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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.138.38721" ID-PMC="PMC6969003" ID-Pensoft-Pub="1314-2003-138-203" ID-Pensoft-UUID="37174C2A6257592B83BED9D2E2CAA79B" ID-PubMed="31988614" ModsDocID="1314-2003-138-203" checkinTime="1578758105156" checkinUser="pensoft" docAuthor="Maw, Mya Bhone, Ding, Hong-Bo, Yang, Bin, Win, Pyae Pyae &amp; Tan, Yun-Hong" docDate="2020" docId="CD6E03078D3950DEB17E0D65E0A617A0" docLanguage="en" docName="PhytoKeys 138: 203-217" docOrigin="PhytoKeys 138" docSource="http://dx.doi.org/10.3897/phytokeys.138.38721" docTitle="Begonia chenii Y. H. Tan, M. B. Maw &amp; H. B. Ding 2020, sp. nov." docType="treatment" docVersion="5" id="37174C2A6257592B83BED9D2E2CAA79B" lastPageNumber="203" masterDocId="37174C2A6257592B83BED9D2E2CAA79B" masterDocTitle="Taxonomic studies on Begonia (Begoniaceae) in Myanmar I: three new species and supplementary description of Begonia rheophytica from Northern Myanmar" masterLastPageNumber="217" masterPageNumber="203" pageNumber="203" updateTime="1668139614508" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic studies on Begonia (Begoniaceae) in Myanmar I: three new species and supplementary description of Begonia rheophytica from Northern Myanmar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maw, Mya Bhone</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ding, Hong-Bo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Bin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Win, Pyae Pyae</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tan, Yun-Hong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>138</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>203</mods:start>
<mods:end>217</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.138.38721</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.138.38721</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-138-203</mods:identifier>
<mods:identifier type="Pensoft-UUID">37174C2A6257592B83BED9D2E2CAA79B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161672084" LSID="urn:lsid:plazi:treatment:CD6E03078D3950DEB17E0D65E0A617A0" httpUri="http://treatment.plazi.org/id/CD6E03078D3950DEB17E0D65E0A617A0" lastPageNumber="203" pageId="0" pageNumber="203">
<subSubSection pageId="0" pageNumber="203" type="nomenclature">
<paragraph pageId="0" pageNumber="203">
<taxonomicName LSID="CD6E0307-8D39-50DE-B17E-0D65E0A617A0" authority="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii" status="sp. nov.">Begonia chenii Y.H.Tan, M.B.Maw &amp; H.B.Ding</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="203">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Begonia chenii Y. H. Tan, M. B. Maw &amp; H. B. Ding, sp. nov. (photographed by H. B. Ding and Y. H. Tan) A habitat B leaves (back view) C inflorescence D staminate flower showing 6 tepals E pistillate flower F infructescence showing monoecious G infructescence showing stigmas H staminate flower showing variation of 4, 5, 6 tepals I tepals of staminate flower J androecium K tepals of pistillate flower L ovary and stigma M ovary (3 - winged) N ovary (4 - winged) O-P serial cross section of ovary (locules 4) Q serial cross section of ovary (locules 4 and wingless) R serial cross section of ovary (locules 3)." figureDoi="10.3897/phytokeys.138.38721.figure1" httpUri="https://binary.pensoft.net/fig/370289" pageId="0" pageNumber="203">Figure 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="diagnosis">
<paragraph pageId="0" pageNumber="203">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="203">
<taxonomicName authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">Begonia chenii</emphasis>
</taxonomicName>
Y.H. Tan, M.B. Maw &amp; H.B. Ding is mostly similar to
<taxonomicName lsidName="B. mariachristinae" pageId="0" pageNumber="203" rank="species" species="mariachristinae">
<emphasis italics="true" pageId="0" pageNumber="203">B. mariachristinae</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.1017/S0960428618000021" author="Wahlsteen, E" journalOrPublisher="Edinburgh Journal of Botany" pageId="0" pageNumber="203" pagination="161 - 166" refId="B17" refString="Wahlsteen, E, 2018. Begonia mariachristinae (Begoniaceae), a new species from Northern Myanmar. Edinburgh Journal of Botany 75 (2): 161 - 166, DOI: https://doi.org/10.1017/S0960428618000021" title="Begonia mariachristinae (Begoniaceae), a new species from Northern Myanmar." url="https://doi.org/10.1017/S0960428618000021" volume="75" year="2018">Wahlsteen (2018</bibRefCitation>
: 1) in lanceolate-ovate leaves with silver patches or dots on the upper surface, but significantly differs by stipules slightly pilose (
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
glabrous), petiole densely reddish pilose (
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
sparsely puberulous), 6 (rarely 4 or 7) tepals of female flower (
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
4 tepals) and red, 3 or 4 locular ovary (
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
green, 2 locular).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="materials_examined">
<paragraph pageId="0" pageNumber="203">Type.</paragraph>
<paragraph pageId="0" pageNumber="203">
Myanmar. Kachin State: Putao District, on the way from Putao to Upper Shankhaung, in tropical rain forest,
<geoCoordinate degrees="27" direction="north" minutes="25" orientation="latitude" precision="1" seconds="36.87" value="27.426908">27°25'36.87&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="16" orientation="longitude" precision="1" seconds="13.56" value="97.27044">97°16'13.56&quot;E</geoCoordinate>
, 512 m, 4 May 2017,
<emphasis italics="true" pageId="0" pageNumber="203">Y.H. Tan</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">B. Yang</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">H.B. Ding</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">X.D. Zeng</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">M.B. Maw &amp; T.S. Tin M1378</emphasis>
(holotype: HITBC!; isotypes: RAF!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="description">
<paragraph pageId="0" pageNumber="203">Description.</paragraph>
<paragraph pageId="0" pageNumber="203">
Perennial herb, dioecious or rarely monoecious, lacking rhizome or tuber.
<emphasis bold="true" pageId="0" pageNumber="203">Stem</emphasis>
erect, 40-60 cm tall, reddish brown, densely white pilose, internode 2-11 cm long, branching.
<emphasis bold="true" pageId="0" pageNumber="203">Stipule</emphasis>
persistent, ovate, 1-15
<normalizedToken originalValue="×">x</normalizedToken>
3-5 mm, papery, keeled, apex cuspidate (1-4 mm), margin entire, slightly pilose.
<emphasis bold="true" pageId="0" pageNumber="203">Leaf</emphasis>
alternate, petiole 1.5-3 cm long, reddish-brown, densely reddish pilose;
<emphasis bold="true" pageId="0" pageNumber="203">blade</emphasis>
asymmetric, lanceolate-ovate, 8-11
<normalizedToken originalValue="×">x</normalizedToken>
2.5-4 cm, apex attenuate, base oblique, margin serrate and with reddish hispid, venation palmate-pinnate, 5-6 pairs of veins;
<emphasis bold="true" pageId="0" pageNumber="203">upper surface</emphasis>
green or dark green with white patches and dots between the veins, bright green shot with metallic blue depending on the angle of the light, especially on young leaves, sparsely reddish hispid, especially along the midrib and lateral veins;
<emphasis bold="true" pageId="0" pageNumber="203">lower surface</emphasis>
deep red or deep red with light green areas both margin linings, scattered reddish hispid and densely along the midrib and lateral veins.
<emphasis bold="true" pageId="0" pageNumber="203">Inflorescence</emphasis>
axillary, flower solitary or in a simple cyme, pendulous;
<emphasis bold="true" pageId="0" pageNumber="203">bract</emphasis>
persistent, ovate to narrow lanceolate, 4-8
<normalizedToken originalValue="×">x</normalizedToken>
2-3 mm.
<emphasis bold="true" pageId="0" pageNumber="203">Staminate flower</emphasis>
: pedicel 0.8-1.1 cm, reddish, glabrous or sparsely pilose; tepals 4 (rarely 5 or 6), reddish with white margins, unequal, inner 2 (rarely 3), ovate, 7-1
<normalizedToken originalValue="×">x</normalizedToken>
6-1 mm, glabrous, outer 2 (rarely 3), ovate, 7-9
<normalizedToken originalValue="×">x</normalizedToken>
7-1 mm, reddish or whitish pilose on the outer surface, margin entire; androecium actinomorphic, stamens numerous, filament free, anther oblong, golden yellow.
<emphasis bold="true" pageId="0" pageNumber="203">Pistillate flower</emphasis>
: pedicel 0.6-0.8 cm, tepals 6 (rarely 4 or 7), unequal, inner 3, elliptic, 8-10
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm, pink to white, glabrous, outer 3, ovate or elliptic, 7-11
<normalizedToken originalValue="×">x</normalizedToken>
5-7 mm, reddish or whitish pilose on outer surface; ovary red, slightly or densely pilose on the surface, triangular or rhomboid winged, 3-5
<normalizedToken originalValue="×">x</normalizedToken>
2-5 mm, placentation axillary, locules 3 or 4, placentae 2 per locule; styles 3, fused at base, stigma bifid with twisted bands, highly convolute, yellow or golden yellow.
<emphasis bold="true" pageId="0" pageNumber="203">Fruit</emphasis>
berry-like, red, reddish or whitish pilose, triangular, rhomboid (8-15
<normalizedToken originalValue="×">x</normalizedToken>
6-9 mm) or suboblate (8-15 mm in diam.), 3 or 4 horned, rarely wingless.
</paragraph>
<caption doi="10.3897/phytokeys.138.38721.figure1" httpUri="https://binary.pensoft.net/fig/370289" pageId="0" pageNumber="203" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="203">
<emphasis bold="true" pageId="0" pageNumber="203">Figure 1.</emphasis>
<taxonomicName authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">Begonia chenii</emphasis>
</taxonomicName>
Y.H.Tan, M.B.Maw &amp; H.B.Ding, sp. nov. (photographed by H.B. Ding and Y.H. Tan)
<emphasis bold="true" pageId="0" pageNumber="203">A</emphasis>
habitat
<emphasis bold="true" pageId="0" pageNumber="203">B</emphasis>
leaves (back view)
<emphasis bold="true" pageId="0" pageNumber="203">C</emphasis>
inflorescence
<emphasis bold="true" pageId="0" pageNumber="203">D</emphasis>
staminate flower showing 6 tepals
<emphasis bold="true" pageId="0" pageNumber="203">E</emphasis>
pistillate flower
<emphasis bold="true" pageId="0" pageNumber="203">F</emphasis>
infructescence showing monoecious
<emphasis bold="true" pageId="0" pageNumber="203">G</emphasis>
infructescence showing stigmas
<emphasis bold="true" pageId="0" pageNumber="203">H</emphasis>
staminate flower showing variation of 4, 5, 6 tepals
<emphasis bold="true" pageId="0" pageNumber="203">I</emphasis>
tepals of staminate flower
<emphasis bold="true" pageId="0" pageNumber="203">J</emphasis>
androecium
<emphasis bold="true" pageId="0" pageNumber="203">K</emphasis>
tepals of pistillate flower
<emphasis bold="true" pageId="0" pageNumber="203">L</emphasis>
ovary and stigma
<emphasis bold="true" pageId="0" pageNumber="203">M</emphasis>
ovary (3-winged)
<emphasis bold="true" pageId="0" pageNumber="203">N</emphasis>
ovary (4-winged)
<emphasis bold="true" pageId="0" pageNumber="203">O-P</emphasis>
serial cross section of ovary (locules 4)
<emphasis bold="true" pageId="0" pageNumber="203">Q</emphasis>
serial cross section of ovary (locules 4 and wingless)
<emphasis bold="true" pageId="0" pageNumber="203">R</emphasis>
serial cross section of ovary (locules 3).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="phenology">
<paragraph pageId="0" pageNumber="203">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="203">Flowering from April to May; fruiting from May to June.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="distribution">
<paragraph pageId="0" pageNumber="203">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="203">The species is only known from the type locality, Putao District, Kachin State, Northern Myanmar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="ecology">
<paragraph pageId="0" pageNumber="203">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="203">The species grows in the moist shaded environment of tropical rain forest, elevation about 512 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="etymology">
<paragraph pageId="0" pageNumber="203">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="203">
The species epithet &quot;
<emphasis italics="true" pageId="0" pageNumber="203">chenii</emphasis>
&quot; is named after Professor Chen Jin, the director of Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, who gave us the opportunity to study the Myanmar flora, which led to the discovery of this new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="conservation status">
<paragraph pageId="0" pageNumber="203">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="203">
Data Deficient (DD).
<taxonomicName authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">Begonia chenii</emphasis>
</taxonomicName>
was collected along the path on the way from Putao to Upper Shankhaung where any signs of major anthropogenic disturbance were noticed in the type locality. However, further explorations are needed for a proper assessment of conservation due to insufficient information on its distribution and population status. Therefore, the species has been preliminarily assigned to Data Deficient (DD) category according to The Guidelines for Using The IUCN Red List Categories and Criteria (
<bibRefCitation author="IUCN" journalOrPublisher="Botanical Bulletin of Academia Sinica" pageId="0" pageNumber="203" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B16" refString="IUCN, 2017. Guidelines for using the IUCN red list categories and criteria. version 13. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN red list categories and criteria. version 13. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="203" type="materials_examined">
<paragraph pageId="0" pageNumber="203">Additional specimens examined</paragraph>
<paragraph pageId="0" pageNumber="203">
<emphasis bold="true" pageId="0" pageNumber="203">(paratypes).</emphasis>
Myanmar. Kachin State: Putao District, Upper Shankhaung, in tropical montane forest,
<geoCoordinate degrees="27" direction="north" minutes="25" orientation="latitude" precision="1" seconds="36.87" value="27.426908">27°25'36.87&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="16" orientation="longitude" precision="1" seconds="13.56" value="97.27044">97°16'13.56&quot;E</geoCoordinate>
, 512 m, 4 May 2017,
<emphasis italics="true" pageId="0" pageNumber="203">Y.H. Tan</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">B. Yang</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">H.B. Ding</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">X.D. Zeng</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="203">M.B. Maw &amp; T.S. Tin M1379</emphasis>
(HITBC!); Kachin State: Putao District, Upper Shankhaung,
<geoCoordinate degrees="27" direction="north" minutes="25" orientation="latitude" precision="15" seconds="35" value="27.426388">27°25'35&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="16" orientation="longitude" precision="15" seconds="14" value="97.27056">97°16'14&quot;E</geoCoordinate>
, 500 m, 29 April 2016,
<emphasis italics="true" pageId="0" pageNumber="203">Y.H. Tan &amp; S.S. Zhou M201627</emphasis>
(HITBC!); Kachin State: Putao District, Upper Shankhaung,
<geoCoordinate degrees="27" direction="north" minutes="25" orientation="latitude" precision="15" seconds="34" value="27.426111">27°25'34&quot;N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="16" orientation="longitude" precision="15" seconds="13" value="97.27028">97°16'13&quot;E</geoCoordinate>
, 520 m, 7 May 2017,
<emphasis italics="true" pageId="0" pageNumber="203">S.S. Zhou &amp; X.D. Zeng M2030</emphasis>
(HITBC!; RAF!)
</paragraph>
<paragraph pageId="0" pageNumber="203">
<emphasis bold="true" pageId="0" pageNumber="203">Affinities.</emphasis>
<taxonomicName authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">Begonia chenii</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName lsidName="B. mariachristinae" pageId="0" pageNumber="203" rank="species" species="mariachristinae">
<emphasis italics="true" pageId="0" pageNumber="203">B. mariachristinae</emphasis>
</taxonomicName>
. But it can be easily distinguished in having 3 or 4 locules (
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
2 locules). See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of key morphological characters of Begonia chenii and B. mariachristinae." httpUri="http://table.plazi.org/id/63A2BA0F873B292CBD3DD4D56F075126" pageId="0" pageNumber="203" tableUuid="63A2BA0F873B292CBD3DD4D56F075126">1</tableCitation>
for detailed comparison of
<taxonomicName lsidName="B. chenii" pageId="0" pageNumber="203" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">B. chenii</emphasis>
</taxonomicName>
with morphologically allied species.
</paragraph>
<caption ID-Table-UUID="63A2BA0F873B292CBD3DD4D56F075126" httpUri="http://table.plazi.org/id/63A2BA0F873B292CBD3DD4D56F075126" pageId="0" pageNumber="203" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="203">
<emphasis bold="true" pageId="0" pageNumber="203">Table 1.</emphasis>
Comparison of key morphological characters of
<taxonomicName authorityName="Y. H. Tan, M. B. Maw &amp; H. B. Ding" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia chenii" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis italics="true" pageId="0" pageNumber="203">Begonia chenii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="B. mariachristinae" pageId="0" pageNumber="203" rank="species" species="mariachristinae">
<emphasis italics="true" pageId="0" pageNumber="203">B. mariachristinae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="203">
<table pageId="0" pageNumber="203">
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">
<emphasis bold="true" pageId="0" pageNumber="203">Attributes</emphasis>
</td>
<td pageId="0" pageNumber="203">
<taxonomicName lsidName="B. chenii" pageId="0" pageNumber="203" rank="species" species="chenii">
<emphasis bold="true" italics="true" pageId="0" pageNumber="203">B. chenii</emphasis>
</taxonomicName>
</td>
<td pageId="0" pageNumber="203">
<taxonomicName lsidName="B. mariachristinae" pageId="0" pageNumber="203" rank="species" species="mariachristinae">
<emphasis bold="true" italics="true" pageId="0" pageNumber="203">B. mariachristinae</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Stem</td>
<td pageId="0" pageNumber="203">40-60 cm tall, reddish brown densely white pilose</td>
<td pageId="0" pageNumber="203">40-60 cm tall dark red to maroon, hairs</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Stipules</td>
<td pageId="0" pageNumber="203">persistent, slightly pilose</td>
<td pageId="0" pageNumber="203">persistent, glabrous</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Petiole</td>
<td pageId="0" pageNumber="203">1.5-3 cm long, densely reddish pilose</td>
<td pageId="0" pageNumber="203">1-5 cm long, sparsely puberulous</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Leaves</td>
<td pageId="0" pageNumber="203">
lanceolate-ovate, 8-11
<normalizedToken originalValue="×">x</normalizedToken>
2.5-4 cm
</td>
<td pageId="0" pageNumber="203">
lanceolate-ovate 6-11.5
<normalizedToken originalValue="×">x</normalizedToken>
2.5-4.5 cm
</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Upper surface</td>
<td pageId="0" pageNumber="203">sparsely reddish hispid especially along the midrib and lateral veins</td>
<td pageId="0" pageNumber="203">slightly reddish hispid</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Lower surface</td>
<td pageId="0" pageNumber="203">scattered reddish hispid and densely along the midrib and lateral veins</td>
<td pageId="0" pageNumber="203">slightly hairy along midrib</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Male flower</td>
<td pageId="0" pageNumber="203">tepals 4 (rarely 5 or 6) reddish with white linings</td>
<td pageId="0" pageNumber="203">tepals 4 pink or white</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Female flower</td>
<td pageId="0" pageNumber="203">tepals 6 (rarely 4 or 7) pink to white</td>
<td pageId="0" pageNumber="203">tepals 4 pink or white</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Ovary</td>
<td pageId="0" pageNumber="203">locules 3 or 4 red, slightly or densely pilose</td>
<td pageId="0" pageNumber="203">locules 2 green, hispid</td>
</tr>
<tr pageId="0" pageNumber="203">
<td pageId="0" pageNumber="203">Style</td>
<td pageId="0" pageNumber="203">3</td>
<td pageId="0" pageNumber="203">2 (or 3)</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>