469 lines
49 KiB
XML
469 lines
49 KiB
XML
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<mods:title id="84C5123BCD604A19FF175628E60C1417">A revision of the genus Mesobuthus Vachon 1950 with a description of 14 new species (Scorpiones Buthidae)</mods:title>
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(Figures 1120–1154, 1155, 1158,
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(in part);
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<bibRefCitation id="621019E810C2F1B993E5520220A9FACE" author="GANTENBEIN" box="[161,510,1329,1353]" pageId="164" pageNumber="163" pagination="412 - 420" refId="ref84304" refString="GANTENBEIN, B., V. FET & A. V. GROMOV. 2003. The first DNA phylogeny of four species of Mesobuthus Vachon, 1950 (Scorpiones: Buthidae) from Eurasia. Journal of Arachnology, 31: 412 - 420." type="journal article" year="2003">Gantenbein et al., 2003: 413</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part;
|
||
<collectingCountry id="7E96248910C2F1B991C052022257FACE" box="[644,768,1329,1353]" name="Uzbekistan" pageId="164" pageNumber="163">Uzbekistan</collectingCountry>
|
||
: Zarafshan).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E9B379210C2F1B9933552A123F8F9CE" pageId="164" pageNumber="163" type="materials_examined">
|
||
<paragraph id="063E641910C2F1B9933552A123F8F9CE" blockId="164.[113,771,1425,1609]" pageId="164" pageNumber="163">
|
||
<materialsCitation id="B6E96E4410C2F1B9933552A123A8FA6E" collectingDate="1998-04-22" collectionCode="TYPE, MATERIAL" collectorName="A. Gromov" country="Uzbekistan" latitude="41.6" location="Tamdy District" longLatPrecision="693" longitude="64.18" pageId="164" pageNumber="163" specimenCount="1" specimenCount-male="1" stateProvince="Navoi Province" typeStatus="holotype">
|
||
<collectionCode id="6090FCDC10C2F1B9933552A12074FA2E" box="[113,291,1425,1449]" pageId="164" pageNumber="163">TYPE MATERIAL</collectionCode>
|
||
.
|
||
<emphasis id="34F5B80B10C2F1B9920652A22096FA2E" bold="true" box="[322,449,1425,1449]" pageId="164" pageNumber="163">
|
||
<collectingCountry id="7E96248910C2F1B9920652A22096FA2E" box="[322,449,1425,1449]" name="Uzbekistan" pageId="164" pageNumber="163">Uzbekistan</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="34F5B80B10C2F1B992A452A223CCFA2E" bold="true" box="[480,667,1425,1449]" italics="true" pageId="164" pageNumber="163">
|
||
<collectingRegion id="C445AAFB10C2F1B992A452A223CCFA2E" box="[480,667,1425,1449]" pageId="164" pageNumber="163">Navoi Province</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="035E32C210C2F1B991FD52A22193FA4E" LSID="urn:lsid:plazi:treatment:8E28D50F10C2F1B7924754FE230BF91C:035E32C210C2F1B991FD52A22193FA4E" country="Uzbekistan" latitude="41.6" longLatPrecision="693" longitude="64.18" name="Tamdy District" pageId="164" pageNumber="163" stateProvince="Navoi Province">Tamdy District</location>
|
||
,
|
||
<locationDeviation id="178C7D3B10C2F1B99391528220B1FA4E" box="[213,486,1457,1481]" location="Zarafshan" pageId="164" pageNumber="163">
|
||
<quantity id="C179C9FC10C2F1B9939152822046FA4E" box="[213,273,1457,1481]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="164" pageNumber="163" unit="km" value="1.0">1 km</quantity>
|
||
NW of Zarafshan
|
||
</locationDeviation>
|
||
,
|
||
<date id="723F42D910C2F1B992B3528223CDFA4E" box="[503,666,1457,1481]" pageId="164" pageNumber="163" value="1998-04-22">
|
||
<collectingDate id="627BBB3110C2F1B992B3528223CDFA4E" box="[503,666,1457,1481]" pageId="164" pageNumber="163" value="1998-04-22">22 April 1998</collectingDate>
|
||
</date>
|
||
,
|
||
<geoCoordinate id="63B502DE10C2F1B991EF52822254FA4E" box="[683,771,1457,1481]" degrees="41" direction="north" minutes="36" orientation="latitude" pageId="164" pageNumber="163" precision="925" value="41.6">41°36'N</geoCoordinate>
|
||
<geoCoordinate id="63B502DE10C2F1B9933552E22192FA6E" box="[113,197,1489,1513]" degrees="64" direction="east" minutes="11" orientation="longitude" pageId="164" pageNumber="163" precision="925" value="64.183334">64°11'E</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="63B502DE10C2F1B9939252E22065FA6E" box="[214,306,1489,1513]" degrees="41.60" direction="north" orientation="latitude" pageId="164" pageNumber="163" precision="555" value="41.6">41.60°N</geoCoordinate>
|
||
<geoCoordinate id="63B502DE10C2F1B9927E52E220C2FA6E" box="[314,405,1489,1513]" degrees="64.18" direction="east" orientation="longitude" pageId="164" pageNumber="163" precision="555" value="64.18">64.18°E</geoCoordinate>
|
||
),
|
||
<specimenCount id="1087AF9010C2F1B992EE52E2209CFA6E" box="[426,459,1489,1513]" pageId="164" pageNumber="163" type="male">1♂</specimenCount>
|
||
(
|
||
<typeStatus id="D93ADABB10C2F1B9929852E22368FA6E" box="[476,575,1489,1513]" pageId="164" pageNumber="163" type="holotype">holotype</typeStatus>
|
||
), leg.
|
||
<collectorName id="AB7401CF10C2F1B991C652E223A8FA6E" box="[642,767,1489,1513]" pageId="164" pageNumber="163">A. Gromov</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6E96E4410C2F1B9933552C22305F9CE" collectingDate="1998-04-18" collectionCode="NMPC" collectorName="A. Gromov" country="Uzbekistan" latitude="41.57" location="Zarafshan" longLatPrecision="693" longitude="64.22" municipality="Tamdy District" pageId="164" pageNumber="163" specimenCount="1" specimenCount-female="1" stateProvince="Navoi Province" typeStatus="paratype">
|
||
<collectionCode id="6090FCDC10C2F1B9933552C22197F98E" box="[113,192,1521,1545]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="164" pageNumber="163">NMPC</collectionCode>
|
||
;
|
||
<collectingMunicipality id="E65AFE6310C2F1B9939452C2202FF98E" box="[208,376,1521,1545]" pageId="164" pageNumber="163">Tamdy District</collectingMunicipality>
|
||
,
|
||
<quantity id="C179C9FC10C2F1B992CC52C2208AF98E" box="[392,477,1521,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="164" pageNumber="163" unit="km" value="1.5" valueMax="2.0" valueMin="1.0">1–2 km</quantity>
|
||
from
|
||
<location id="035E32C210C2F1B9916D52C223C1F98E" LSID="urn:lsid:plazi:treatment:8E28D50F10C2F1B7924754FE230BF91C:035E32C210C2F1B9916D52C223C1F98E" box="[553,662,1521,1545]" country="Uzbekistan" latitude="41.57" longLatPrecision="693" longitude="64.22" municipality="Tamdy District" name="Zarafshan" pageId="164" pageNumber="163" stateProvince="Navoi Province">Zarafshan</location>
|
||
,
|
||
<date id="723F42D910C2F1B991E252C221FEF9AE" pageId="164" pageNumber="163" value="1998-04-18">
|
||
<collectingDate id="627BBB3110C2F1B991E252C221FEF9AE" pageId="164" pageNumber="163" value="1998-04-18">18 April 1998</collectingDate>
|
||
</date>
|
||
,
|
||
<geoCoordinate id="63B502DE10C2F1B993FC51222047F9AE" box="[184,272,1553,1577]" degrees="41" direction="north" minutes="34" orientation="latitude" pageId="164" pageNumber="163" precision="925" value="41.566666">41°34'N</geoCoordinate>
|
||
<geoCoordinate id="63B502DE10C2F1B9925F51222027F9AE" box="[283,368,1553,1577]" degrees="64" direction="east" minutes="13" orientation="longitude" pageId="164" pageNumber="163" precision="925" value="64.21667">64°13'E</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="63B502DE10C2F1B992C751222089F9AE" box="[387,478,1553,1577]" degrees="41.57" direction="north" orientation="latitude" pageId="164" pageNumber="163" precision="555" value="41.57">41.57°N</geoCoordinate>
|
||
<geoCoordinate id="63B502DE10C2F1B992AD51222313F9AE" box="[489,580,1553,1577]" degrees="64.22" direction="east" orientation="longitude" pageId="164" pageNumber="163" precision="555" value="64.22">64.22°E</geoCoordinate>
|
||
),
|
||
<specimenCount id="1087AF9010C2F1B9911F51222326F9AE" box="[603,625,1553,1577]" pageId="164" pageNumber="163" type="female">1♀</specimenCount>
|
||
(incomplete;
|
||
<typeStatus id="D93ADABB10C2F1B9933551022186F9CE" box="[113,209,1585,1609]" pageId="164" pageNumber="163" type="paratype">paratype</typeStatus>
|
||
), leg.
|
||
<collectorName id="AB7401CF10C2F1B99254510220DEF9CE" box="[272,393,1585,1609]" pageId="164" pageNumber="163">A. Gromov</collectorName>
|
||
(DNA-165, 7359)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="6090FCDC10C2F1B99118510223FCF9CE" box="[604,683,1585,1609]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="164" pageNumber="163">NMPC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E9B379210C2F1B9933551422002F86E" pageId="164" pageNumber="163" type="etymology">
|
||
<paragraph id="063E641910C2F1B9933551422034F8EE" blockId="164.[113,771,1649,1897]" pageId="164" pageNumber="163">
|
||
ETYMOLOGY. The new species is named in honor of our colleague and friend Sergei Zonstein (Tel Aviv, Israel’ originally from
|
||
<collectingCountry id="7E96248910C2F1B99261518220F6F94E" box="[293,417,1713,1737]" name="Kyrgyzstan" pageId="164" pageNumber="163">Kyrgyzstan</collectingCountry>
|
||
) who made great contributions to arachnology over the decades. Among many other things, Sergei was instrumental in re-examining one of the most interesting scorpions known (and possibly extinct), a troglobitic
|
||
<taxonomicName id="C1811F9A10C2F1B993A85002239BF8CE" authority="Levy, 2007" authorityName="Levy" authorityYear="2007" box="[236,716,1841,1866]" class="Arachnida" family="Akravidae" genus="Akrav" kingdom="Animalia" order="Scorpiones" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="israchanani">
|
||
<emphasis id="34F5B80B10C2F1B993A8500220E0F8CE" box="[236,439,1841,1865]" italics="true" pageId="164" pageNumber="163">Akrav israchanani</emphasis>
|
||
Levy, 2007 (Akravidae)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="621019E810C2F1B9933550612045F8EE" author="FET" box="[113,274,1873,1897]" pageId="164" pageNumber="163" pagination="1 - 49" refId="ref84106" refString="FET, V., M. E. SOLEGLAD & S. L. ZONSTEIN. 2011. The genus Akrav Levy, 2007 (Scorpiones: Akravidae) revisited. Euscorpius, 134: 1 - 49." type="journal article" year="2011">Fet et al., 2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="621019E810C2F1B9925950622000F8EE" author="FET" box="[285,343,1873,1897]" pageId="164" pageNumber="163" pagination="1 - 12" refId="ref84146" refString="FET, V., M. E. SOLEGLAD, S. L. ZONSTEIN, I. NAAMAN, S. LUBATON, B. LANGFORD & A. FRUMKIN. 2017. The second record of a relict Akrav israchanani Levy, 2007 (Scorpiones: Akravidae) from Levana Cave, Israel. Euscorpius, 247: 1 - 12." type="journal article" year="2017">2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="063E641910C2F1B9933550A22002F86E" blockId="164.[113,771,1937,2025]" pageId="164" pageNumber="163">
|
||
DNA DATA.
|
||
<taxonomicName id="C1811F9A10C2F1B993BE50A220B3F82E" authority="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov, 2022" authorityName="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov" authorityYear="2022" box="[250,484,1937,1961]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="164" pageNumber="163" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="34F5B80B10C2F1B993BE50A220B3F82E" box="[250,484,1937,1961]" italics="true" pageId="164" pageNumber="163">Mesobuthus zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FC6057010C2F1B992AA50A2237AF82E" box="[494,557,1937,1961]" pageId="164" pageNumber="163" rank="species">
|
||
<emphasis id="34F5B80B10C2F1B992AA50A22350F82E" bold="true" box="[494,519,1937,1961]" pageId="164" pageNumber="163">sp</emphasis>
|
||
.
|
||
<emphasis id="34F5B80B10C2F1B9915350A22371F82E" bold="true" box="[535,550,1937,1961]" pageId="164" pageNumber="163">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was represented in our survey by a single DNA sequence from
|
||
<collectingCountry id="7E96248910C2F1B99178508223E3F84E" box="[572,692,1969,1993]" name="Uzbekistan" pageId="164" pageNumber="163">Uzbekistan</collectingCountry>
|
||
(Navoi Province) (
|
||
<tableCitation id="4B0351A210C2F1B993A350E2201FF86E" box="[231,328,2001,2025]" captionStart="Table 16" captionStartId="177.[116,172,1991,2012]" captionText="Table 16. Label data for ingroup specimen vouchers, labels used on phylogenetic tree, and GenBank accession numbers for samples used in the phylogenetic analysis." pageId="164" pageNumber="163">Table 16</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E9B379210C2F1B9907457D82406FB85" pageId="164" pageNumber="163" type="diagnosis">
|
||
<paragraph id="063E641910C2F1B9907457D82406FB85" blockId="164.[816,1474,235,1027]" pageId="164" pageNumber="163">
|
||
DIAGNOSIS. Total length of adult male
|
||
<quantity id="C179C9FC10C2F1B9978757D82446FE85" box="[1219,1297,235,259]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="164" pageNumber="163" unit="mm" value="41.0">41 mm</quantity>
|
||
. Trichobothrium
|
||
<emphasis id="34F5B80B10C2F1B990745638221CFEA4" box="[816,843,267,291]" italics="true" pageId="164" pageNumber="163">db</emphasis>
|
||
on fixed finger of pedipalp situated between trichobothria
|
||
<emphasis id="34F5B80B10C2F1B9907456182219FEC5" box="[816,846,299,322]" italics="true" pageId="164" pageNumber="163">est</emphasis>
|
||
and
|
||
<emphasis id="34F5B80B10C2F1B9903A561822F5FEC4" box="[894,930,299,323]" italics="true" pageId="164" pageNumber="163">esb</emphasis>
|
||
, very near to
|
||
<emphasis id="34F5B80B10C2F1B997685618251DFEC5" box="[1068,1098,299,322]" italics="true" pageId="164" pageNumber="163">est</emphasis>
|
||
. Both sexes with fingers proximally slightly twisted. Pedipalp chela length/width ratio
|
||
<quantity id="C179C9FC10C2F1B9963556782496FEE4" box="[1393,1473,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="7.5692" pageId="164" pageNumber="163" unit="in" value="2.98">2.98 in</quantity>
|
||
female and
|
||
<quantity id="C179C9FC10C2F1B990EE565822A3FE04" box="[938,1012,363,387]" metricMagnitude="-1" metricUnit="m" metricValue="1.08712" pageId="164" pageNumber="163" unit="in" value="4.28">4.28 in</quantity>
|
||
male. Pectinal teeth number
|
||
<quantity id="C179C9FC10C2F1B99660565824D5FE04" box="[1316,1410,363,387]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.366" metricValueMin="7.112" pageId="164" pageNumber="163" unit="in" value="28.5" valueMax="29.0" valueMin="28.0">28–29 in</quantity>
|
||
male,
|
||
<quantity id="C179C9FC10C2F1B9907456B922C0FE24" box="[816,919,394,419]" metricMagnitude="-1" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" pageId="164" pageNumber="163" unit="in" value="24.5" valueMax="25.0" valueMin="24.0">24–25 in</quantity>
|
||
female. Chelicerae yellow, without reticulation. Pedipalps and metasoma very sparsely hirsute. Carapace and tergites yellowish brown, black pigmented; metasoma, telson, pedipalps and legs yellowish brown with spots sparsely and black part of metasoma V. Femur of pedipalp with 4–5 granulated carinae. Patella with 8 carinae. Chela lacks carinae. Movable fingers of pedipalps with 12 cutting rows of denticles and 5 terminal denticles. Central lateral and posterior lateral carinae of carapace not joined to form a continuous linear series of granules to posterior margin. Sternite VII smooth, with 4 well marked granulated carinae. Metasomal segment I with 10 carinae; segments II to IV with 8 carinae, other two carinae are indicated by incomplete row of denticles on metasomal segments II and III; segment V with 5 carinae. Metasoma with all carinae granulated. Intercarinal surfaces on metasoma I laterally smooth. Length to depth ratio of metasoma III 1.3, metasoma IV
|
||
<quantity id="C179C9FC10C2F1B990BE54B8256FFC24" box="[1018,1080,907,931]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="164" pageNumber="163" unit="in" value="1.8">1.8 in</quantity>
|
||
male. Telotarsus III ventral setation represented by short and strong spiniform setae. Tarsi hirsute, in adults with 10–15 retroinferior macrosetae on basitarsus III. Telson elongated. Anal lobe divided into two parts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E9B379210C2F1B790745318205AF9DD" lastPageId="170" lastPageNumber="169" pageId="164" pageNumber="163" type="description">
|
||
<paragraph id="063E641910C2F1B9907453182469FB24" blockId="164.[816,1474,1067,2019]" pageId="164" pageNumber="163">
|
||
DESCRIPTION.Totallengthofadultmale
|
||
<quantity id="C179C9FC10C2F1B9978D53182443FBC5" box="[1225,1300,1067,1091]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="164" pageNumber="163" unit="mm" value="41.0">41 mm</quantity>
|
||
.Trichobothrium
|
||
<emphasis id="34F5B80B10C2F1B990745378221CFBE4" box="[816,843,1099,1123]" italics="true" pageId="164" pageNumber="163">db</emphasis>
|
||
on fixed finger of pedipalp situated between trichobothria
|
||
<emphasis id="34F5B80B10C2F1B9907453582219FB05" box="[816,846,1131,1154]" italics="true" pageId="164" pageNumber="163">est</emphasis>
|
||
and
|
||
<emphasis id="34F5B80B10C2F1B990C3535822FCFB04" box="[903,939,1131,1155]" italics="true" pageId="164" pageNumber="163">esb</emphasis>
|
||
, near to
|
||
<emphasis id="34F5B80B10C2F1B9975453582579FB05" box="[1040,1070,1131,1154]" italics="true" pageId="164" pageNumber="163">est</emphasis>
|
||
. Both sexes have fingers proximally slightly twisted. For measurements, see
|
||
<tableCitation id="4B0351A210C2F1B9979F53B8246DFB24" box="[1243,1338,1163,1187]" captionStart="Table 13" captionStartId="153.[115,171,989,1010]" captionText="Table 13. Comparative measurements of Mesobuthus vignolii sp. n. and M. zonsteini sp. n. types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="164" pageNumber="163">Table 13</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="063E641910C2F1B990745399252FFAE4" blockId="164.[816,1474,1067,2019]" pageId="164" pageNumber="163">
|
||
<emphasis id="34F5B80B10C2F1B99074539922FCFB45" bold="true" box="[816,939,1194,1218]" pageId="164" pageNumber="163">Coloration</emphasis>
|
||
(Figs. 1120–1121). Carapace and tergites are yellowish brown with dark pigmentation. Telson is yellow. Metasoma, pedipalps and legs are yellowish brown with several dark spots, part of metasomal segment V black. Chelicerae yellow, without reticulation, the tips of denticles on cheliceral fingers are black.
|
||
</paragraph>
|
||
<paragraph id="063E641910C2F1B9907452582430F964" blockId="164.[816,1474,1067,2019]" pageId="164" pageNumber="163">
|
||
<emphasis id="34F5B80B10C2F1B9907452582537FA04" bold="true" box="[816,1120,1387,1411]" pageId="164" pageNumber="163">Mesosoma and carapace</emphasis>
|
||
(Figs. 1147–1150). Carapace carinate, unevenly covered by granules of varying size; much of the granulation is fine but some granules are larger and distinctly rounded. Tergites I–VI with three carinae and are sparsely granulated. Tergite VII is pentacarinate. Pectinal tooth count
|
||
<quantity id="C179C9FC10C2F1B990EA5138255CF9A4" box="[942,1035,1547,1571]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.366" metricValueMin="7.112" pageId="164" pageNumber="163" unit="in" value="28.5" valueMax="29.0" valueMin="28.0">28–29 in</quantity>
|
||
male,
|
||
<quantity id="C179C9FC10C2F1B99714513925FAF9A4" box="[1104,1197,1546,1571]" metricMagnitude="-1" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" pageId="164" pageNumber="163" unit="in" value="24.5" valueMax="25.0" valueMin="24.0">24–25 in</quantity>
|
||
female. Pectinal marginal tips extend to the end of the sternite IV in male and female. Pectines have three marginal lamellae and eight to nine middle lamellae. Lamellae with numerous long setae, each fulcrum with three to five dark setae. All sternites are smooth and sparsely hirsute. Sternite VII with four well marked granulated carinae. Other sternites have two furrows.
|
||
</paragraph>
|
||
<paragraph id="063E641910C2F1B9907451D922CEF864" blockId="164.[816,1474,1067,2019]" pageId="164" pageNumber="163">
|
||
<emphasis id="34F5B80B10C2F1B9907451D922F6F885" bold="true" box="[816,929,1770,1794]" pageId="164" pageNumber="163">Pedipalps</emphasis>
|
||
(Figs. 1127–1146). Pedipalps hirsute and smooth. Femur has four to five granulated carinae, the middle carina on internal surface consists of strong irregular granules. Patella with eight carinae, of which internal and dorsal are granulated. The chela is smooth. Movable fingers of pedipalps have 11–12 cutting rows of denticles, each with external and internal denticles present, and five terminal denticles.
|
||
</paragraph>
|
||
<caption id="52FE349110C3F1B8933550EF2441F875" box="[113,1302,2012,2034]" pageId="165" pageNumber="164" startId="165.[113,190,2012,2034]">
|
||
<paragraph id="063E641910C3F1B8933550EF2441F875" blockId="165.[113,1302,2012,2034]" box="[113,1302,2012,2034]" pageId="165" pageNumber="164">
|
||
<emphasis id="34F5B80B10C3F1B8933550EF2079F876" bold="true" box="[113,302,2012,2034]" pageId="165" pageNumber="164">Figures 1120–1121</emphasis>
|
||
:
|
||
<taxonomicName id="C1811F9A10C3F1B8927E50EF235EF875" authority="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov, 2022" authorityName="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov" authorityYear="2022" box="[314,521,2012,2034]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="165" pageNumber="164" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="34F5B80B10C3F1B8927E50EF235EF875" box="[314,521,2012,2034]" italics="true" pageId="165" pageNumber="164">Mesobuthus zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FC6057010C3F1B8915450EF2312F875" box="[528,581,2012,2034]" pageId="165" pageNumber="164" rank="species">
|
||
<emphasis id="34F5B80B10C3F1B8915450EF2370F876" bold="true" box="[528,551,2012,2033]" pageId="165" pageNumber="164">sp</emphasis>
|
||
.
|
||
<emphasis id="34F5B80B10C3F1B8917650EF2368F876" bold="true" box="[562,575,2012,2033]" pageId="165" pageNumber="164">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype, dorsal (1120) and ventral (1121) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="52FE349110C0F1BB933050F024FAF871" ID-DOI="http://doi.org/10.5281/zenodo.7163175" ID-Zenodo-Dep="7163175" httpUri="https://zenodo.org/record/7163175/files/figure.png" pageId="166" pageNumber="165" startId="166.[116,193,1987,2009]" targetBox="[117,1466,277,1903]" targetPageId="166">
|
||
<paragraph id="063E641910C0F1BB933050F024FAF871" blockId="166.[116,1474,1987,2038]" pageId="166" pageNumber="165">
|
||
<emphasis id="34F5B80B10C0F1BB933050F02078F85F" bold="true" box="[116,303,1987,2009]" pageId="166" pageNumber="165">Figures 1122–1126</emphasis>
|
||
:
|
||
<taxonomicName id="C1811F9A10C0F1BB927E50F0235FF85E" authority="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov, 2022" authorityName="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov" authorityYear="2022" box="[314,520,1987,2009]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="34F5B80B10C0F1BB927E50F0235FF85E" box="[314,520,1987,2009]" italics="true" pageId="166" pageNumber="165">Mesobuthus zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FC6057010C0F1BB914950F02315F85E" box="[525,578,1987,2009]" pageId="166" pageNumber="165" rank="species">
|
||
<emphasis id="34F5B80B10C0F1BB914950F02373F85F" bold="true" box="[525,548,1987,2008]" pageId="166" pageNumber="165">sp</emphasis>
|
||
.
|
||
<emphasis id="34F5B80B10C0F1BB916B50F0236BF85F" bold="true" box="[559,572,1987,2008]" pageId="166" pageNumber="165">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="34F5B80B10C0F1BB910350F02255F85F" bold="true" box="[583,770,1987,2009]" pageId="166" pageNumber="165">Figures 1122–1125</emphasis>
|
||
. Male holotype, telson lateral (1122), metasoma and telson, lateral (1123), dorsal (1124), and ventral (1125) views.
|
||
<emphasis id="34F5B80B10C0F1BB92BE50D32324F872" bold="true" box="[506,627,2016,2038]" pageId="166" pageNumber="165">Figure 1126</emphasis>
|
||
. Female paratype, metasoma IV–V and telson lateral. Scale bar: 10 mm (1123–1125).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="52FE349110C1F1BA9337505E250EF871" ID-DOI="http://doi.org/10.5281/zenodo.7163177" ID-Zenodo-Dep="7163177" httpUri="https://zenodo.org/record/7163177/files/figure.png" pageId="167" pageNumber="166" startId="167.[115,192,1901,1923]" targetBox="[130,1466,231,1890]" targetPageId="167">
|
||
<paragraph id="063E641910C1F1BA9337505E250EF871" blockId="167.[115,1473,1901,2039]" pageId="167" pageNumber="166">
|
||
<emphasis id="34F5B80B10C1F1BA9337505E2066F805" bold="true" box="[115,305,1901,1923]" pageId="167" pageNumber="166">Figures 1127–1146</emphasis>
|
||
:
|
||
<taxonomicName id="C1811F9A10C1F1BA9204505E2346F804" authority="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov, 2022" authorityName="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov" authorityYear="2022" box="[320,529,1901,1923]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="34F5B80B10C1F1BA9204505E2346F804" box="[320,529,1901,1923]" italics="true" pageId="167" pageNumber="166">Mesobuthus zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FC6057010C1F1BA915E505D2306F804" box="[538,593,1902,1923]" pageId="167" pageNumber="166" rank="species">
|
||
<emphasis id="34F5B80B10C1F1BA915E505D2366F804" bold="true" box="[538,561,1902,1923]" pageId="167" pageNumber="166">sp</emphasis>
|
||
.
|
||
<emphasis id="34F5B80B10C1F1BA917B505D231BF804" bold="true" box="[575,588,1902,1923]" pageId="167" pageNumber="166">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, pedipalp.
|
||
<emphasis id="34F5B80B10C1F1BA9185505E22D7F805" bold="true" box="[705,896,1901,1923]" pageId="167" pageNumber="166">Figures 1127–1136</emphasis>
|
||
. Male holotype, chela, dorsal (1127), external (1128), and ventral (1129) views. Patella, dorsal (1130), external (1131) and ventral (1132) views. Femur and trochanter, internal (1133), dorsal (1134), and ventral (1135) views. Movable finger (1136).
|
||
<emphasis id="34F5B80B10C1F1BA9169509423BDF83B" bold="true" box="[557,746,1959,1981]" pageId="167" pageNumber="166">Figures 1137–1146</emphasis>
|
||
. Female paratype, chela, dorsal (1137), external (1138), and ventral (1139) views. Patella, dorsal (1140), external (1141) and ventral (1142) views. Femur and trochanter, internal (1143), dorsal (1144), and ventral (1145) views. Movable finger (1146). The trichobothrial pattern is indicated in Figures 1127–1131, 1133–1134.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="52FE349110CEF1B593305095221CF871" ID-DOI="http://doi.org/10.5281/zenodo.7163179" ID-Zenodo-Dep="7163179" httpUri="https://zenodo.org/record/7163179/files/figure.png" pageId="168" pageNumber="167" startId="168.[116,193,1958,1980]" targetBox="[119,1474,242,1948]" targetPageId="168">
|
||
<paragraph id="063E641910CEF1B593305095221CF871" blockId="168.[116,1475,1958,2038]" pageId="168" pageNumber="167">
|
||
<emphasis id="34F5B80B10CEF1B5933050952067F83C" bold="true" box="[116,304,1958,1980]" pageId="168" pageNumber="167">Figures 1147–1154</emphasis>
|
||
:
|
||
<taxonomicName id="C1811F9A10CEF1B5927F5095235DF83B" authority="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov, 2022" authorityName="Kovařík & Fet & Gantenbein & Graham & Aydin & Yağmur & Šťáhlavský & Nikita & Poverennyi & Novruzov" authorityYear="2022" box="[315,522,1958,1980]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="168" pageNumber="167" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
|
||
<emphasis id="34F5B80B10CEF1B5927F5095235DF83B" box="[315,522,1958,1980]" italics="true" pageId="168" pageNumber="167">Mesobuthus zonsteini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FC6057010CEF1B5914B50952313F83B" box="[527,580,1958,1980]" pageId="168" pageNumber="167" rank="species">
|
||
<emphasis id="34F5B80B10CEF1B5914B50952371F83C" bold="true" box="[527,550,1958,1979]" pageId="168" pageNumber="167">sp</emphasis>
|
||
.
|
||
<emphasis id="34F5B80B10CEF1B5917550952369F83C" bold="true" box="[561,574,1958,1979]" pageId="168" pageNumber="167">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="34F5B80B10CEF1B5910E5095239CF83C" bold="true" box="[586,715,1958,1980]" pageId="168" pageNumber="167">Figures 1147</emphasis>
|
||
,
|
||
<emphasis id="34F5B80B10CEF1B5919250952252F83C" bold="true" box="[726,773,1958,1979]" pageId="168" pageNumber="167">1149</emphasis>
|
||
,
|
||
<emphasis id="34F5B80B10CEF1B590545095222EF83C" bold="true" box="[784,889,1958,1980]" pageId="168" pageNumber="167">1151–1154</emphasis>
|
||
. Male holotype, chelicerae, carapace and tergites I–V (1147), sternopectinal region and sternites (1149), and left legs I–IV, retrolateral aspect (1151–1154).
|
||
<emphasis id="34F5B80B10CEF1B590A850F0253AF85F" bold="true" box="[1004,1133,1987,2009]" pageId="168" pageNumber="167">Figures 1148</emphasis>
|
||
,
|
||
<emphasis id="34F5B80B10CEF1B5973C50F025F0F85F" bold="true" box="[1144,1191,1987,2008]" pageId="168" pageNumber="167">1150</emphasis>
|
||
. Female paratype, chelicerae, carapace and tergites (1148), and sternopectinal region and sternites (1150).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="52FE349110CFF1B4933750E825D0F876" ID-DOI="http://doi.org/10.5281/zenodo.7163181" ID-Zenodo-Dep="7163181" box="[115,1159,2011,2033]" httpUri="https://zenodo.org/record/7163181/files/figure.png" pageId="169" pageNumber="168" startId="169.[115,181,2011,2033]" targetBox="[114,1474,234,1911]" targetPageId="169">
|
||
<paragraph id="063E641910CFF1B4933750E825D0F876" blockId="169.[115,1159,2011,2033]" box="[115,1159,2011,2033]" pageId="169" pageNumber="168">
|
||
<emphasis id="34F5B80B10CFF1B4933750E821BFF877" bold="true" box="[115,232,2011,2033]" pageId="169" pageNumber="168">Figure 1155</emphasis>
|
||
. Type localities of
|
||
<taxonomicName id="C1811F9A10CFF1B492DE50E82359F877" authorityName="Vachon" authorityYear="1950" box="[410,526,2011,2032]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="169" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="34F5B80B10CFF1B492DE50E82359F877" box="[410,526,2011,2032]" italics="true" pageId="169" pageNumber="168">Mesobuthus</emphasis>
|
||
</taxonomicName>
|
||
taxa (except
|
||
<taxonomicName id="C1811F9A10CFF1B491CB50E822D8F877" authorityName="Birula" authorityYear="1911" box="[655,911,2011,2033]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="169" pageNumber="168" phylum="Arthropoda" rank="subSpecies" species="eupeus" subSpecies="mongolicus">
|
||
<emphasis id="34F5B80B10CFF1B491CB50E822D8F877" box="[655,911,2011,2033]" italics="true" pageId="169" pageNumber="168">Buthus eupeus mongolicus</emphasis>
|
||
</taxonomicName>
|
||
which is in Figure 1158).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="52FE349110CCF1B79337531C24D1FBC2" ID-DOI="http://doi.org/10.5281/zenodo.7163185" ID-Zenodo-Dep="7163185" box="[115,1414,1071,1093]" httpUri="https://zenodo.org/record/7163185/files/figure.png" pageId="170" pageNumber="169" startId="170.[115,181,1071,1093]" targetBox="[113,1474,234,1062]" targetPageId="170">
|
||
<paragraph id="063E641910CCF1B79337531C24D1FBC2" blockId="170.[115,1414,1071,1093]" box="[115,1414,1071,1093]" pageId="170" pageNumber="169">
|
||
<emphasis id="34F5B80B10CCF1B79337531C21BFFBC3" bold="true" box="[115,232,1071,1093]" pageId="170" pageNumber="169">Figure 1156</emphasis>
|
||
. Distribution of
|
||
<taxonomicName id="C1811F9A10CCF1B792C7531C2372FBC2" box="[387,549,1071,1093]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="170" pageNumber="169" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="34F5B80B10CCF1B792C7531C20A0FBC3" box="[387,503,1071,1092]" italics="true" pageId="170" pageNumber="169">Mesobuthus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
in Armenia, Azerbaijan, Georgia, Iraq, Syria, and Turkey. Based on studied material only.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="063E641910CCF1B79337535023E7FA9C" blockId="170.[115,772,1123,1627]" pageId="170" pageNumber="169">
|
||
<emphasis id="34F5B80B10CCF1B79337535021FFFBFC" bold="true" box="[115,168,1123,1147]" pageId="170" pageNumber="169">Legs</emphasis>
|
||
(Figs. 1151–1154). Telotarsus III ventral setation represented by short and strong spiniform setae. Tarsi hirsute, in adults with 10–15 retroinferior macrosetae on basitarsus III. Femur with only several macrosetae. Femur and patella with carinae well developed. Tibial spurs present and media to strong on legs III and IV and absent on the other legs.
|
||
</paragraph>
|
||
<paragraph id="063E641910CCF1B793375210205AF9DD" blockId="170.[115,772,1123,1627]" pageId="170" pageNumber="169">
|
||
<emphasis id="34F5B80B10CCF1B7933752102023FABC" bold="true" box="[115,372,1315,1339]" pageId="170" pageNumber="169">Metasoma and telson</emphasis>
|
||
(Figs. 1122–1126). All metasomal segments only sparsely hirsute. Metasomal segment I with 10 carinae, II–III with 8 carinae but other two lateromedian carinae are indicated by incomplete row of granules, IV with 8 carinae, and V with 5 carinae. Metasoma with all carinae granulated. Intercarinal surfaces on metasoma smooth. Ventrolateral carinae of metasoma V posteriorly and metasoma II–III laterally with several large lobate denticles. Telson is very sparsely hirsute, elongate and smooth. Anal lobe divided into two parts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E9B379210CCF1B7933751B0230BF91C" box="[115,604,1667,1691]" pageId="170" pageNumber="169" type="distribution">
|
||
<paragraph id="063E641910CCF1B7933751B0230BF91C" blockId="170.[115,604,1667,1691]" box="[115,604,1667,1691]" pageId="170" pageNumber="169">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="7E96248910CCF1B7925751B020DCF91C" box="[275,395,1667,1691]" name="Uzbekistan" pageId="170" pageNumber="169">Uzbekistan</collectingCountry>
|
||
(Figs. 1155, 1158).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |