicName id="93B84D1E3B44FFCCB0B13816E5E82305" box="[393,491,260,284]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="species" species="jaredi">
<emphasis id="66CCEA8F3B44FFCCB0B13816E5E82305" box="[393,491,260,284]" italics="true" pageId="9" pageNumber="488">E. jaredi</emphasis>
</taxonomicName>
, as well as the presence of lateral lobes, shorter genal spines, LA not bulging much further laterally than LO, and the length of LA relative to LO and L1 is longer in
<taxonomicName id="93B84D1E3B44FFCCB509383AE0902359" box="[1073,1171,296,320]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="species" species="jaredi">
<emphasis id="66CCEA8F3B44FFCCB509383AE0902359" box="[1073,1171,296,320]" italics="true" pageId="9" pageNumber="488">E. jaredi</emphasis>
</taxonomicName>
. Due to the similarities between
<taxonomicName id="93B84D1E3B44FFCCB039385EE56A237D" box="[257,361,332,356]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="species" species="jaredi">
<emphasis id="66CCEA8F3B44FFCCB039385EE56A237D" box="[257,361,332,356]" italics="true" pageId="9" pageNumber="488">E. jaredi</emphasis>
</taxonomicName>
and
<taxonomicName id="93B84D1E3B44FFCCB094385EE651237D" box="[428,594,332,356]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="species" species="walcottana">
<emphasis id="66CCEA8F3B44FFCCB094385EE651237D" box="[428,594,332,356]" italics="true" pageId="9" pageNumber="488">W. walcottana</emphasis>
</taxonomicName>
, it is likely that
<taxonomicName id="93B84D1E3B44FFCCB21F385EE78C237D" box="[807,911,332,356]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="species" species="jaredi">
<emphasis id="66CCEA8F3B44FFCCB21F385EE78C237D" box="[807,911,332,356]" italics="true" pageId="9" pageNumber="488">E. jaredi</emphasis>
</taxonomicName>
is positioned basally within
<taxonomicName id="93B84D1E3B44FFCCB5C8385FE196237D" box="[1264,1429,332,356]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B44FFCCB5C8385FE196237D" box="[1264,1429,332,356]" italics="true" pageId="9" pageNumber="488">Elliptocephala</emphasis>
</taxonomicName>
. However, further phylogenetic investigation is needed to test this, and the fact that LA does not bulge much further laterally than LO is somewhat distinctive for
<taxonomicName id="93B84D1E3B44FFCCB3AB3887E73B23B5" box="[659,824,404,428]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B44FFCCB3AB3887E73B23B5" box="[659,824,404,428]" italics="true" pageId="9" pageNumber="488">Elliptocephala</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5407369D3B44FFCCB1FF38A4E19A2001" blockId="9.[151,1437,151,536]" pageId="9" pageNumber="488">
<emphasis id="66CCEA8F3B44FFCCB1FF38A4E55A23C9" bold="true" box="[199,345,439,464]" pageId="9" pageNumber="488">Occurrence.</emphasis>
<taxonomicName id="93B84D1E3B44FFCCB05A38ABE60923D6" ID-CoL="67KY" authority="Zone" authorityName="Zone" box="[354,522,440,464]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="9" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B44FFCCB05A38ABE5CE23C9" box="[354,461,440,464]" italics="true" pageId="9" pageNumber="488">Olenellus</emphasis>
zone
</taxonomicName>
or Waucoban Series, Dyeran stage,
<emphasis id="66CCEA8F3B44FFCCB29038AAE7E523C9" box="[936,998,441,464]" italics="true" pageId="9" pageNumber="488">sensu</emphasis>
<bibRefCitation id="30294B6C3B44FFCCB2D638A4E0A223C9" author="Webster" box="[1006,1185,439,464]" pageId="9" pageNumber="488" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian &quot; Lower Cambrian &quot;) in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
,
<bibRefCitation id="30294B6C3B44FFCCB59638A4E0BE23C9" author="Webster" box="[1198,1213,439,464]" pageId="9" pageNumber="488" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
) and Webster
<emphasis id="66CCEA8F3B44FFCCB45A38AAE19F23C9" box="[1378,1436,440,464]" italics="true" pageId="9" pageNumber="488">et al.</emphasis>
(2011), early Cambrian, Sekwi Formation, Mackenzie Mountains, Northwest Territories,
<collectingCountry id="2CAF760D3B44FFCCB58438CFE11023EC" box="[1212,1299,476,501]" name="Canada" pageId="9" pageNumber="488">Canada</collectingCountry>
, Section 2, 260
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above the base of section, Section 3, 700
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above the base of the section, and Section 4, in float).
</paragraph>
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